Checklist for Subfamily Scarabaeinae Latreille, 1802 (Dung Beetles)
and region Zambia (East Africa)
Class
| Insecta
Linnaeus, 1758
Insects, True insects | 333
3740
9137 | Families
Genera
Species |
Taxom
|
Subclass
| Pterygota
Lang, 1888
Winged insects | 331
3738
9135 | Families
Genera
Species |
Taxom
|
Infraclass
| Neoptera
van der Wulp, 1890
Modern Wing-folding insects | 309
3639
8842 | Families
Genera
Species |
Taxom
|
Superorder
| Holometabola
Heider, 1889
Holometabolous Insects | 202
3129
7992 | Families
Genera
Species |
Taxom
|
Order
| Coleoptera
Linnaeus, 1758
Beetles, Coleopterans | 53
1092
2610 | Families
Genera
Species |
Taxom
|
Suborder
| Polyphaga
Emery, 1886
Polyphags | 48
914
1934 | Families
Genera
Species |
Taxom
|
Infraorder
| Scarabaeiformia
Crowson, 1960
| 8
231
634 | Families
Genera
Species |
Taxom
|
Superfamily
| Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles | 8
231
634 | Families
Genera
Species |
Taxom
| | Family | Scarabaeidae
Latreille, 1802
Dung beetles, Flower beetles, Fruit chafers, Rain beetles | 214
597 | Genera
Species |
Taxom
| | Subfamily | Scarabaeinae
Latreille, 1802
Dung Beetles, True dung beetles | 50
288 | Genera
Species |
Taxom
|
Genera for Zambia
50 Genera with 288 Species #ACDEGHJKLMNOPST
| Acanthonitis Janssens, 2012 | | 2 | Species | Taxom | | Afrodrepanus Krikken, 1857 | | 1 | Species | Taxom | | Allogymnopleurus Janssens, 1855 | | 2 | Species | Taxom | | Anachalcos Hope, 1871 | | 2 | Species | Taxom | | Caccobius Thomson, 2014 | | 12 | Species | Taxom | | Canthidium Erichson, 1867 | | 1 | Species | Taxom | | Canthon Hoffmannsegg, 1868 | Tumblebug | 1 | Species | Taxom | | Catharsius Hope, 2009 | | 16 | Species | Taxom | | Cheironitis van Lansberge, 1789 | | 2 | Species | Taxom | | Cleptocaccobius Cambefort, 1902 | | 2 | Species | Taxom | | Copridaspidus Bloucomont, 1878 | | 1 | Species | Taxom | | Copris Geoffroy, 1987 | Mexican dung beetle | 17 | Species | Taxom | | Coptodactyla Burmeister, 1877 | | 1 | Species | Taxom | | Coptorhina Hope, 1857 | | 3 | Species | Taxom | | Delopleurus Erichson, 1939 | | 2 | Species | Taxom | | Diastellopalpus van Lansberge, 2000 | | 5 | Species | Taxom | | Digitonthophagus Balthasar, 1787 | | 2 | Species | Taxom | | Eodrepanus Barbero, Pal..., 1877 | | 2 | Species | Taxom | | Euoniticellus Janssens, 1873 | | 2 | Species | Taxom | | Euonthophagus Balthasar, 1901 | | 2 | Species | Taxom | | Eusaproecius Branco, 1923 | | 2 | Species | Taxom | | Garreta Janssens, 1857 | | 6 | Species | Taxom | | Gilletellus Janssens, 1937 | | 1 | Species | Taxom | | Gymnopleurus Illiger, 1843 | | 2 | Species | Taxom | | Heliocopris Hope, 1893 | | 15 | Species | Taxom | | Heteroclitopus Péringuey, 2007 | | 1 | Species | Taxom | | Heteronitis Gillet, 1862 | | 1 | Species | Taxom | | Hyalonthophagus Palestrini &..., 2021 | | 3 | Species | Taxom | | Janssensantus Paulian, 2022 | | 1 | Species | Taxom | | Kheper Janssens, 1886 | | 4 | Species | Taxom | | Latodrepanus Krikken, 2009 | | 3 | Species | Taxom | | Metacatharsius Paulian, 1857 | | 6 | Species | Taxom | | Milichus Péringuey, 1907 | | 2 | Species | Taxom | | Neosisyphus Müller, 1974 | | 2 | Species | Taxom | | Onitis Fabricius, 2012 | | 18 | Species | Taxom | | Onthophagus Latreille, 2022 | | 74 | Species | Taxom | | Pachylomera Griffith & P..., 1874 | | 2 | Species | Taxom | | Pedaria Laporte, 2003 | | 17 | Species | Taxom | | Phalops Erichson, 2003 | | 6 | Species | Taxom | | Pinacotarsus Harold, 1875 | | 1 | Species | Taxom | | Pleuronitis Lansberge, 2011 | | 1 | Species | Taxom | | Proagoderus Lansberge, 1901 | | 14 | Species | Taxom | | Pseudocopris Ferreira, 1910 | | 2 | Species | Taxom | | Sarophorus Erichson, 1901 | | 1 | Species | Taxom | | Scaptocnemis Péringuey, 1901 | | 1 | Species | Taxom | | Scarabaeus Linnaeus, 1901 | | 6 | Species | Taxom | | Sceliages Westwood, 1908 | | 1 | Species | Taxom | | Sisyphus Latreille, 2015 | | 16 | Species | Taxom | | Stiptopodius Harold, 1901 | | 2 | Species | Taxom | | Tiniocellus Péringuey, 1851 | | 2 | Species | Taxom |
Species list for Zambia
288 Species and 9 Subspecies | Acanthonitis nicolasi Cambefort, 1995 | | | Acanthonitis spissus Josso, 2012 | | | Afrodrepanus impressicollis Boheman, 1857 | | | Allogymnopleurus splendidus Bertoloni, 1849 | | | Allogymnopleurus thalassinus Klug, 1855 | | | Anachalcos convexus Boheman, 1857 | | | Anachalcos procerus Gerstaecker, 1871 | | | Caccobius auberti d´Orbigny, 1902 | | | Caccobius cavatus d´Orbigny, 1908 | | | Caccobius ferrugineus Fahraeus, 1857 | | | Caccobius globaticeps d´Orbigny, 1905 | | | Caccobius inconspicuus (Fahraeus, 1857) | ⇒ Caccophilus fuliginosus Roth, 1851 | | Caccobius inops Péringuey, 1901 | | | Caccobius longipennis d´Orbigny, 1904 | | | Caccobius nigritulus Klug, 1855 | | | Caccobius pentagonus d´Orbigny, 1908 | | | Caccobius pseudocorniceps Josso, 2018 | | | Caccobius tuberifrons d´Orbigny, 1902 | | | Caccobius zambiensis Walter, 2014 | | | Canthidium smaragdinum Harold, 1867 | | | Canthon dives Harold, 1868 | | | Catharsius aegeus Génier, 2017 | | | Catharsius brittoni Ferreira, 1963 | | | Catharsius coptorhinaformis Josso & Prév..., 2009 | | | Catharsius dux Harold, 1878 | | | Catharsius folicornis Josso & Prév..., 2019 | | | Catharsius heros Boheman, 1860 | | | Catharsius juheli Josso, 2011 | | | Catharsius melancholicus (Boheman, 1860) | ⇒ Catharsius longiceps Gillet, 1907 | | Catharsius pallas Génier, 2017 | | | Catharsius pandion Harold, 1877 | | | Catharsius platicornis Ferreira, 1960 | | | Catharsius sesostris Waterhouse, 1888 | | | Catharsius stricticornis Josso & Prév..., 2019 | | | Catharsius takanoi Josso & Prév..., 2019 | | | Catharsius tricornutus DeGeer, 1778 | | | Catharsius tridens Josso & Prév..., 2009 | | | Cheironitis hungaricus Herbst, 1789 | | | Cheironitis ungaricus