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Checklist for Subfamily Scarabaeinae Latreille, 1802 (Dung Beetles)
and region Taiwan (East Asia)


Class
 
Insecta
Linnaeus, 1758
Insects, True insects
650
12552
43810
Families
Genera
Species
 
Taxom
 

Subclass
 
Pterygota
Lang, 1888
Winged insects
648
12549
43801
Families
Genera
Species
 
Taxom
 

Infraclass
 
Neoptera
van der Wulp, 1890
Modern Wing-folding insects
622
12414
43343
Families
Genera
Species
 
Taxom
 

Superorder
 
Holometabola
Heider, 1889
Holometabolous Insects
402
9699
34661
Families
Genera
Species
 
Taxom
 

Order
 
Coleoptera
Linnaeus, 1758
Beetles, Coleopterans
112
3405
12308
Families
Genera
Species
 
Taxom
 

Suborder
 
Polyphaga
Emery, 1886
Polyphags
105
3138
11411
Families
Genera
Species
 
Taxom
 

Infraorder
 
Scarabaeiformia
Crowson, 1960
12
309
1436
Families
Genera
Species
 
Taxom
 

Superfamily
 
Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles
12
309
1436
Families
Genera
Species
 
Taxom
 
FamilyScarabaeidae
Latreille, 1802
Dung beetles, Flower beetles, Fruit chafers, Rain beetles
239
1155
Genera
Species
 
Taxom
 
SubfamilyScarabaeinae
Latreille, 1802
Dung Beetles, True dung beetles
25
164
Genera
Species
 
Taxom
 


Genera for Taiwan
25 Genera with 164 Species
#CDEGHKLOPST
Caccobius Thomson, 19389SpeciesTaxom
Cassolus Sharp, 19861SpeciesTaxom
Catharsius Hope, 17789SpeciesTaxom
Cheironitis van Lansberge, 18681SpeciesTaxom
Cleptocaccobius Cambefort, 19421SpeciesTaxom
Copris Geoffroy, 1857Mexican dung beetle14SpeciesTaxom
Drepanocerus Kirby, 19122SpeciesTaxom
Euoniticellus Janssens, 17811SpeciesTaxom
Gymnopleurus Illiger, 17854SpeciesTaxom
Haroldius Boucomont, 19392SpeciesTaxom
Kheper Janssens, 18671SpeciesTaxom
Liatongus Reitter, 17983SpeciesTaxom
Oniticellus Serville, 19342SpeciesTaxom
Onitis Fabricius, 19314SpeciesTaxom
Onthophagus Latreille, 201488SpeciesTaxom
Panelus Lewis, 20114SpeciesTaxom
Parachorius Harold, 19861SpeciesTaxom
Paragymnopleurus Shipp, 05SpeciesTaxom
Paraphytus Harold, 18951SpeciesTaxom
Phanaeus MacLeay, 1860Burrowing Dung Beetles1SpeciesTaxom
Proagoderus Lansberge, 18882SpeciesTaxom
Scarabaeus Linnaeus, 17896SpeciesTaxom
Sinodrepanus Simonis, 20042SpeciesTaxom
Synapsis Forster, 18754SpeciesTaxom
Tibiodrepanus Krikken, 18681SpeciesTaxom


