Checklist for Subfamily Scarabaeinae Latreille, 1802 (Dung Beetles)
and region Paraguay (Northern South America)
Class
| Insecta
Linnaeus, 1758
Insects, True insects | 403
4047
8334 | Families
Genera
Species |
Taxom
|
Subclass
| Pterygota
Lang, 1888
Winged insects | 402
4046
8333 | Families
Genera
Species |
Taxom
|
Infraclass
| Neoptera
van der Wulp, 1890
Modern Wing-folding insects | 385
3986
8213 | Families
Genera
Species |
Taxom
|
Superorder
| Holometabola
Heider, 1889
Holometabolous Insects | 250
3207
6803 | Families
Genera
Species |
Taxom
|
Order
| Coleoptera
Linnaeus, 1758
Beetles, Coleopterans | 77
1517
3212 | Families
Genera
Species |
Taxom
|
Suborder
| Polyphaga
Emery, 1886
Polyphags | 72
1377
2875 | Families
Genera
Species |
Taxom
|
Infraorder
| Scarabaeiformia
Crowson, 1960
| 10
193
670 | Families
Genera
Species |
Taxom
|
Superfamily
| Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles | 10
193
670 | Families
Genera
Species |
Taxom
| | Family | Scarabaeidae
Latreille, 1802
Dung beetles, Flower beetles, Fruit chafers, Rain beetles | 151
545 | Genera
Species |
Taxom
| | Subfamily | Scarabaeinae
Latreille, 1802
Dung Beetles, True dung beetles | 41
210 | Genera
Species |
Taxom
|
Genera for Paraguay
41 Genera with 210 Species #ABCDEGHMOPSTUZ
| Agamopus Bates, 1928 | | 1 | Species | Taxom | | Anisocanthon Martinez & P..., 1868 | | 1 | Species | Taxom | | Anomiopus Westwood, 1842 | | 7 | Species | Taxom | | Ateuchus Weber, 1935 | | 7 | Species | Taxom | | Besourenga Vaz-de-Mello, 1935 | | 3 | Species | Taxom | | Bolbites Harold, 1868 | | 1 | Species | Taxom | | Canthidium Erichson, 1939 | | 12 | Species | Taxom | | Canthon Hoffmannsegg, 1939 | Tumblebug | 41 | Species | Taxom | | Chalcocopris Burmeister, 2015 | | 2 | Species | Taxom | | Copris Geoffroy, 1959 | Mexican dung beetle | 1 | Species | Taxom | | Coprophanaeus Olsoufieff, 1847 | | 14 | Species | Taxom | | Deltochilum Eschscholtz, 1939 | | 19 | Species | Taxom | | Dendropaemon Perty, 1831 | | 12 | Species | Taxom | | Diabroctis Gistl, 1877 | | 2 | Species | Taxom | | Dichotomius Hope, 1869 | | 22 | Species | Taxom | | Digitonthophagus Balthasar, 1787 | | 1 | Species | Taxom | | Eudinopus Burmeister, 1802 | | 1 | Species | Taxom | | Eurysternus Dalman, 1988 | | 10 | Species | Taxom | | Eutrichillum | | 1 | Species | Taxom | | Genieridium Vaz-de-Mello, 1913 | | 2 | Species | Taxom | | Gromphas Brullé, 1837 | | 5 | Species | Taxom | | Holocanthon | | 1 | Species | Taxom | | Holocephalus Hope, 2016 | | 2 | Species | Taxom | | Malagoniella Martínez, 0 | | 5 | Species | Taxom | | Ontherus Erichson, 1775 | | 8 | Species | Taxom | | Onthocharis Westwood, 1955 | | 1 | Species | Taxom | | Onthophagus Latreille, 1859 | | 6 | Species | Taxom | | Oxysternon Laporte, 1840 | | 2 | Species | Taxom | | Pedaridium Harold, 1951 | | 1 | Species | Taxom | | Phanaeus MacLeay, 1846 | Burrowing Dung Beetles | 6 | Species | Taxom | | Pinotus Erichson, 1929 | | 1 | Species | Taxom | | Pseudocanthon Bates, 1846 | | 3 | Species | Taxom | | Scatonomus Erichson, 1959 | | 2 | Species | Taxom | | Scybalocanthon Martínez, 1855 | | 3 | Species | Taxom | | Sulcophanaeus Olsoufieff, 1840 | | 4 | Species | Taxom | | Sylvicanthon Halffter & M..., 1920 | | 1 | Species | Taxom | | Tetraechma Agassiz, 1939 | | 3 | Species | Taxom | | Trichillidium Vaz-de-Mello, 1932 | | 1 | Species | Taxom | | Trichillum Harold, 2005 | | 7 | Species | Taxom | | Uroxys Westwood, 1940 | | 3 | Species | Taxom | | Zonocopris Arrow, 2007 | | 2 | Species | Taxom |
Species list for Paraguay
210 Species and 20 Subspecies | Agamopus viridis Boucomont, 1928 | | | Anisocanthon villosus Harold, 1868 | | | Anomiopus birai Canhedo, 2006 | | | Anomiopus germari Harold, 1867 | | | Anomiopus nigrocoeruleus Martínez, 1955 | | | Anomiopus paraguaiensis Canhedo, 2004 | | | Anomiopus pereirai Martínez, 1955 | | | Anomiopus tuberifrons Canhedo, 2004 | | | Anomiopus virescens Westwood, 1842 | | | Ateuchus apicatus Harold, 1867 | | | Ateuchus balthasari Martínez, 1952 | | | Ateuchus breve Harold, 1868 | | | Ateuchus contractus Balthasar, 1939 | | | Ateuchus ovale Boucomont, 1928 | | | Ateuchus ovalis Boucomont, 1928 | | | Ateuchus viridimicans Boucomont, 1935 | | | Besourenga amarillai Aguilar-Julio, 2001 | | | Besourenga minutum Saylor, 1935 | | | Besourenga minutus Saylor, 1935 | | | Bolbites onitoides Harold, 1868 | | | Canthidium barbacenicum Preudhomme d..., 1886 | | | Canthidium bokermanni Martinez, Ha..., 1964 | | | Canthidium breve Germar, 1824 | | | Canthidium cupreum Blanchard, 1846 | | | Canthidium dispar Harold, 1867 | | | Canthidium femoratum Boucomont, 1935 | | | Canthidium gigas Balthasar, 1939 | | | Canthidium kelleri Martinez, Ha..., 1964 | | | Canthidium multipunctatum Balthasar, 1939 | | | Canthidium pinotoides (Balthasar, 1939) | ⇒ Canthidium barbacenicum Preudhomme d..., 1886 | | Canthidium prasinum Blanchard, 1846 | | | Canthidium titschacki Balthasar, 1939 | | | Canthon angularis Harold, 1868 | | | Canthon angularis angularis | | | Canthon arriagadai Vieira, Vaz-..., 2019 | | | Canthon cobosi Pereira & Ma..., 1960 | | | Canthon curvodilatatus Schmidt, 1920 | | | Canthon denticulatus Schmidt, 1922 | | | Canthon edentulus Harold, 1868 | | | Canthon forcipatus Harold, 1868 | | | Canthon hendrichsi Halffter & M..., 1968 | | | Canthon ibarragrassoi Martínez, 1952 | | | Canthon laminatus Balthasar, 1939 | | | Canthon lituratus Germar, 1813 | | | Canthon lividus Blanchard, 1846 | | | Canthon lividus lividus | | | Canthon lividus seminitens Harold, 1868 | | | Canthon luctuosus Harold, 1868 | | | Canthon mutabilis Lucas, 1857 | | | Canthon muticus Harold, 1867 | | | Canthon octodentatus Schmidt, 1920 | | | Canthon oliverioi Pereira & Ma..., 1956 | | | Canthon ornatus Redtenbacher, 1867 | | | Canthon ornatus ornatus | | | Canthon podagricus Harold, 1868 | | | Canthon quadratus Blanchard, 1846 | | | Canthon quinquemaculatus Castelnau, 1840 | | | Canthon rutilans (Castelnau, 1840) | ⇒ Francmonrosia rutilans Castelnau, 1840 | | Canthon rutilans cyanescens Harold, 1868 | | | Canthon rutilans rutilans | | | Canthon septemmaculatus Latreille, 1812 | | | Canthon septemmaculatus septemmaculatus | | | Canthon smaragdulus Fabricius, 1781 | | | Canthon smaragdulus smaragdulus | | | Canthon staigi Pereira, 1953 | | | Canthon substriatus Harold, 1868 | | | Canthon triangularis Drury, 1770 | | | Canthon unicolor Blanchard, 1846 | | | Canthon unicolor fortemarginatus Balthasar, 1939 | | | Canthon unicolor unicolor | | | Canthon velutinus Harold, 1868 | | | Canthon virens Mannerheim, 1829 | | | Canthon virens paraguayanus Balthasar, 1939 | | | Chalcocopris hesperus Olivier, 1789 | | | Chalcocopris inexpectatus Rossini & Va..., 2015 | | | Copris megasoma Matthews & H..., 1959 | | | Coprophanaeus (Megaphanaeus) bonariensis Gory, 1844 | | | Coprophanaeus (Megaphanaeus) ensifer Germar, 1821 | | | Coprophanaeus (Megaphanaeus) lancifer Linnaeus, 1767 | | | Coprophanaeus (Metallophanaeus) horus Waterhouse, 1891 | | | Coprophanaeus (Metallophanaeus) pessoai Pereira, 1949 | | | Coprophanaeus (Metallophanaeus) saphirinus Sturm, 1826 | | | Coprophanaeus saphirinus saphirinus | | | Coprophanaeus (Metallophanaeus) thalassinus Perty, 1830 | | | Coprophanaeus cyanescens Olsufieff, 1924 | | | Coprophanaeus jasius Olivier, 1789 | | | Coprophanaeus milon Blanchard, 1846 | | | Coprophanaeus rigoutorum Arnaud, 2002 | | | Coprophanaeus spitzi Pessôa, 1934 | | | Coprophanaeus telamon Erichson, 1847 | | | Deltochilum amandaarcanjoae Silva, Louza..., 2015 | | | Deltochilum aspericolle Bates, 1870 | | | Deltochilum brasiliense Castelnau, 1840 | | | Deltochilum cristatum Paulian, 1938 | | | Deltochilum cupreicolle Blanchard, 1846 | | | Deltochilum dentipes Eschscholtz, 1822 | | | Deltochilum elongatum Felsche, 1907 | | | Deltochilum icariforme Paulian, 1938 | | | Deltochilum icaroides Balthasar, 1939 | | | Deltochilum icarus Olivier, 1789 | | | Deltochilum irroratum Castelnau, 1840 | | | Deltochilum kolbei Paulian, 1938 | | | Deltochilum komareki Balthasar, 1939 | | | Deltochilum pseudoicarus Balthasar, 1939 | | | Deltochilum rubripenne Gory, 1831 | | | Deltochilum valgum Burmeister, 1873 | | | Deltochilum variolosum Burmeister, 1873 | | | Deltochilum viridescens Martinez, 1948 | | | Deltochilum viridicupreum Balthasar, 1939 | | | Dendropaemon (Coprophanaeoides) renatii Olsoufieff, 1924 | | | Dendropaemon (Glaphyropaemon) bahianus (Harold, 1868) | ⇒ Coprophanaeoides bahianus Harold, 1868 | | Dendropaemon (Rutilopaemon) refulgens (Waterhouse, 1891) | ⇒ Coprophanaeoides refulgens Waterhouse, 1891 | | Dendropaemon (Sulcopaemon) fasces Blut, 1939 | | | Dendropaemon (Sulcopaemon) latistriatus Génier & Arn..., 2016 | | | Dendropaemon (Titthopaemon) denticollis Felsche, 1909 | | | Dendropaemon batrachites Blut, 1939 | | | Dendropaemon pilosissimus Génier & Arn..., 2016 | | | Dendropaemon quadratus Laporte, 1832 | | | Dendropaemon similis Blut, 1939 | | | Dendropaemon tenuitarsis Felsche, 1909 | | | Dendropaemon viridipennis Laporte, 1831 | | | Diabroctis mimas Linnaeus, 1758 | | | Diabroctis mirabilis Harold, 1877 | | | Dichotomius (Luederwaldtinia) bosqui Pereira, 1941 | | | Dichotomius (Luederwaldtinia) carbonarius Mannerheim, 1829 | | | Dichotomius (Luederwaldtinia) crinicollis Germar, 1824 | | | Dichotomius (Luederwaldtinia) glaucus Harold, 1869 | | | Dichotomius (Luederwaldtinia) micans Luederwaldt, 1924 | | | Dichotomius (Luederwaldtinia) nisus Olivier, 1789 | | | Dichotomius (Luederwaldtinia) nobilis Waterhouse, 1891 | | | Dichotomius (Luederwaldtinia) opacipennis Luederwaldt, 1931 | | | Dichotomius (Luederwaldtinia) punctatostriatus (Felsche, 1901) | ⇒ Luederwaldtinia punctatostriatus Felsche, 1901 | | Dichotomius (Luederwaldtinia) sericeus Harold, 1867 | | | Dichotomius (Selenocopris) ascanius Harold, 1869 | | | Dichotomius (Selenocopris) bicuspis Germar, 1824 | | | Dichotomius (Selenocopris) guaranii Gandini & Ag..., 2009 | | | Dichotomius (Selenocopris) inflaticollis Felsche, 1910 | | | Dichotomius anthrax Felsche, 1901 | | | Dichotomius bos Blanchard, 1846 | | | Dichotomius depressicollis Harold, 1867 | | | Dichotomius fornicatus Luederwaldt, 1931 | | | Dichotomius joelus Gandini & Ag..., 2009 | | | Dichotomius mormon Ljungh, 1799 | | | Dichotomius nitidissimus Waterhouse, 1891 | | | Dichotomius reichei Harold, 1869 | | | Digitonthophagus gazella Fabricius, 1787 | Brown dung beetle | | Eudinopus dytiscoides Schreibers, 1802 | | | Eurysternus aeneus Génier, 2009 | | | Eurysternus caribaeus Herbst, 1789 | | | Eurysternus cyanescens Balthasar, 1939 | | | Eurysternus foedus Guérin-Ménev..., 1844 | | | Eurysternus hirtellus Dalman, 1824 | | | Eurysternus jessopi Martínez, 1988 | | | Eurysternus navajasi Martínez, 1988 | | | Eurysternus nigrovirens Génier, 2009 | | | Eurysternus parallelus Castelnau, 1840 | | | Eurysternus rivaldii Martínez, 1988 | | | Eutrichillum hirsutum Boucomont, 1928 | | | Genieridium bidens Balthasar, 1938 | | | Genieridium cryptops Arrow, 1913 | | | Gromphas aeruginosa Perty, 1830 | | | Gromphas amazonica Bates, 1870 | | | Gromphas dichroa Blanchard, 1846 | | | Gromphas inermis Harold, 1869 | | | Gromphas lacordairei (Brullé, 1837) | ⇒ Gromphas inermis Harold, 1869 | | Holocanthon fuscorubrus Blanchard, 1845 | | | Holocephalus cristatus Gillet, 1907 | | | Holocephalus pokornyi Nunes & Vaz-..., 2016 | | | Malagoniella astyanax punctatostriata Blanchard, 1846 | | | Malagoniella magnifica Balthasar, 1939 | | | Malagoniella puncticollis Blanchard, 1846 | | | Malagoniella puncticollis aeneicollis Waterhouse, 1890 | | | Malagoniella puncticollis