Checklist of Family Scarabaeidae Latreille, 1802 (Dung beetles)
for Paraguay (Northern South America)
Class
| Insecta
Linnaeus, 1758
Insects, True insects | 403
4043
8341 | Families
Genera
Species |
Taxom
|
Subclass
| Pterygota
Lang, 1888
Winged insects | 402
4042
8340 | Families
Genera
Species |
Taxom
|
Infraclass
| Neoptera
van der Wulp, 1890
Modern Wing-folding insects | 385
3982
8220 | Families
Genera
Species |
Taxom
|
Superorder
| Holometabola
Heider, 1889
Holometabolous Insects | 250
3204
6812 | Families
Genera
Species |
Taxom
|
Order
| Coleoptera
Linnaeus, 1758
Beetles, Coleopterans | 77
1514
3205 | Families
Genera
Species |
Taxom
|
Suborder
| Polyphaga
Emery, 1886
Polyphags | 72
1374
2870 | Families
Genera
Species |
Taxom
|
Infraorder
| Scarabaeiformia
Crowson, 1960
| 10
193
670 | Families
Genera
Species |
Taxom
|
Superfamily
| Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles | 10
193
670 | Families
Genera
Species |
Taxom
| | Family | Scarabaeidae
Latreille, 1802
Dung beetles, Flower beetles, Fruit chafers, Rain beetles | 151
545 | Genera
Species |
Taxom
|
Genera for Paraguay
151 Genera with 545 Species
| Aceratus Prell, 1847 | | 1 | Species | Taxom | | Actinobolus Westwood, 1841 | | 1 | Species | Taxom | | Agamopus Bates, 1928 | | 1 | Species | Taxom | | Agaocephala Mannerheim, 1828 | | 1 | Species | Taxom | | Aidophus Balthasar, 1909 | | 1 | Species | Taxom | | Allorrhina Burmeister, 1829 | | 4 | Species | Taxom | | Anahi Martínez, 1958 | | 1 | Species | Taxom | | Anisocanthon Martinez & P..., 1868 | | 1 | Species | Taxom | | Anomala Samouelle, 1916 | Shining Leaf Chafer | 4 | Species | Taxom | | Anomiopus Westwood, 1842 | | 8 | Species | Taxom | | Anoplosiagum Blanchard, 1921 | | 1 | Species | Taxom | | Aphodius Hellwig, 1798 | | 1 | Species | Taxom | | Apterodemidea Gutierrez Al..., 1903 | | 1 | Species | Taxom | | Archophileurus Kolbe, 1847 | | 7 | Species | Taxom | | Astaena Erichson, 1918 | | 4 | Species | Taxom | | Ataeniopsis Petrovitz, 1877 | | 2 | Species | Taxom | | Ataenius Harold, 1887 | | 30 | Species | Taxom | | Ateuchus Weber, 1935 | | 7 | Species | Taxom | | Auperia Chevrolat, 1864 | | 1 | Species | Taxom | | Badelina J.Thomson, 1833 | | 1 | Species | Taxom | | Besourenga Vaz-de-Mello, 1935 | | 3 | Species | Taxom | | Blackburneus Schmidt, 1911 | | 5 | Species | Taxom | | Blaesia Burmeister, 1905 | | 2 | Species | Taxom | | Blepharotoma Blanchard, 1919 | | 1 | Species | Taxom | | Bolbites Harold, 1868 | | 1 | Species | Taxom | | Bothynus Hope, 1968 | | 12 | Species | Taxom | | Bruchaphodius Martínez, 1871 | | 1 | Species | Taxom | | Bryantella Britton, 0 | | 1 | Species | Taxom | | Byrsopolis Burmeister, 2022 | | 1 | Species | Taxom | | Canthidium Erichson, 1939 | | 12 | Species | Taxom | | Canthon Hoffmannsegg, 1939 | Tumblebug | 42 | Species | Taxom | | Cartwrightia Islas, 1967 | | 1 | Species | Taxom | | Ceraspis Serville, 1924 | | 1 | Species | Taxom | | Chalcocopris Burmeister, 2015 | | 2 | Species | Taxom | | Chalcoplethis Burmeister, 0 | | 1 | Species | Taxom | | Chalepides Casey, 1963 | | 9 | Species | Taxom | | Cnemida Kirby, 1824 | | 1 | Species | Taxom | | Coelosis Hope, 1846 | | 3 | Species | Taxom | | Copris Geoffroy, 1959 | Mexican dung beetle | 1 | Species | Taxom | | Coprophanaeoides Edmonds, 1891 | | 1 | Species | Taxom | | Coprophanaeus Olsoufieff, 1847 | | 14 | Species | Taxom | | Cotinis Burmeister, 1860 | Green June Beetles | 1 | Species | Taxom | | Cyclidius McLeay, 1789 | | 1 | Species | Taxom | | Cyclocephala Dejean, 1923 | Masked Chafers | 35 | Species | Taxom | | Deltochilum Eschscholtz, 1939 | | 20 | Species | Taxom | | Dendropaemon Perty, 1831 | | 12 | Species | Taxom | | Diabroctis Gistl, 1877 | | 2 | Species | Taxom | | Dichotomius Hope, 1869 | | 22 | Species | Taxom | | Dicrania Lepeletier &..., 1829 | | 3 | Species | Taxom | | Digitonthophagus Balthasar, 1787 | | 1 | Species | Taxom | | Diloboderus Reiche, 1826 | | 1 | Species | Taxom | | Dyscinetus Harold, 1847 | Rice Beetles | 3 | Species | Taxom | | Enema Hope, 1775 | | 1 | Species | Taxom | | Erioscelis Burmeister, 1829 | | 1 | Species | Taxom | | Eudinopus Burmeister, 1802 | | 1 | Species | Taxom | | Euetheola Bates, 2014 | | 3 | Species | Taxom | | Euphoria Burmeister, 1775 | | 1 | Species | Taxom | | Eurysternus Dalman, 1988 | | 10 | Species | Taxom | | Eutrichillum | | 1 | Species | Taxom | | Ferrerianus | | 1 | Species | Taxom | | Francmonrosia Pereira & Ma..., 1840 | | 1 | Species | Taxom | | Geniates Kirby, 1911 | | 2 | Species | Taxom | | Genieridium Vaz-de-Mello, 1913 | | 2 | Species | Taxom | | Gibboryctes Endrödi, 1974 | | 2 | Species | Taxom | | Glaphyrocanthon Martínez, 1952 | | 1 | Species | Taxom | | Golofa Hope, 1984 | | 2 | Species | Taxom | | Gonaphodioides Dellacasa M...., 1973 | | 1 | Species | Taxom | | Gromphas Brullé, 1837 | | 6 | Species | Taxom | | Gymnetis MacLeay, 1833 | | 22 | Species | Taxom | | Hemiphileurus Kolbe, 1978 | | 1 | Species | Taxom | | Heterogomphus Burmeister, 1830 | | 5 | Species | Taxom | | Heteropelidnota Ohaus, 1912 | | 1 | Species | Taxom | | Heterosternus Moure, 1940 | | 1 | Species | Taxom | | Hiekeianus Endrödi, 1978 | | 1 | Species | Taxom | | Holocanthon | | 1 | Species | Taxom | | Holocephalus Hope, 2016 | | 2 | Species | Taxom | | Hologymnetis Martínez, 1825 | | 1 | Species | Taxom | | Homophileurus Kolbe, 1910 | | 1 | Species | Taxom | | Homothermon Ohaus, 1855 | | 1 | Species | Taxom | | Hoplopyga Thomson, 1833 | | 9 | Species | Taxom | | Iarupea Martínez, 1973 | | 2 | Species | Taxom | | Inca Lepeletier &..., 1789 | | 3 | Species | Taxom | | Junkia Dalla Torre, 1894 | | 1 | Species | Taxom | | Labarrus Mulsant and Rey, 1941 | | 2 | Species | Taxom | | Lagochile Hoffmannsegg, 1914 | | 3 | Species | Taxom | | Leiopsammodius Rakovic, 1955 | | 1 | Species | Taxom | | Leucothyreus MacLeay, 1917 | | 9 | Species | Taxom | | Lichniops Gutierrez Al..., 1946 | | 1 | Species | Taxom | | Ligyrus Erichson, 1872 | | 1 | Species | Taxom | | Liogenys Guérin-Ménév..., 1851 | | 10 | Species | Taxom | | Lobogeniates Ohaus, 1917 | | 2 | Species | Taxom | | Lomanoxia Martínez, 1867 | | 2 | Species | Taxom | | Lomanoxoides Stebnicka, 1963 | | 1 | Species | Taxom | | Luederwaldtinia Martínez, 1901 | | 1 | Species | Taxom | | Macraspis MacLeay, 1844 | | 6 | Species | Taxom | | Macrocranius | | 1 | Species | Taxom | | Macrodactylus Dejean, 1921 | Rose Chafers | 4 | Species | Taxom | | Malagoniella Martínez, 0 | | 5 | Species | Taxom | | Marmarina Kirby, 1833 | | 2 | Species | Taxom | | Martineziana Chalumeau & ..., 1963 | | 5 | Species | Taxom | | Megaphanaeus | | 4 | Species | Taxom | | Megasoma Kirby, 2003 | Elephant beetles | 8 | Species | Taxom | | Microrutela Bates, 1832 | | 1 | Species | Taxom | | Neocorvicoana | | 1 | Species | Taxom | | Neodiapterna Dellacasa, 1861 | | 1 | Species | Taxom | | Nialaphodius Kolbe, 1801 | | 1 | Species | Taxom | | Nobius Mulsant & Rey, 1851 | | 1 | Species | Taxom | | Odontolytes Koshantschikov, 1963 | | 1 | Species | Taxom | | Omaloplia Schönherr, 1817 | | 1 | Species | Taxom | | Ontherus Erichson, 1775 | | 9 | Species | Taxom | | Onthocharis Westwood, 1955 | | 1 | Species | Taxom | | Onthophagus Latreille, 1859 | | 6 | Species | Taxom | | Oxyligyrus Arrow, 1847 | | 1 | Species | Taxom | | Oxysternon Laporte, 1840 | | 2 | Species | Taxom | | Paracotalpa Ohaus, 1915 | | 1 | Species | Taxom | | Parataenius Balthasar, 2009 | | 2 | Species | Taxom | | Pedaridium Harold, 1951 | | 2 | Species | Taxom | | Pelidnota MacLeay, 1782 | Precious Metal Scarabs | 13 | Species | Taxom | | Phanaeus MacLeay, 1846 | Burrowing Dung Beetles | 6 | Species | Taxom | | Phileurus Sieg, 1758 | | 7 | Species | Taxom | | Philochloenia Dejean, 1921 | | 1 | Species | Taxom | | Philoscaptus Brèthes, 1847 | | 1 | Species | Taxom | | Phyllophaga Harris, 1921 | June beetles | 5 | Species | Taxom | | Pinotus Erichson, 1929 | | 1 | Species | Taxom | | Platytomus Mulsant, 1948 | | 2 | Species | Taxom | | Plectris Lepeletier a..., 1967 | | 17 | Species | Taxom | | Protaetia Burmeister, 1841 | | 1 | Species | Taxom | | Pseudocanthon Bates, 1846 | | 3 | Species | Taxom | | Pseudodorysthetus | | 2 | Species | Taxom | | Pseudoliogenys Moser, 1892 | | 1 | Species | Taxom | | Raysymmela Saylor, 1921 | | 2 | Species | Taxom | | Rhizogeniates Ohaus, 1845 | | 1 | Species | Taxom | | Rutela Latreille, 1758 | | 2 | Species | Taxom | | Saprosites Redtenbacher, 1869 | | 3 | Species | Taxom | | Scatonomus Erichson, 1959 | | 2 | Species | Taxom | | Scybalocanthon Martínez, 1855 | | 3 | Species | Taxom | | Stenocrates Burmeister, 1911 | | 9 | Species | Taxom | | Strategus Kirby, 1775 | Ox Beetles | 5 | Species | Taxom | | Sulcophanaeus Olsoufieff, 1840 | | 4 | Species | Taxom | | Sylvicanthon Halffter & M..., 1920 | | 1 | Species | Taxom | | Symmela Erichson, 1921 | | 3 | Species | Taxom | | Tetraechma Agassiz, 1939 | | 3 | Species | Taxom | | Tomarus Erichson, 1774 | Carrot Beetles | 4 | Species | Taxom | | Trichillidium Vaz-de-Mello, 1932 | | 1 | Species | Taxom | | Trichillum Harold, 2005 | | 7 | Species | Taxom | | Trichiopsammobius Petrovitz, 1963 | | 1 | Species | Taxom | | Trigonopeltastes Burmeister &..., 1818 | | 1 | Species | Taxom | | Trioplus Burmeister, 1829 | | 1 | Species | Taxom | | Trizogeniates Ohaus, 1857 | | 2 | Species | Taxom | | Uroxys Westwood, 1940 | | 3 | Species | Taxom | | Zonocopris Arrow, 2007 | | 2 | Species | Taxom |