Species list for Taiwan
164 Species and 7 Subspecies
Caccobius (Caccobius) chujoi Paulian, 1942
Caccobius (Caccophilus) scrofa (Balthasar, 1942)⇒ Cleptocaccobius scrofa Balthasar, 1942
Caccobius (Caccophilus) tortus Sharp, 1875
Caccobius tortus matsudai Ochi & Araya, 1996
Caccobius (Caccophilus) unicornis Fabricius, 1798
Caccobius demangei Boucomont, 1919
Caccobius flavolimbatus Balthasar, 1942
Caccobius gonoderus Fairmaire, 1888
Caccobius tsunoellus (Matsumara, 1938)⇒ Caccophilus tortus Sharp, 1875
Cassolus gotoi Masumoto, 1986
Catharsius borneensis (Paulian, 1936)⇒ Catharsius molossus Linnaeus, 1758
Catharsius dubius (Péringuey, 1901)⇒ Catharsius molossus Linnaeus, 1758
Catharsius javanus Lansberge, 1886
Catharsius kangeanus (Paulian, 1936)⇒ Catharsius molossus Linnaeus, 1758
Catharsius molossus Linnaeus, 1758
Catharsius pithecius Fabricius, 1775
Catharsius renaudpauliani Ochi & Kon, 1996
Catharsius timorensis (Lansberge, 1879)⇒ Catharsius molossus Linnaeus, 1758
Catharsius tricornutus DeGeer, 1778
Cheironitis hoplosternus Harold, 1868
Cleptocaccobius scrofa Balthasar, 1942
Copris (Copris) fukiensis Balthasar, 1952
Copris (Copris) punctatus Gillet, 1910
Copris punctatus formosanus Gillet, 1922
Copris (Copris) tripartitus Waterhouse, 1875
Copris (Microcopris) propinquus Felsche, 1910
Copris (Paracopris) cariniceps Felsche, 1910
Copris (Sinocopris) pecuarius Lewis, 1884
Copris acutidens Motschulsky, 1860
Copris confucius Harold, 1877
Copris minutus Drury, 1773Small Black Dung Beetle
Copris ochus Motschulsky, 1860
Copris reflexus (Fabricius, 1787)⇒ Copris minutus Drury, 1773
Copris sinicus Hope, 1842
Copris urus (Boheman, 1857)⇒ Catharsius molossus Linnaeus, 1758
Drepanocerus arrowi (Bathasar, 1932)⇒ Sinodrepanus rex Boucomont, 1912
Drepanocerus rex (Boucomont, 1912)⇒ Sinodrepanus rex Boucomont, 1912
Euoniticellus pallipes Fabricius, 1781Southern sandy dung beetle
Gymnopleurus aethiops (Sharp, 1875)⇒ Paragymnopleurus melanarius Harold, 1867
Gymnopleurus geoffroyi Fuessly, 1775
Gymnopleurus leei Fabricius, 1792
Gymnopleurus sinuatus (Geoffroy, 1785)⇒ Gymnopleurus geoffroyi Fuessly, 1775
Haroldius hwangi Masumoto, Le..., 2005
Haroldius perroti Paulian, 1939
Kheper erichsoni Harold, 1867
Liatongus minutus Motschulsky, 1860
Liatongus phanaeoides Westwood, 1839
Liatongus vertagus Fabricius, 1798
Oniticellus cinctus Fabricius, 1775Bordered dung beetle
Oniticellus yohenai (Matsumura, 1934)⇒ Liatongus phanaeoides Westwood, 1839
Onitis crenatus Reiche, 1847
Onitis falcatus Wulfen, 1786
Onitis philemon (Fabricius, 1801)⇒ Onitis subopacus Arrow, 1931
Onitis subopacus Arrow, 1931
Onthophagus (Colobonthophagus) quadridentatus (Fabricius, 1798)⇒ Onthophagus fissicornis Stevens, 1809
Onthophagus (Colobonthophagus) tragus Fabricius, 1792
Onthophagus (Gibbonthophagus) kentingensis Nomura, 1973
Onthophagus (Gibbonthophagus) luridipennis Boheman, 1858
Onthophagus (Gibbonthophagus) taurinus White, 1844
Onthophagus (Gibbonthophagus) viduus (Harold, 1875)⇒ Onthophagus orbus Boucomont, 1924
Onthophagus (Indachorius) hsui Masumoto, Ch..., 2004