ungaricus Herbst, 1789 | | | Cleptocaccobius convexifrons Raffray, 1877 | | | Cleptocaccobius schaedlei d´Orbigny, 1902 | | | Copridaspidus peregrinus Harold, 1878 | | | Copris amyntor (Klug, 1855) | ⇒ Copris harrisi Waterhouse, 1891 | | Copris armiger Gillet, 1910 | | | Copris bootes Klug, 1855 | | | Copris cassius Péringuey, 1901 | | | Copris elphenor Klug, 1855 | | | Copris evanidus Klug, 1855 | | | Copris fallaciosus Gillet, 1907 | Kenyan dung beetle | | Copris harrisi Waterhouse, 1891 | | | Copris inhalatus perturbator Péringuey, 1901 | | | Copris kasanka Josso & Prév..., 2016 | | | Copris mesacanthus mesacanthus | | | Copris misellus Péringuey, 1901 | | | Copris nepos Gillet, 1908 | | | Copris obesus Boheman, 1857 | | | Copris prevosti Josso, 2011 | | | Copris puncticollis Boheman, 1857 | | | Copris serius Nguyen-phung, 1987 | | | Coptodactyla subaenea Harold, 1877 | | | Coptorhina auspicata Péringuey, 1901 | | | Coptorhina davidi Janssens, 1939 | | | Coptorhina nitidipennis Boheman, 1857 | | | Delopleurus darrenmanni Frolov, 2014 | | | Delopleurus gilletti Janssens, 1939 | | | Diastellopalpus ikelenge Josso & Pré..., 2014 | | | Diastellopalpus infernalis Lansberge, 1887 | | | Diastellopalpus lamellicollis Quedenfeldt, 1884 | | | Diastellopalpus quinquedens Bates, 1888 | | | Diastellopalpus walteri Josso & Pré..., 2000 | | | Digitonthophagus biflagellatus Génier, 2017 | | | Digitonthophagus gazella Fabricius, 1787 | Brown dung beetle | | Eodrepanus bechynei Janssens, 1953 | | | Eodrepanus parallelus Raffray, 1877 | | | Euoniticellus intermedius Reiche, 1848 | Intermediate sandy dung beetle | | Euoniticellus triangulatus Harold, 1873 | | | Euonthophagus carbonarius Klug, 1855 | | | Euonthophagus consobrinus Péringuey, 1901 | | | Eusaproecius magnigena d´Orbigny, 1913 | | | Eusaproecius tinantae Boucomont, 1923 | | | Garreta azareus | | | Garreta dejeani Castelnau, 1840 | | | Garreta laetus olivaceus Quedenfeldt, 1884 | | | Garreta malleolus Kolbe, 1895 | | | Garreta nitens Olivier, 1789 | | | Garreta unicolor Fahraeus, 1857 | | | Gilletellus collarti Janssens, 1937 | | | Gymnopleurus sericeifrons Fairmaire, 1887 | | | Gymnopleurus virens Erichson, 1843 | | | Heliocopris atropos Boheman, 1860 | | | Heliocopris biimpressus Kolbe, 1893 | | | Heliocopris colossus Bates, 1868 | | | Heliocopris eryx ikelenge Moretto, 2017 | | | Heliocopris hamadryas Fabricius, 1775 | | | Heliocopris hamifer Harold, 1878 | | | Heliocopris japetus Klug, 1855 | | | Heliocopris kolbei Felsche, 1901 | | | Heliocopris letiranti Moretto, 2021 | | | Heliocopris marshalli Péringuey, 1901 | | | Heliocopris minos Gillet, 1907 | | | Heliocopris myrmidon Kolbe, 1893 | | | Heliocopris neptunus Boheman, 1857 | | | Heliocopris samson Harold, 1878 | | | Heliocopris staudingeri Kolbe, 1893 | | | Heteroclitopus foveolatus Branco & Josso, 2007 | | | Heteronitis castelnaui Harold, 1862 | | | Hyalonthophagus