Species list for Paraguay
545 Species and 44 Subspecies | Aceratus davus Burmeister, 1847 | | | Actinobolus radians Westwood, 1841 | | | Agamopus viridis Boucomont, 1928 | | | Agaocephala cornigera Lepeletier &..., 1828 | | | Aidophus infuscatopennis Schmidt, 1909 | | | Allorrhina carmelita Burmeister, 1842 | | | Allorrhina cornifrons Gory & Perch..., 1833 | | | Allorrhina menetriesi Mannerheim, 1829 | | | Anahi guaraniticus Martínez, 1958 | | | Anisocanthon villosus Harold, 1868 | | | Anomala inconstans Burmeister, 1844 | | | Anomala nitidula Blanchard, 1851 | | | Anomala testaceipennis (Blanchard, 1851) | ⇒ Anomala nitidula Blanchard, 1851 | | Anomala toliensis Ohaus, 1916 | | | Anomiopus birai Canhedo, 2006 | | | Anomiopus germari Harold, 1867 | | | Anomiopus nigrocoeruleus Martínez, 1955 | | | Anomiopus paraguaiensis Canhedo, 2004 | | | Anomiopus pereirai Martínez, 1955 | | | Anomiopus tuberifrons Canhedo, 2004 | | | Anomiopus virescens Westwood, 1842 | | | Anoplosiagum macrophyllum Moser, 1921 | | | Apterodemidea paraguayensis Arrow, 1903 | | | Archophileurus alternans Endrödi, 1977 | | | Archophileurus chaconus Kolbe, 1910 | | | Archophileurus elatus Prell, 1914 | | | Archophileurus fodiens Kolbe, 1910 | | | Archophileurus latipennis Burmeister, 1847 | | | Archophileurus spinosus Dechambre, 2006 | | | Archophileurus vervex Burmeister, 1847 | | | Astaena conformis Blanchard, 1850 | | | Astaena cujabana Moser, 1918 | | | Astaena cuyabana Moser, 1918 | | | Astaena tenellula Moser, 1918 | | | Ataeniopsis haroldi Steinheil, 1872 | | | Ataeniopsis pusillus Burmeister, 1877 | | | Ataenius aequalis Harold, 1880 | | | Ataenius bispinulosus Schmidt, 1911 | | | Ataenius blapoides Balthasar, 1947 | | | Ataenius catarinaensis Stebnicka, 2007 | | | Ataenius clitellarius Petrovitz, 1973 | | | Ataenius columbicus Harold, 1880 | | | Ataenius complicatus Harold, 1869 | | | Ataenius crenulatus (Schmidt, 1910) | ⇒ Parataenius derbesis Solier, 1851 | | Ataenius forsteri Balthasar, 1960 | | | Ataenius granchacoensis (Balthasar, 1939) | ⇒ Ataenius platensis Blanchard, 1846 | | Ataenius imbricatus Melsheimer, 1845 | | | Ataenius impiger Schmidt, 1916 | | | Ataenius insculptus Horn, 1887 | | | Ataenius koelleri Balthasar, 1963 | | | Ataenius londrinae Stebnicka, 2007 | | | Ataenius luctuosus Burmeister, 1877 | | | Ataenius morator Harold, 1869 | | | Ataenius noques Stebnicka, 2007 | | | Ataenius opacipennis Schmidt, 1910 | | | Ataenius petrovitzi Balthasar, 1960 | | | Ataenius picinus Harold, 1861 | Antarctic blennies | | Ataenius platensis Blanchard, 1846 | | | Ataenius purator Harold, 1868 | | | Ataenius schmidti Stebnicka, 2003 | | | Ataenius sculptilis (Harold, 1868) | ⇒ Ataenius insculptus Horn, 1887 | | Ataenius stercorator (Fabricius, 1775) | ⇒ Ataenius strigatus Say, 1823 | | Ataenius strigatus Say, 1823 | Black turfgrass ataenius | | Ataenius strigicauda Bates, 1887 | | | Ataenius strigicaudus Bates, 1887 | | | Ateuchus apicatus Harold, 1867 | | | Ateuchus balthasari Martínez, 1952 | | | Ateuchus breve Harold, 1868 | | | Ateuchus contractus Balthasar, 1939 | | | Ateuchus ovale Boucomont, 1928 | | | Ateuchus ovalis Boucomont, 1928 | | | Ateuchus viridimicans Boucomont, 1935 | | | Auperia denominata Chevrolat, 1864 | | | Badelina aterrima Gory & Perch..., 1833 | | | Besourenga amarillai Aguilar-Julio, 2001 | | | Besourenga minutum Saylor, 1935 | | | Besourenga minutus Saylor, 1935 | | | Blackburneus argentinensis Schmidt, 1909 | | | Blackburneus caracaensis Petrovitz, 1970 | | | Blackburneus furcatus Schmidt, 1909 | | | Blackburneus laxepunctatus Schmidt, 1910 | | | Blackburneus richteri (Schmidt, 1911) | ⇒ Nobius hepaticus Roth, 1851 | | Blaesia atra Burmeister, 1842 | | | Blaesia subrugosa