Checklist for Subfamily Scarabaeinae Latreille, 1802 (Dung Beetles)
and region Pakistan (South Asia)
Class
| Insecta
Linnaeus, 1758
Insects, True insects | 341
2930
6223 | Families
Genera
Species |
Taxom
|
Subclass
| Pterygota
Lang, 1888
Winged insects | 341
2930
6223 | Families
Genera
Species |
Taxom
|
Infraclass
| Neoptera
van der Wulp, 1890
Modern Wing-folding insects | 331
2906
6190 | Families
Genera
Species |
Taxom
|
Superorder
| Holometabola
Heider, 1889
Holometabolous Insects | 221
2259
4909 | Families
Genera
Species |
Taxom
|
Order
| Coleoptera
Linnaeus, 1758
Beetles, Coleopterans | 50
720
1882 | Families
Genera
Species |
Taxom
|
Suborder
| Polyphaga
Emery, 1886
Polyphags | 48
551
1298 | Families
Genera
Species |
Taxom
|
Infraorder
| Scarabaeiformia
Crowson, 1960
| 11
142
401 | Families
Genera
Species |
Taxom
|
Superfamily
| Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles | 11
142
401 | Families
Genera
Species |
Taxom
| | Family | Scarabaeidae
Latreille, 1802
Dung beetles, Flower beetles, Fruit chafers, Rain beetles | 112
339 | Genera
Species |
Taxom
| | Subfamily | Scarabaeinae
Latreille, 1802
Dung Beetles, True dung beetles | 24
102 | Genera
Species |
Taxom
|
Genera for Pakistan
24 Genera with 102 Species #CDEGHLMOPST
| Caccobius Thomson, 1931 | | 3 | Species | Taxom | | Catharsius Hope, 1806 | | 5 | Species | Taxom | | Cheironitis van Lansberge, 1856 | | 3 | Species | Taxom | | Cleptocaccobius Cambefort, 1979 | | 1 | Species | Taxom | | Copris Geoffroy, 1848 | Mexican dung beetle | 7 | Species | Taxom | | Digitonthophagus Balthasar, 2017 | | 4 | Species | Taxom | | Euoniticellus Janssens, 1781 | | 2 | Species | Taxom | | Euonthophagus Balthasar, 1892 | | 4 | Species | Taxom | | Garreta Janssens, 1819 | | 2 | Species | Taxom | | Gymnopleurus Illiger, 1931 | | 6 | Species | Taxom | | Haroldius Boucomont, 1918 | | 1 | Species | Taxom | | Heliocopris Hope, 1775 | | 3 | Species | Taxom | | Liatongus Reitter, 1839 | | 1 | Species | Taxom | | Metacatharsius Paulian, 1840 | | 1 | Species | Taxom | | Oniticellus Serville, 1775 | | 1 | Species | Taxom | | Onitis Fabricius, 1875 | | 11 | Species | Taxom | | Onthophagus Latreille, 1798 | | 33 | Species | Taxom | | Paragymnopleurus Shipp, 1890 | | 1 | Species | Taxom | | Phalops Erichson, 1883 | | 3 | Species | Taxom | | Proagoderus Lansberge, 1875 | | 1 | Species | Taxom | | Scarabaeus Linnaeus, 1840 | | 5 | Species | Taxom | | Sisyphus Latreille, 1833 | | 3 | Species | Taxom | | Tibiodrepanus Krikken, 1868 | | 2 | Species | Taxom | | Tiniocellus Péringuey, 1851 | | 2 | Species | Taxom |
Species list for Pakistan
102 Species and 6 Subspecies | Caccobius balthasari (Tesar, 1969) | ⇒ Cleptocaccobius balthasarianus Cambefort, 1979 | | Caccobius denticollis Harold, 1867 | | | Caccobius pantherinus Arrow, 1931 | | | Catharsius granulatus Sharp, 1875 | | | Catharsius molossus Linnaeus, 1758 | | | Catharsius pithecius Fabricius, 1775 | | | Catharsius platypus Sharp, 1875 | | | Catharsius sagax Quenstedt, 1806 | | | Cheironitis arrowi Janssens, 1937 | | | Cheironitis indicus (Lansberge, 1875) | ⇒ Cheironitis arrowi Janssens, 1937 | | Cheironitis osiridis Reiche, 1856 | | | Cleptocaccobius balthasarianus Cambefort, 1979 | | | Copris excisus Waterhouse, 1891 | | | Copris fricator Fabricius, 1787 | | | Copris fricator fricator | | | Copris hispanus Linnaeus, 1764 | | | Copris hispanus cavolinii Petagna, 1792 | | | Copris punjabensis Gillet, 1921 | | | Copris sacontala Redtenbacher, 1848 | | | Digitonthophagus bonasus Fabricius, 1775 | | | Digitonthophagus catta Fabricius, 1787 | Beetle | | Digitonthophagus gazella Fabricius, 1787 | Brown dung beetle | | Digitonthophagus uks Génier, 2017 | | | Euoniticellus pallens Olivier, 1789 | | | Euoniticellus pallipes Fabricius, 1781 | Southern sandy dung beetle | | Euonthophagus laghmanicus Kabakov, 1977 | | | Euonthophagus pertinax Balthasar, 1959 | | | Euonthophagus schnabeli Splichal, 1910 | | | Euonthophagus