alcedo Orbigny, 1913 | | | Hyalonthophagus alcyon Klug, 1855 | | | Hyalonthophagus novalcedo Josso, 2021 | | | Janssensantus alatus Josso, 2022 | | | Kheper festivus Harold, 1868 | | | Kheper lamarcki MacLeay, 1821 | | | Kheper prodigiosus Erichson, 1843 | | | Kheper vanderkelleni Lansberge, 1886 | | | Latodrepanus caelatus Gerstaecker, 1871 | | | Latodrepanus laticollis Fahraeus, 1857 | | | Latodrepanus nicolasi Barbero, Pal..., 2009 | | | Metacatharsius dentinum Ferreira, 1964 | | | Metacatharsius exiguus Boheman, 1860 | | | Metacatharsius ferreirae Balthasar, 1965 | | | Metacatharsius opacus Waterhouse, 1891 | | | Metacatharsius peleus Olivier, 1789 | | | Metacatharsius troglodytes (Boheman, 1857) | ⇒ Metacatharsius anderseni Waterhouse, 1891 | | Milichus dudleyae Cambefort, 1996 | | | Milichus jugatus d´Orbigny, 1907 | | | Neosisyphus calcaratus Klug, 1855 | | | Neosisyphus rubrus Paschalidis, 1974 | | | Onitis albertcollarti Génier & Mor..., 2019 | | | Onitis alexis (Klug, 1835) | ⇒ Cheironitis ungaricus ungaricus Herbst, 1789 | | Onitis alexis alexis | | | Onitis brucemillerorum Josso & Mont..., 2017 | | | Onitis cryptodus Gillet, 1918 | | | Onitis cupreus Castelnau, 1840 | | | Onitis deceptor Péringuey, 1901 | | | Onitis inversidens Lansberge, 1875 | | | Onitis kasanka Josso, 2012 | | | Onitis lycophron Klug, 1855 | | | Onitis mendax Gillet, 1918 | | | Onitis perturbator Péringuey, 1901 | | | Onitis pseudosetosus Ferreira, 1976 | | | Onitis robustus Boheman, 1857 | | | Onitis uncinatus Klug, 1855 | | | Onitis viridulus (Boheman, 1857) | ⇒ Onitis pecuarius Lansberge, 1875 | | Onitis westermanni Lansberge, 1886 | | | Onitis zambiensis Josso, 2012 | | | Onthophagus (Afrostrandius) plebejus Klug, 1855 | | | Onthophagus (Euonthophagus) consobrinus Péringuey, 1901 | | | Onthophagus marginalis przewalski Kabakov, 2006 | | | Onthophagus (Proagoderus) alcedo d´Orbigny, 1913 | | | Onthophagus (Proagoderus) aureiceps d´Orbigny, 1902 | | | Onthophagus aequatus Péringuey, 1901 | | | Onthophagus aeruginosus Roth, 1851 | | | Onthophagus alainmonforti Josso, 2022 | | | Onthophagus aliceae Josso, 2019 | | | Onthophagus apiciosus d´Orbigny, 1902 | | | Onthophagus atrosplendens Frey, 1961 | | | Onthophagus axillaris (Boheman, 1860) | ⇒ Onthophagus criniger d´Orbigny, 1904 | | Onthophagus bifidus Reiche, 1849 | | | Onthophagus bituberifrons Josso, 2018 | | | Onthophagus bufonidius Josso, 2019 | | | Onthophagus chalcostolus d´Orbigny, 1902 | | | Onthophagus convexipennis Dierkens, 2022 | | | Onthophagus corniculiger d´Orbigny, 1913 | | | Onthophagus delahayei Josso, 2011 | | | Onthophagus dudleyianus Josso, 2019 | | | Onthophagus ebenicolor d´Orbigny, 1902 | | | Onthophagus emeritus Péringuey, 1901 | | | Onthophagus estelleae Josso, 2022 | | | Onthophagus extraneus Josso, 2021 | | | Onthophagus filicornis Harold, 1873 | | | Onthophagus fimetarius Roth, 1851 | | | Onthophagus