Moser, 1905 | | | Blepharotoma heynei Moser, 1919 | | | Bolbites onitoides Harold, 1868 | | | Bothynus araya Duarte & Grossi, 2020 | | | Bothynus ascanius Kirby, 1818 | | | Bothynus cunctator Mannerheim, 1829 | | | Bothynus cyclops Burmeister, 1847 | | | Bothynus dasypleurus Germar, 1824 | | | Bothynus exaratus Burmeister, 1847 | | | Bothynus horridus Endrödi, 1968 | | | Bothynus medon Germar, 1824 | | | Bothynus spinophallicus Ratcliffe & ..., 2021 | | | Bothynus striatellus Fairmaire, 1878 | | | Bothynus villiersi Endrödi, 1968 | | | Bruchaphodius ovalipennis Harold, 1871 | | | Bryantella smaragda | Jumping spiders | | Byrsopolis vazdemelloi Medeiros, Se..., 2022 | | | Canthidium barbacenicum Preudhomme d..., 1886 | | | Canthidium bokermanni Martinez, Ha..., 1964 | | | Canthidium breve Germar, 1824 | | | Canthidium cupreum Blanchard, 1846 | | | Canthidium dispar Harold, 1867 | | | Canthidium femoratum Boucomont, 1935 | | | Canthidium gigas Balthasar, 1939 | | | Canthidium kelleri Martinez, Ha..., 1964 | | | Canthidium multipunctatum Balthasar, 1939 | | | Canthidium pinotoides (Balthasar, 1939) | ⇒ Canthidium barbacenicum Preudhomme d..., 1886 | | Canthidium prasinum Blanchard, 1846 | | | Canthidium titschacki Balthasar, 1939 | | | Canthon angularis Harold, 1868 | | | Canthon angularis angularis | | | Canthon arriagadai Vieira, Vaz-..., 2019 | | | Canthon cobosi Pereira & Ma..., 1960 | | | Canthon curvodilatatus Schmidt, 1920 | | | Canthon denticulatus Schmidt, 1922 | | | Canthon edentulus Harold, 1868 | | | Canthon forcipatus Harold, 1868 | | | Canthon hendrichsi Halffter & M..., 1968 | | | Canthon ibarragrassoi Martínez, 1952 | | | Canthon laminatus Balthasar, 1939 | | | Canthon lituratus Germar, 1813 | | | Canthon lividus Blanchard, 1846 | | | Canthon lividus lividus | | | Canthon lividus seminitens Harold, 1868 | | | Canthon luctuosus Harold, 1868 | | | Canthon mutabilis Lucas, 1857 | | | Canthon muticus Harold, 1867 | | | Canthon octodentatus Schmidt, 1920 | | | Canthon oliverioi Pereira & Ma..., 1956 | | | Canthon ornatus Redtenbacher, 1867 | | | Canthon ornatus ornatus | | | Canthon podagricus Harold, 1868 | | | Canthon quadratus Blanchard, 1846 | | | Canthon quinquemaculatus Castelnau, 1840 | | | Canthon rutilans (Castelnau, 1840) | ⇒ Francmonrosia rutilans Castelnau, 1840 | | Canthon rutilans cyanescens Harold, 1868 | | | Canthon rutilans rutilans | | | Canthon septemmaculatus Latreille, 1812 | | | Canthon septemmaculatus septemmaculatus | | | Canthon smaragdulus Fabricius, 1781 | | | Canthon smaragdulus smaragdulus | | | Canthon staigi Pereira, 1953 | | | Canthon substriatus Harold, 1868 | | | Canthon triangularis Drury, 1770 | | | Canthon unicolor Blanchard, 1846 | | | Canthon unicolor fortemarginatus Balthasar, 1939 | | | Canthon unicolor unicolor | | | Canthon velutinus Harold, 1868 | | | Canthon virens Mannerheim, 1829 | | | Canthon virens paraguayanus Balthasar, 1939 | | | Cartwrightia cartwrighti Cartwright, 1967 | | | Ceraspis squamulata Moser, 1924 | | | Chalcocopris hesperus Olivier, 1789 | | | Chalcocopris inexpectatus Rossini & Va..., 2015 | | | Chalcoplethis kirbii kirbii | | | Chalepides barbatus Fabricius, 1781 | | | Chalepides barbatus argentinus Prell, 1938 | | | Chalepides comes Prell, 1936 | | | Chalepides eucephalus Casey, 1915 | | | Chalepides hydrophiloides Burmeister, 1847 | | | Chalepides luridus Burmeister, 1847 | | | Chalepides narcisoi Martínez, 1978 | | | Chalepides unicolor Endrödi, 1963 | | | Cnemida lacerata Germar, 1824 | | | Coelosis bicornis Leske, 1779 | | | Coelosis biloba Linnaeus, 1767 | | | Coelosis hippocrates Blanchard, 1846 | | | Copris megasoma Matthews & H..., 1959 | | | Coprophanaeoides refulgens Waterhouse, 1891 | | | Coprophanaeus (Megaphanaeus) bonariensis Gory, 1844 | | | Coprophanaeus (Megaphanaeus) ensifer Germar, 1821 | | | Coprophanaeus (Megaphanaeus) lancifer Linnaeus, 1767 | | | Coprophanaeus (Metallophanaeus) horus Waterhouse, 1891 | | | Coprophanaeus (Metallophanaeus) pessoai Pereira, 1949 | | | Coprophanaeus (Metallophanaeus) saphirinus Sturm, 1826 | | | Coprophanaeus saphirinus saphirinus | | | Coprophanaeus (Metallophanaeus) thalassinus Perty, 1830 | | | Coprophanaeus cyanescens Olsufieff, 1924 | | | Coprophanaeus jasius Olivier, 1789 | | | Coprophanaeus milon Blanchard, 1846 | | | Coprophanaeus rigoutorum Arnaud, 2002 | | | Coprophanaeus spitzi Pessôa, 