sulcicollis Reitter, 1892 | | | Garreta dejeani Castelnau, 1840 | | | Garreta mundus Wiedemann, 1819 | | | Gymnopleurus bicallosus Felsche, 1909 | | | Gymnopleurus cyaneus (Fabricius, 1798) | ⇒ Gymnopleurus sericeifrons Fairmaire, 1887 | | Gymnopleurus flagellatus flagellatus Fabricius, 1787 | | | Gymnopleurus flagellatus hornei Waterhouse, 1890 | | | Gymnopleurus sericeifrons Fairmaire, 1887 | | | Gymnopleurus sindensis Arrow, 1931 | | | Haroldius heimi Wasmann, 1918 | | | Heliocopris bucephalus Fabricius, 1775 | | | Heliocopris gigas Linnaeus, 1758 | | | Heliocopris midas Fabricius, 1775 | | | Liatongus phanaeoides Westwood, 1839 | | | Metacatharsius inermis Castelnau, 1840 | | | Oniticellus cinctus Fabricius, 1775 | Bordered dung beetle | | Onitis castaneus Redtenbacher, 1848 | | | Onitis excavatus Arrow, 1931 | | | Onitis falcatus Wulfen, 1786 | | | Onitis humerosus Pallas, 1771 | | | Onitis lama Lansberge, 1875 | | | Onitis niger (Lansberge, 1875) | ⇒ Liatongus femoratus Illiger, 1800 | | Onitis philemon (Fabricius, 1801) | ⇒ Onitis subopacus Arrow, 1931 | | Onitis punctatostriatus Janssens, 1937 | | | Onitis singhalensis Lansberge, 1875 | | | Onitis subopacus Arrow, 1931 | | | Onitis virens Lansberge, 1875 | | | Onthophagus (Colobonthophagus) aenescens Wiedemann, 1823 | | | Onthophagus (Colobonthophagus) paliceps Arrow, 1931 | | | Onthophagus (Colobonthophagus) quadridentatus (Fabricius, 1798) | ⇒ Onthophagus fissicornis Stevens, 1809 | | Onthophagus (Onthophagus) fissicornis Stevens, 1809 | | | Onthophagus (Palaeonthophagus) fissicornis Steven, 1809 | | | Onthophagus (Trichonthophagus) tarandus Fabricius, 1792 | | | Onthophagus amphicoma Boucomont, 1914 | | | Onthophagus asperrimus d´Orbigny, 1902 | | | Onthophagus bicolor (Raffray, 1877) | ⇒ Onthophagus flavolimbatus Klug, 1855 | | Onthophagus capitosus Harold, 1867 | | | Onthophagus concolor Sharp, 1878 | | | Onthophagus cyaneus Kabakov, 2006 | | | Onthophagus ensifer Boucomont, 1914 | | | Onthophagus falsus Gillet, 1925 | | | Onthophagus filicornis Harold, 1873 | | | Onthophagus flavolimbatus Klug, 1855 | | | Onthophagus frontalis Raffray, 1877 | | | Onthophagus hindu Arrow, 1931 | | | Onthophagus incertus (d´Orbigny, 1897) | ⇒ Onthophagus asperrimus d´Orbigny, 1902 | | Onthophagus lahorensis Kabakov, 2008 | | | Onthophagus mopsus Fabricius, 1792 | | | Onthophagus nitidulus Klug, 1845 | | | Onthophagus ochreatus d´Orbigny, 1897 | | | Onthophagus parvulus (Fabricius, 1798) | ⇒ Onthophagus frontalis Raffray, 1877 | | Onthophagus piffli Petrovitz, 1961 | | | Onthophagus ramosellus Bates, 1891 | | | Onthophagus ramosus Wiedemann, 1823 | | | Onthophagus semicinctus d´Orbigny, 1897 | | | Onthophagus spinifex Fabricius, 1781 | | | Onthophagus sutleinensis Splichal, 1910 | | | Onthophagus transcaspicus Koenig, 1888 | Gazella scarab | | Onthophagus troglodyta Wiedemann, 1823 | | | Onthophagus variegatus (Fabricius, 1798) | ⇒ Onthophagus filicornis Harold, 1873 | | Paragymnopleurus sinuatus assamensis Waterhouse, 1890 | | | Phalops cyanescens d´Orbigny, 1897 | | | Phalops divisus Wiedemann, 1823 | | | Phalops olivaceus Lansberge, 1883 | | | Proagoderus amplexus Sharp, 1875 | | | Scarabaeus (Scarabaeus) babori Balthasar, 1934 | | | Scarabaeus (Scarabaeus) sacer (Linnaeus, 1758) | ⇒ Scarabaeus pius Illiger, 1803 | | Scarabaeus (Scarabaeus) winkleri Stolfa, 1938 | | | Scarabaeus acuticollis Motschulsky, 1849 | | | Scarabaeus gangeticus Castelnau, 1840 | | | Sisyphus crispatus hirtus Wiedemann, 1823 | | | Sisyphus longipes Olivier, 1789 | | | Sisyphus neglectus Gory, 1833 | | | Tibiodrepanus setosus Wiedemann, 1823 | | | Tibiodrepanus sinicus Harold, 1868 | | | Tiniocellus imbellis Bates, 1891 | | | Tiniocellus spinipes Roth, 1851 | |
Subnodes of Scarabaeinae with occurrences in Pakistan
Links, sources and further information Scarabaeinae at artfakta.se
Scarabaeinae at dyntaxa.se
Scarabaeinae at itis.gov
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