flexifrons d´Orbigny, 1908 | | | Onthophagus francoisi Josso, 2013 | | | Onthophagus geminatus d´Orbigny, 1907 | | | Onthophagus granulifer (Harold, 1886) | ⇒ Onthophagus ponticus Harold, 1883 | | Onthophagus granulifrons Frey, 1961 | | | Onthophagus horridus d´Orbigny, 1908 | | | Onthophagus ikelengeensis Josso, 2018 | | | Onthophagus impressicollis Boheman, 1860 | | | Onthophagus inermicollis d´Orbigny, 1908 | | | Onthophagus infuscatus Klug, 1845 | | | Onthophagus insignis (Péringuey, 1896) | ⇒ Proagoderus fossidorsis d´Orbigny, 1905 | | Onthophagus jugicola d´Orbigny, 1902 | | | Onthophagus juniperensis Josso, 2019 | | | Onthophagus juvencus Klug, 1835 | | | Onthophagus kaomaensis Josso, 2022 | | | Onthophagus kapangaensis Dierkens, 2022 | | | Onthophagus katangensis Frey, 1961 | | | Onthophagus lacustris Harold, 1877 | | | Onthophagus lamelliger (Gerstaecker, 1871) | ⇒ Onthophagus ochropygus d´Orbigny, 1902 | | Onthophagus laminidorsis d´Orbigny, 1902 | | | Onthophagus latestriatus d´Orbigny, 1908 | | | Onthophagus marginipennis Josso, 2018 | | | Onthophagus minans d´Orbigny, 1913 | | | Onthophagus minettianus Josso, 2019 | | | Onthophagus mixticeps d´Orbigny, 1905 | | | Onthophagus mwenseensis Dierkens, 2022 | | | Onthophagus nchilaensis Josso, 2022 | | | Onthophagus nigroflavus Josso, 2022 | | | Onthophagus nyikaensis Josso, 2018 | | | Onthophagus obtusicornis (Fahraeus, 1857) | ⇒ Onthophagus marginalis przewalski Kabakov, 2006 | | Onthophagus ochropygus d´Orbigny, 1902 | Common snipe fly | | Onthophagus okahandjanus Balthasar, 1974 | | | Onthophagus parumnotatus Fahraeus, 1857 | | | Onthophagus picatus d´Orbigny, 1902 | | | Onthophagus pronocarinatus Josso, 2019 | | | Onthophagus pseudocribripennis Josso, 2019 | | | Onthophagus pullus (Roth, 1851) | ⇒ Onthophagus parenthesis Boucomont, 1913 | | Onthophagus rojkoffianus Josso, 2018 | | | Onthophagus rufobasalis Fairmaire, 1887 | | | Onthophagus simplex Raffray, 1877 | | | Onthophagus splendidoides Josso, 2013 | | | Onthophagus subdivisus d´Orbigny, 1908 | | | Onthophagus valentineae Josso, 2019 | | | Onthophagus variatus d´Orbigny, 1902 | | | Onthophagus verae Josso, 2013 | | | Onthophagus verticalis (Fahraeus, 1857) | ⇒ Onthophagus praedatus Harold, 1862 | | Onthophagus vinctus Erichson, 1843 | | | Onthophagus zambundaticeps Josso, 2022 | | | Pachylomera femoralis Kirby, 1828 | | | Pachylomera opaca Lansberge, 1874 | | | Pedaria armata Raffray, 1877 | | | Pedaria catherinae Josso & Prév..., 2009 | | | Pedaria conformis Péringuey, 1901 | | | Pedaria cylindrica Fahraeus, 1857 | | | Pedaria decorsei (Boucomont, 1922) | ⇒ Pedaria taylori Waterhouse, 1890 | | Pedaria demeyeri Josso & Prév..., 2003 | | | Pedaria dudleyi Josso & Prév..., 2003 | | | Pedaria hanae Balthasar, 1963 | | | Pedaria montreuili Josso & Prév..., 2015 | | | Pedaria murphyi Josso & Prév..., 2003 | | | Pedaria murphyi murphyi | | | Pedaria puncticollis