1934 | | | Coprophanaeus telamon Erichson, 1847 | | | Cotinis paraguayanus Thomson, 1860 | | | Cyclidius elongatus Olivier, 1789 | | | Cyclocephala amazona Linnaeus, 1767 | | | Cyclocephala amazona amazona | | | Cyclocephala atricapilla Mannerheim, 1829 | | | Cyclocephala celata Dechambre, 1980 | | | Cyclocephala concolor Burmeister, 1847 | | | Cyclocephala conspicua Sharp, 1877 | | | Cyclocephala discolor Herbst, 1792 | | | Cyclocephala dispar Herbst, 1792 | | | Cyclocephala epistomalis Bates, 1888 | | | Cyclocephala erotylina Arrow, 1914 | | | Cyclocephala forsteri Endrödi, 1963 | | | Cyclocephala forsteri forsteri | | | Cyclocephala gravis Bates, 1888 | | | Cyclocephala laminata Burmeister, 1847 | | | Cyclocephala latericia Höhne, 1923 | | | Cyclocephala lizeri Martínez, 1964 | | | Cyclocephala lunulata Burmeister, 1847 | Twister | | Cyclocephala mechae Martínez, 1978 | | | Cyclocephala mecynotarsis Höhne, 1923 | | | Cyclocephala meinanderi Endrödi, 1964 | | | Cyclocephala melanocephala (Fabricius, 1775) | ⇒ Cyclocephala olivieri Arrow, 1911 | | Cyclocephala metrica Steinheil, 1872 | | | Cyclocephala minuta Burmeister, 1847 | | | Cyclocephala modesta Hansen, 1916 | | | Cyclocephala ochracea Prell, 1937 | | | Cyclocephala olivieri Arrow, 1911 | | | Cyclocephala ovulum (Bates, 1888) | ⇒ Cyclocephala testacea Burmeister, 1847 | | Cyclocephala paraguayensis Arrow, 1903 | | | Cyclocephala paraguayensis marginella Endrödi, 1966 | | | Cyclocephala putrida Burmeister, 1847 | | | Cyclocephala simulatrix Höhne, 1923 | | | Cyclocephala suturalis Ohaus, 1911 | | | Cyclocephala testacea Burmeister, 1847 | Twister | | Cyclocephala tucumana Brèthes, 1904 | | | Cyclocephala vestita Höhne, 1923 | | | Deltochilum amandaarcanjoae Silva, Louza..., 2015 | | | Deltochilum aspericolle Bates, 1870 | | | Deltochilum brasiliense Castelnau, 1840 | | | Deltochilum cristatum Paulian, 1938 | | | Deltochilum cupreicolle Blanchard, 1846 | | | Deltochilum dentipes Eschscholtz, 1822 | | | Deltochilum elongatum Felsche, 1907 | | | Deltochilum icariforme Paulian, 1938 | | | Deltochilum icaroides Balthasar, 1939 | | | Deltochilum icarus Olivier, 1789 | | | Deltochilum irroratum Castelnau, 1840 | | | Deltochilum kolbei Paulian, 1938 | | | Deltochilum komareki Balthasar, 1939 | | | Deltochilum pseudoicarus Balthasar, 1939 | | | Deltochilum rubripenne Gory, 1831 | | | Deltochilum valgum Burmeister, 1873 | | | Deltochilum variolosum Burmeister, 1873 | | | Deltochilum viridescens Martinez, 1948 | | | Deltochilum viridicupreum Balthasar, 1939 | | | Dendropaemon (Coprophanaeoides) renatii Olsoufieff, 1924 | | | Dendropaemon (Glaphyropaemon) bahianus (Harold, 1868) | ⇒ Coprophanaeoides bahianus Harold, 1868 | | Dendropaemon (Rutilopaemon) refulgens (Waterhouse, 1891) | ⇒ Coprophanaeoides refulgens Waterhouse, 1891 | | Dendropaemon (Sulcopaemon) fasces Blut, 1939 | | | Dendropaemon (Sulcopaemon) latistriatus Génier & Arn..., 2016 | | | Dendropaemon (Titthopaemon) denticollis Felsche, 1909 | | | Dendropaemon batrachites Blut, 1939 | | | Dendropaemon pilosissimus Génier & Arn..., 2016 | | | Dendropaemon quadratus Laporte, 1832 | | | Dendropaemon similis Blut, 1939 | | | Dendropaemon tenuitarsis Felsche, 1909 | | | Dendropaemon viridipennis Laporte, 1831 | | | Diabroctis mimas Linnaeus, 1758 | | | Diabroctis mirabilis Harold, 1877 | | | Dichotomius (Luederwaldtinia) bosqui Pereira, 1941 | | | Dichotomius (Luederwaldtinia) carbonarius Mannerheim, 1829 | | | Dichotomius (Luederwaldtinia) crinicollis Germar, 1824 | | | Dichotomius (Luederwaldtinia) glaucus Harold, 1869 | | | Dichotomius (Luederwaldtinia) micans Luederwaldt, 1924 | | | Dichotomius (Luederwaldtinia) nisus Olivier, 1789 | | | Dichotomius (Luederwaldtinia) nobilis Waterhouse, 1891 | | | Dichotomius (Luederwaldtinia) opacipennis Luederwaldt, 1931 | | | Dichotomius (Luederwaldtinia) punctatostriatus (Felsche, 1901) | ⇒ Luederwaldtinia punctatostriatus Felsche, 1901 | | Dichotomius (Luederwaldtinia) sericeus Harold, 1867 | | | Dichotomius (Selenocopris) ascanius Harold, 1869 | | | Dichotomius (Selenocopris) bicuspis Germar, 1824 | | | Dichotomius (Selenocopris) guaranii Gandini & Ag..., 2009 | | | Dichotomius (Selenocopris) inflaticollis Felsche, 1910 | | | Dichotomius anthrax Felsche, 1901 | | | Dichotomius bos Blanchard, 1846 | | | Dichotomius depressicollis Harold, 1867 | | | Dichotomius fornicatus Luederwaldt, 