Onthophagus (Indachorius) koshunensis Balthasar, 1941
Onthophagus (Matashia) anguliceps (Boucomont, 1914)⇒ Onthophagus avocetta Arrow, 1933
Onthophagus (Matashia) yubarinus (Matsumura, 1937)⇒ Onthophagus fodiens Waterhouse, 1875
Onthophagus (Micronthophagus) konoi Matsumura, 1938
Onthophagus (Micronthophagus) wangi Masumoto, Ch..., 2004
Onthophagus (Onthophagiellus) taiyaruensis Masumoto, 1977
Onthophagus (Onthophagus) acuticollis (Gillet, 1927)⇒ Onthophagus amphioxus Arrow, 1933
Onthophagus (Onthophagus) anguicorius Boucomont, 1914
Onthophagus (Onthophagus) dapcauensis Boucomont, 1921
Onthophagus (Onthophagus) dubernardi Boucomont, 1914
Onthophagus (Onthophagus) fissicornis Stevens, 1809
Onthophagus (Onthophagus) formosanus Gillet, 1924
Onthophagus (Onthophagus) hajimei Masumoto, 1984
Onthophagus (Onthophagus) hastifer Lansberge, 1885
Onthophagus (Onthophagus) hayashii Masumoto, 1991
Onthophagus (Onthophagus) kuatunensis Balthasar, 1942
Onthophagus (Onthophagus) lutosopictus Fairmaire, 1897
Onthophagus (Onthophagus) masumotoi Ochi, 1985
Onthophagus (Onthophagus) mendicus Gillet, 1924
Onthophagus (Onthophagus) orientalis (Harold, 1868)⇒ Onthophagus fissinasus Fairmaire, 1895
Onthophagus (Onthophagus) proletarius Harold, 1875
Onthophagus (Onthophagus) pseudoarmatus Balthasar, 1944
Onthophagus (Onthophagus) roubali Balthasar, 1935
Onthophagus (Onthophagus) rugulosus Harold, 1886
Onthophagus (Onthophagus) sakainoi Masumoto, 1991
Onthophagus (Onthophagus) sauteri Gillet, 1924
Onthophagus (Onthophagus) setchan Masumoto, 1984
Onthophagus (Onthophagus) sobrius Balthasar, 1959
Onthophagus (Onthophagus) striatulus Palisot de B..., 1809Surf blenny
Onthophagus (Onthophagus) tatsienluensis Balthasar, 1942
Onthophagus (Onthophagus) terminatus Eschscholtz, 1822
Onthophagus (Onthophagus) trituber Wiedemann, 1823
Onthophagus (Onthophagus) yangi Masumoto, Ts..., 2006
Onthophagus (Palaeonthophagus) fissicornis Steven, 1809
Onthophagus gibbulus gibbulus
Onthophagus (Palaeonthophagus) vacca Linnaeus, 1767
Onthophagus (Paraphanaeomorphus) argyropygus Gillet, 1927
Onthophagus (Parascatonomus) kiuchianus Masumoto, Ya..., 2004
Onthophagus (Parascatonomus) miyakei Ochi & Araya, 1992
Onthophagus (Parascatonomus) tricornis (Wiedemann, 1823)⇒ Onthophagus vacca Linnaeus, 1767
Onthophagus (Phanaeomorphus) ater Waterhouse, 1875
Onthophagus (Phanaeomorphus) cernyi (Balthasar, 1935)⇒ Onthophagus ater Waterhouse, 1875
Onthophagus (Phanaeomorphus) lenzi Harold, 1875
Onthophagus potanini hiurai Ochi, 1984
Onthophagus (Serrophorus) rectecornutus Lansberge, 1883
Onthophagus (Serrophorus) sagittarius Fabricius, 1775Sagittarius dung beetle
Onthophagus (Strandius) japonicus Harold, 1875
Onthophagus (Strandius) kuraruanus (Matsumura, 1938)⇒ Onthophagus lenzi Harold, 1875
Onthophagus (Strandius) potanini R. C. Brusca..., 1990
Onthophagus potanini hiurai Ochi, 1984
Onthophagus (Strandius) taiwanus Nomura, 1973
Onthophagus agilis (Matsumura, 1938)⇒ Onthophagus hastifer Lansberge, 1885
Onthophagus amphioxus Arrow, 1933
Onthophagus armatus Blanchard, 1853
Onthophagus avocetta Arrow, 1933
Onthophagus campestris Masumoto, Ki..., 2018