puncticollis | | | Ontherus androgynus Génier, 1996 | | | Ontherus aphodioides Burmeister, 1874 | | | Ontherus appendiculatus Mannerheim, 1829 | | | Ontherus azteca Harold, 1869 | | | Ontherus carinicollis Luederwaldt, 1930 | | | Ontherus digitatus Harold, 1868 | | | Ontherus erosioides Luederwaldt, 1930 | | | Ontherus sulcator Fabricius, 1775 | | | Onthocharis melancholica (Martínez, 1955) | ⇒ Anomiopus virescens Westwood, 1842 | | Onthophagus buculus Mannerheim, 1829 | | | Onthophagus carpophilus Pereira & Ha..., 1961 | | | Onthophagus chacoensis Rossini, Vaz..., 2018 | | | Onthophagus hirculus (Mannerheim, 1829) | ⇒ Onthophagus mutatus Harold, 1859 | | Onthophagus hircus Billberg, 1815 | | | Onthophagus mutatus Harold, 1859 | | | Oxysternon festivum festivum | | | Oxysternon palaemon Castelnau, 1840 | | | Pedaridium mansosotoi (Martínez, 1951) | ⇒ Genieridium cryptops Arrow, 1913 | | Phanaeus (Notiophanaeus) kirbyi Vigors, 1825 | | | Phanaeus kirbyi kirbyi | | | Phanaeus bogotensis (Kirsch, 1871) | ⇒ Phanaeus hermes Harold, 1868 | | Phanaeus hermes Harold, 1868 | | | Phanaeus menelas (Castelnau, 1840) | ⇒ Sulcophanaeus menelas Castelnau, 1840 | | Phanaeus palaeno Blanchard, 1846 | | | Pinotus transiens (Luederwaldt, 1929) | ⇒ Dichotomius bicuspis Germar, 1824 | | Pseudocanthon chaquensis Nazaré-Silva..., 2021 | | | Pseudocanthon pantanensis Nazaré-Silva..., 2021 | | | Pseudocanthon xanthurus Blanchard, 1846 | | | Scatonomus viridis viridis | | | Scatonomus viridis walzi Pereira & Ma..., 1959 | | | Scybalocanthon luctuosus Harold, 1868 | | | Scybalocanthon nigriceps Harold, 1868 | | | Scybalocanthon sexspilotus Guérin-Ménev..., 1855 | | | Sulcophanaeus faunus Fabricius, 1775 | | | Sulcophanaeus imperator Chevrolat, 1844 | | | Sulcophanaeus imperator imperator | | | Sulcophanaeus menelas Castelnau, 1840 | | | Sylvicanthon foveiventris Schmidt, 1920 | | | Tetraechma chlorophana Mannerheim, 1828 | | | Tetraechma liturata Germar, 1813 | | | Tetraechma tarsalis Balthasar, 1939 | | | Trichillidium quadridens Arrow, 1932 | | | Trichillum arrowi (Saylor, 1935) | ⇒ Trichillum pauliani Balthasar, 1939 | | Trichillum boucomonti (Saylor, 1935) | ⇒ Eutrichillum hirsutum Boucomont, 1928 | | Trichillum externepunctatum Preudhomme d..., 1880 | | | Trichillum heydeni Harold, 1868 | | | Trichillum minutum (Saylor, 1935) | ⇒ Besourenga minutum Saylor, 1935 | | Trichillum pauliani Balthasar, 1939 | | | Trichillum pseudoarrowi Vaz-de-Mello..., 2005 | | | Uroxys corporaali Balthasar, 1940 | | | Uroxys histeroides Arrow, 1933 | | | Uroxys thoracalis Balthasar, 1940 | | | Zonocopris gibbicollis Harold, 1868 | | | Zonocopris machadoi Vaz-de-Mello, 2007 | |
Subnodes of Scarabaeinae with occurrences in Paraguay
Links, sources and further information Scarabaeinae at artfakta.se
Scarabaeinae at dyntaxa.se
Scarabaeinae at itis.gov
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