1931 | | | Dichotomius joelus Gandini & Ag..., 2009 | | | Dichotomius mormon Ljungh, 1799 | | | Dichotomius nitidissimus Waterhouse, 1891 | | | Dichotomius reichei Harold, 1869 | | | Dicrania flavipennis Kraatz, 1896 | | | Dicrania fraudulenta Frey, 1972 | | | Dicrania nigriceps Guérin-Ménev..., 1829 | | | Digitonthophagus gazella Fabricius, 1787 | Brown dung beetle | | Diloboderus abderus Sturm, 1826 | | | Dyscinetus dubius Olivier, 1789 | | | Dyscinetus rugifrons Burmeister, 1847 | | | Enema pan Fabricius, 1775 | | | Erioscelis emarginata Mannerheim, 1829 | | | Eudinopus dytiscoides Schreibers, 1802 | | | Euetheola humilis Burmeister, 1847 | Spotted kelpfish | | Euetheola latipennis Arrow, 1911 | | | Euetheola paraguayensis Prokofiev, 2014 | | | Euphoria lurida Fabricius, 1775 | | | Eurysternus aeneus Génier, 2009 | | | Eurysternus caribaeus Herbst, 1789 | | | Eurysternus cyanescens Balthasar, 1939 | | | Eurysternus foedus Guérin-Ménev..., 1844 | | | Eurysternus hirtellus Dalman, 1824 | | | Eurysternus jessopi Martínez, 1988 | | | Eurysternus navajasi Martínez, 1988 | | | Eurysternus nigrovirens Génier, 2009 | | | Eurysternus parallelus Castelnau, 1840 | | | Eurysternus rivaldii Martínez, 1988 | | | Eutrichillum hirsutum Boucomont, 1928 | | | Ferrerianus biimpressus Schmidt, 1909 | | | Francmonrosia rutilans Castelnau, 1840 | | | Geniates impressicollis Ohaus, 1911 | | | Genieridium bidens Balthasar, 1938 | | | Genieridium cryptops Arrow, 1913 | | | Gibboryctes bollei Dechambre, 2006 | | | Gibboryctes szelenyii Endrödi, 1974 | | | Glaphyrocanthon ibarragrassoi Martínez, 1952 | | | Golofa horizontalis Voirin, 1984 | | | Gonaphodioides sincerus Petrovitz, 1973 | | | Gromphas aeruginosa Perty, 1830 | | | Gromphas amazonica Bates, 1870 | | | Gromphas dichroa Blanchard, 1846 | | | Gromphas inermis Harold, 1869 | | | Gromphas lacordairei (Brullé, 1837) | ⇒ Gromphas inermis Harold, 1869 | | Gymnetis bomplandi Schaum, 1844 | | | Gymnetis bonplandi Schaum, 1844 | | | Gymnetis carbo Schürhoff, 1937 | | | Gymnetis chalcipes Gory & Perch..., 1833 | | | Gymnetis chalcipes undata Olivier, 1789 | | | Gymnetis drogoni Ratcliffe, 2018 | | | Gymnetis flava (Burmeister, 1842) | ⇒ Gymnetis xanthospila Schaum, 1844 | | Gymnetis flaveola schistacea Burmeister, 1847 | | | Gymnetis flavomarginata Blanchard, 1846 | | | Gymnetis hebraica Drapiez, 1820 | | | Gymnetis hepatica Di Iorio, 2013 | | | Gymnetis hieroglyphica Vigors, 1825 | | | Gymnetis holosericea (Voet, 1779) | ⇒ Gymnetis xanthospila Schaum, 1844 | | Gymnetis litigiosa Gory & Perch., 1833 | | | Gymnetis pantherina (Burmeister, 1842) | ⇒ Gymnetis hieroglyphica rubrocincta Schürhoff, 1937 | | Gymnetis pantherina meleagris Burmeister, 1842 | | | Gymnetis pudibunda Burmeister, 1866 | | | Gymnetis punctipennis Burmeister, 1842 | | | Gymnetis rufilatris Illiger, 1800 | | | Gymnetis schistacea (Burmeister, 1847) | ⇒ Gymnetis flaveola schistacea Burmeister, 1847 | | Gymnetis tigrina (Gory & Perch..., 1833) | ⇒ Marmarina tigrina Gory & Perch..., 1833 | | Hemiphileurus costatus Endrödi, 1978 | | | Heterogomphus achilles Burmeister, 1847 | | | Heterogomphus aidoneus Perty, 1830 | | | Heterogomphus monotuberculatus Dupuis & Dec..., 2008 | | | Heterogomphus pauson Perty, 1830 | | | Heteropelidnota kuhnti Ohaus, 1912 | | | Heterosternus dubitabilis Hustache, 1940 | | | Hiekeianus tuberculatus Endrödi, 1978 | | | Holocanthon fuscorubrus Blanchard, 1845 | | | Holocephalus cristatus Gillet, 1907 | | | Holocephalus pokornyi Nunes & Vaz-..., 2016 | | | Hologymnetis undulata Vigors, 1825 | | | Homophileurus luederwaldti Ohaus, 1910 | | | Homothermon praemorsus Burmeister, 1855 | | | Hoplopyga albiventris Gory & Perch..., 1833 | | | Hoplopyga brasiliensis Gory & Perch..., 1833 | | | Hoplopyga gosseti Antoine, 2008 | | | Hoplopyga liturata Olivier, 1789 | | | Hoplopyga miliaris Gory & Perch..., 1833 | | | Hoplopyga multipunctata Gory & Perch..., 1833 | | | Hoplopyga ocellata Gory & Perch..., 1833 | | | Hoplopyga singularis Gory & Perch..., 1833 | | | Iarupea lopeteguii Martínez, 1953 | | | Iarupea serratipennis Petrovitz, 1973 | | | Inca bonplandi Gyllenhal, 1817 | | | Inca pulverulenta Olivier, 1789 | | | Inca pulverulentus Olivier, 1789 | | | Junkia ceylanica Nonfried, 1894 | | | Labarrus parnaguaensis Petrovitz, 1961 | | | Labarrus pseudolividus Balthasar, 1941 | | | Lagochile bipunctata heliophila