Onthophagus capitosus Harold, 1867
Onthophagus chokakurianus Matsumura, 1938
Onthophagus egurianus Matsumura, 1938
Onthophagus euplynes Bates, 1888
Onthophagus excavatus (Redtenbacher, 1848)⇒ Liatongus phanaeoides Westwood, 1839
Onthophagus fodiens Waterhouse, 1875
Onthophagus gagates Hope, 1831
Onthophagus jiupengensis Masumoto, La..., 2017
Onthophagus liui Masumoto, Oc..., 2014
Onthophagus lunatus Harold, 1868
Onthophagus midorianus (Matsumura, 1938)⇒ Parascatonomus anguicorius Boucomont, 1914
Onthophagus mushanus Matsumura, 1938
Onthophagus mushensis Matsumura, 1938
Onthophagus nagasawai Matsumura, 1938
Onthophagus nitidus Waterhouse, 1875
Onthophagus ohbayashii Nomura, 1939
Onthophagus orbus Boucomont, 1924
Onthophagus phanaeoides (Westwood, 1839)⇒ Proagoderus lanista Castelnau, 1840
Onthophagus putealis (Matsumura, 1938)⇒ Onthophagus hastifer Lansberge, 1885
Onthophagus senescens Péringuey, 1908
Onthophagus seniculus (Fabricius, 1781)⇒ Onthophagus senescens Péringuey, 1908
Onthophagus sonani Miwa, 1930
Onthophagus spitsbergeniensis Krell, 2010
Onthophagus yanoi Matsumura, 1938
Onthophagus yaoi Masumoto, Oc..., 2014
Panelus crenatus Nomura, 1973
Panelus maedai Nomura, 1973
Panelus manmiaoae Masumoto, Oc..., 2011
Panelus wangi Masumoto, Oc..., 2011
Parachorius gotoi Masumoto, 1986
Paragymnopleurus ambiguus Janssens, 1945
Paragymnopleurus melanarius Harold, 1867
Paragymnopleurus sinuatus Olivier, 1789
Paragymnopleurus sinuatus productus Sharp, 1875
Paragymnopleurus sinuatus sinuatus
Paraphytus dentifrons Lewis, 1895
Phanaeus minutus (Motschulsky, 1860)⇒ Liatongus phanaeoides Westwood, 1839
Proagoderus lanista Castelnau, 1840
Proagoderus panoplus Bates, 1888
Scarabaeus abbreviatus (Herbst, 1789)⇒ Catharsius molossus Linnaeus, 1758
Scarabaeus berbiceus (Herbst, 1789)⇒ Catharsius molossus Linnaeus, 1758
Scarabaeus janus (Olivier, 1789)⇒ Catharsius molossus Linnaeus, 1758
Scarabaeus leei (Donovan, 1798)⇒ Gymnopleurus leei Fabricius, 1792
Scarabaeus molossus (Linnaeus, 1758)⇒ Catharsius molossus Linnaeus, 1758
Scarabaeus sinuatus (Olivier, 1789)⇒ Paragymnopleurus sinuatus Olivier, 1789
Sinodrepanus rex Boucomont, 1912
Sinodrepanus tsaii Masumoto, Ya..., 2004
Synapsis davidi (Fairmaire, 1878)⇒ Synapsis davidis Fairmaire, 1878
Synapsis davidis Fairmaire, 1878
Synapsis masumotoi Ochi, 1992
Synapsis simplex Sharp, 1875
Tibiodrepanus sinicus Harold, 1868

Subnodes of Scarabaeinae with occurrences in Taiwan
TribeAteuchini
Montreuil, 1998
1
1
Genus
Species

Taxom

TribeCanthonini
Péringuey, 1900
2
5
Genera
Species

Taxom

TribeCoprini
Kolbe, 1805
3
26
Genera
Species

Taxom

TribeGymnopleurini
Lacordaire, 1856
2
7
Genera
Species

Taxom

TribeOniticellini
d´Orbigny, 1916
6
11
Genera
Species

Taxom

TribeOnitini
Hope, 1837
2
5
Genera
Species

Taxom

TribeOnthophagini
Lacordaire, 1856
6
101
Genera
Species

Taxom

TribePhanaeini
MacLeay, 1819
1
1
Genus
Species

Taxom

TribeScarabaeini
Latreille, 1802
2
7
Genera
Species

Taxom



Links, sources and further information
Scarabaeinae at artfakta.se
Scarabaeinae at dyntaxa.se
Scarabaeinae at itis.gov