Waterhouse, 1890 | | | Pedaria rohani Boucomont, 1922 | | | Pedaria sudrei Josso & Prév..., 2003 | | | Pedaria taylori Waterhouse, 1890 | | | Pedaria tenebrosa Balthasar, 1937 | | | Pedaria werneri Josso & Prév..., 2003 | | | Phalops ardea Klug, 1855 | | | Phalops boschas Klug, 1855 | | | Phalops flavocinctus Klug, 1855 | | | Phalops laminifrons Fairmaire, 1882 | | | Phalops smaragdinus Harold, 1875 | | | Phalops zuninoi Barbero & Pa..., 2003 | | | Pinacotarsus dohrni Harold, 1875 | | | Pleuronitis minettii Josso, 2011 | | | Proagoderus bicallosus Klug, 1855 | | | Proagoderus brucei (d´Orbigny, 1902) | ⇒ Onthophagus chalcostolus d´Orbigny, 1902 | | Proagoderus brucei brucei | | | Proagoderus cyanopterus d´Orbigny, 1905 | | | Proagoderus dives Harold, 1877 | | | Proagoderus edithae Josso, 2014 | | | Proagoderus gemmatus Péringuey, 1901 | | | Proagoderus gibbiramus d´Orbigny, 1902 | | | Proagoderus kafueensis Josso, 2014 | | | Proagoderus lanista Castelnau, 1840 | | | Proagoderus louisi Josso, 2014 | | | Proagoderus pyramidalis Klug, 1855 | | | Proagoderus rectefurcatus Fairmaire, 1891 | | | Proagoderus sapphirinus (Péringuey, 1901) | ⇒ Onthophagus aureiceps d´Orbigny, 1902 | | Pseudocopris lunaticeps Gillet, 1910 | | | Pseudocopris tubericollis Felsche, 1910 | | | Sarophorus cicatricosus Péringuey, 1901 | | | Scaptocnemis segregis Péringuey, 1901 | | | Scarabaeus ermienae Deschodt & D..., 2015 | | | Scarabaeus galenus Westwood, 1847 | | | Scarabaeus goryi Castelnau, 1840 | | | Scarabaeus profanus Fabricius, 1792 | | | Scarabaeus sulcipennis Quedenfeldt, 1888 | | | Scarabaeus zambesianus Péringuey, 1901 | | | Sceliages augias Gillet, 1908 | | | Sisyphus (Sisyphus) struwei Schafer & Fi..., 1992 | | | Sisyphus alveatus Boucomont, 1935 | | | Sisyphus bicuariensis Daniel & Davis, 2018 | | | Sisyphus calcaratus (Boucomont, 1937) | ⇒ Neosisyphus rugosus Gory, 1833 | | Sisyphus crispatus Gory, 1833 | | | Sisyphus gazanus Arrow, 1909 | | | Sisyphus goryi (Harold, 1859) | ⇒ Sisyphus caffer goryi Harold, 1859 | | Sisyphus impressipennis Lansberge, 1886 | | | Sisyphus infuscatus (Klug, 1855) | ⇒ Neosisyphus spinipes Thunberg, 1818 | | Sisyphus jossoi Montreuil, 2015 | | | Sisyphus juheli Montreuil, 2015 | | | Sisyphus morettoi Montreuil, 2015 | | | Sisyphus nanniscus Péringuey, 1901 | | | Sisyphus rubrus (Paschalidis, 1974) | ⇒ Neosisyphus rubrus Paschalidis, 1974 | | Sisyphus spinipes (Thunberg, 1818) | ⇒ Neosisyphus spinipes Thunberg, 1818 | | Sisyphus trichodichromicus Montreuil, 2015 | | | Stiptopodius granulosus Branco, 2009 | | | Stiptopodius singularis Péringuey, 1901 | | | Tiniocellus dolosus Branco, 2010 | | | Tiniocellus spinipes Roth, 1851 | |
Subnodes of Scarabaeinae with occurrences in Zambia
Links, sources and further information Scarabaeinae at artfakta.se
Scarabaeinae at dyntaxa.se
Scarabaeinae at itis.gov
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