Ohaus, 1914 | | | Lagochile castanicolor Ohaus, 1914 | | | Leiopsammodius bolivianus Cartwright, 1955 | | | Leucothyreus bohlsi Ohaus, 1918 | | | Leucothyreus cribratipennis Blanchard, 1851 | | | Leucothyreus femoralis Blanchard, 1851 | | | Leucothyreus femoralis femoralis | | | Leucothyreus flavipes Eschscholtz, 1822 | | | Leucothyreus marginaticollis Blanchard, 1846 | | | Leucothyreus sosthenes Ohaus, 1918 | | | Leucothyreus spinifer Ohaus, 1918 | | | Leucothyreus viridiaeneus Ohaus, 1917 | | | Lichniops barberoi Gutierrex Al..., 1946 | | | Ligyrus burmeisteri Steinheil, 1872 | | | Liogenys bidenticeps Moser, 1919 | | | Liogenys elegans Nonfried, 1891 | | | Liogenys fusca Blanchard, 1851 | | | Liogenys latitarsis Moser, 1918 | | | Liogenys nigrofusca Moser, 1918 | | | Liogenys opthalmica Frey, 1973 | | | Liogenys pallens Blanchard, 1851 | | | Liogenys rufocastanea Moser, 1918 | | | Liogenys rufoflava Moser, 1918 | | | Liogenys suturalis Blanchard, 1851 | | | Lobogeniates flavipes Ohaus, 1917 | | | Lobogeniates nigripennis Ohaus, 1917 | | | Lomanoxia chacocola Krikken, 1972 | | | Lomanoxia costulata Harold, 1867 | | | Lomanoxoides tesari Balthasar, 1963 | | | Luederwaldtinia punctatostriatus Felsche, 1901 | | | Macraspis chrysis sudista Soula, 2006 | | | Macraspis cincta Drury, 1782 | | | Macraspis festiva Burmeister, 1844 | | | Macraspis forsteri Soula, 1998 | | | Macraspis forsteri forsteri | | | Macraspis morio Burmeister, 1844 | | | Macrocranius similis Schürhoff, 1935 | | | Macrodactylus fulvipes Moser, 1918 | | | Macrodactylus obscuritarsis Moser, 1926 | | | Macrodactylus pallens Blanchard, 1850 | | | Macrodactylus paraguayensis Moser, 1921 | | | Malagoniella astyanax punctatostriata Blanchard, 1846 | | | Malagoniella magnifica Balthasar, 1939 | | | Malagoniella puncticollis Blanchard, 1846 | | | Malagoniella puncticollis aeneicollis Waterhouse, 1890 | | | Malagoniella puncticollis puncticollis | | | Marmarina insculpta Kirby, 1818 | | | Marmarina tigrina Gory & Perch..., 1833 | | | Martineziana argentina Harold, 1867 | | | Martineziana dutertrei Chalumeau, 1983 | | | Martineziana excavaticollis Blanchard, 1846 | Beetle | | Martineziana separata Schmidt, 1909 | | | Martineziana simplex Balthasar, 1963 | | | Megaphanaeus bonariensis Gory, 1844 | | | Megaphanaeus ensifer (Germar, 1821) | ⇒ Coprophanaeus ensifer Germar, 1821 | | Megaphanaeus lancifer Linnaeus, 1767 | | | Megaphanaeus vicinus (Martínez, 1944) | ⇒ Coprophanaeus bonariensis Gory, 1844 | | Megasoma actaeon Linnaeus, 1758 | Actaeon beetle | | Megasoma anubis Chevrolat, 1836 | | | Megasoma bollei Dechambre, 2006 | | | Megasoma janus (Felsche, 1906) | ⇒ Megasoma actaeon Linnaeus, 1758 | | Megasoma janus argentinum Höhne, 1923 | | | Megasoma janus janus | | | Megasoma joergenseni Bruch, 1910 | | | Megasoma joergenseni penyai Nagai, 2003 | | | Microrutela coerulea Perty, 1832 | | | Neocorvicoana reticulata Kirby, 1818 | | | Neodiapterna erichsoni Harold, 1861 | | | Nialaphodius nigrita Fabricius, 1801 | | | Nobius hepaticus Roth, 1851 | | | Odontolytes landai Balthasar, 1963 | | | Ontherus androgynus Génier, 1996 | | | Ontherus aphodioides Burmeister, 1874 | | | Ontherus appendiculatus Mannerheim, 1829 | | | Ontherus azteca Harold, 1869 | | | Ontherus carinicollis Luederwaldt, 1930 | | | Ontherus digitatus Harold, 1868 | | | Ontherus erosioides Luederwaldt, 1930 | | | Ontherus sulcator Fabricius, 1775 | | | Onthocharis melancholica (Martínez, 1955) | ⇒ Anomiopus virescens Westwood, 1842 | | Onthophagus buculus Mannerheim, 1829 | | | Onthophagus carpophilus Pereira & Ha..., 1961 | | | Onthophagus chacoensis Rossini, Vaz..., 2018 | | | Onthophagus hirculus (Mannerheim, 1829) | ⇒ Onthophagus mutatus Harold, 1859 | | Onthophagus hircus Billberg, 1815 | | | Onthophagus mutatus Harold, 1859 | | | Oxyligyrus nasutus Burmeister, 1847 | | | Oxysternon festivum festivum | | | Oxysternon palaemon Castelnau, 1840 | | | Parataenius derbesis Solier, 1851 | | | Parataenius martinezi Stebnicka & ..., 2009 | | | Pedaridium mansosotoi (Martínez, 1951) | ⇒ Genieridium cryptops Arrow, 1913 | | Pelidnota (Strigidia) cuprea Germar, 1824 | | | Pelidnota crassipes Ohaus, 1905 | | | Pelidnota fulva Blanchard, 1851 | | | Pelidnota glaberrima meridionalis Soula, 2006 | | | Pelidnota gracilis gracilis | | | Pelidnota kuhnti Ohaus, 1912 | | | Pelidnota liturella assumpta Ohaus, 1928 | | | Pelidnota paraguayensis Bates, 1904 | | | Pelidnota pulchella altoparanaensis Soula, 2006 | | | Pelidnota sordida Germar, 1824 | | | Pelidnota sumptuosa Vigors, 1825 | | | Pelidnota unicolor Drury, 1782 | | | Phanaeus (Notiophanaeus) kirbyi Vigors, 1825 | | | Phanaeus kirbyi kirbyi | | | Phanaeus bogotensis (Kirsch, 1871) | ⇒ Phanaeus hermes Harold, 1868 | | Phanaeus hermes Harold, 1868 | | | Phanaeus menelas (Castelnau, 1840) | ⇒ Sulcophanaeus menelas Castelnau, 1840 | | Phanaeus palaeno Blanchard, 1846 | | | Phileurus affinis (Burmeister, 1847) | ⇒ Phileurus didymus Linnaeus, 1758 | | Phileurus angustatus Kolbe, 1910 | | | Phileurus carinatus Prell, 1914 | | | Phileurus carinatus carinatus | | | Phileurus didymus Linnaeus, 1758 | | | Phileurus kaszabi Endrödi, 1978 | | | Phileurus valgus Linnaeus, 1758 | | | Philochloenia macrophylla Moser, 1921 | | | Philoscaptus bonariensis Burmeister, 1847 | | | Phyllophaga boliviensis Blanchard, 1851 | | | Phyllophaga paraguayana Moser, 1921 | | | Phyllophaga paraguayensis Moser, 1918 | | | Phyllophaga propinqua Moser, 1918 | | | Phyllophaga trichoderma Moser, 1921 | | | Pinotus transiens (Luederwaldt, 1929) | ⇒ Dichotomius bicuspis Germar, 1824 | | Platytomus gregalis Cartwright, 1948 | | | Platytomus longulus Cartwright, 1948 | Horned blenny | | Plectris (Plectris) tomentosa Lepeletier &..., 1825 | | | Plectris aliena Chapin, 1934 | Scarab beetle | | Plectris comata Blanchard, 1850 | | | Plectris corrientensis Moser, 1919 | | | Plectris corvicoana Moser, 1919 | | | Plectris decipiens Burmeister, 1855 | | | Plectris guayrana Frey, 1967 | | | Plectris kriegi Frey, 1973 | | | Plectris laeviscutata Moser, 1918 | | | Plectris lindneri Frey, 1967 | | | Plectris maculicollis Arrow, 1925 | | | Plectris nigrita Moser, 1918 | | | Plectris paraguayensis Moser, 1924 | | | Plectris schoolmeestersi Smith, 2015 | | | Plectris setiventris Moser, 1918 | | | Plectris tacoma Smith, 2008 | | | Plectris tenuevestita Frey, 1967 | | | Protaetia (Protaetia) anovittata Chevrolat, 1841 | | | Pseudocanthon chaquensis Nazaré-Silva..., 2021 | | | Pseudocanthon pantanensis Nazaré-Silva..., 2021 | | | Pseudocanthon xanthurus Blanchard, 1846 | | | Pseudodorysthetus calcaratus Spinola, 1835 | | | Pseudodorysthetus calcaratus calcaratus | | | Pseudoliogenys bidentula Fairmaire, 1892 | | | Raysymmela curtula Erichson, 1835 | | | Raysymmela seticollis Moser, 1921 | | | Rhizogeniates antennatus Curtis, 1845 | | | Rutela histrio Sahlberg, 1823 | | | Rutela lineola Linnaeus, 1758 | | | Saprosites breviusculus Harold, 1867 | | | Saprosites dentipes Harold, 1867 | | | Saprosites sulcatus Harold, 1869 | | | Scatonomus viridis viridis | | | Scatonomus viridis walzi Pereira & Ma..., 1959 | | | Scybalocanthon luctuosus Harold, 1868 | | | Scybalocanthon nigriceps Harold, 1868 | | | Scybalocanthon sexspilotus Guérin-Ménev..., 1855 | | | Stenocrates agricola Dechambre & ..., 2004 | | | Stenocrates clipeatus Endrödi, 1966 | | | Stenocrates cultor Burmeister, 1847 | | | Stenocrates cultor cultor | | | Stenocrates holomelanus Germar, 1824 | | | Stenocrates laborator Fabricius, 1775 | | | Stenocrates laborator australis Endrödi, 1973 | | | Stenocrates ligneus Arrow, 1911 | | | Strategus centaurus Kolbe, 1906 | | | Strategus mandibularis Sternberg, 1910 | | | Strategus surinamensis Burmeister, 1847 | | | Strategus surinamensis hirtus Sternberg, 1910 | | | Strategus validus Fabricius, 1775 | | | Sulcophanaeus faunus Fabricius, 1775 | | | Sulcophanaeus imperator Chevrolat, 1844 | | | Sulcophanaeus imperator imperator | | | Sulcophanaeus menelas Castelnau, 1840 | | | Sylvicanthon foveiventris Schmidt, 1920 | | | Symmela corumbana Moser, 1921 | | | Symmela paschoali Pacheco, Wip..., 2022 | | | Symmela seticollis Moser, 1921 | | | Tetraechma chlorophana Mannerheim, 1828 | | | Tetraechma liturata Germar, 1813 | | | Tetraechma tarsalis Balthasar, 1939 | | | Tomarus burmeisteri Steinheil, 1872 | | | Tomarus fossor Latreille, 1813 | | | Tomarus gibbosus De Geer, 1774 | Carrot beetle | | Trichillidium quadridens Arrow, 1932 | | | Trichillum arrowi (Saylor, 1935) | ⇒ Trichillum pauliani Balthasar, 1939 | | Trichillum boucomonti (Saylor, 1935) | ⇒ Eutrichillum hirsutum Boucomont, 1928 | | Trichillum externepunctatum Preudhomme d..., 1880 | | | Trichillum heydeni Harold, 1868 | | | Trichillum minutum (Saylor, 1935) | ⇒ Besourenga minutum Saylor, 1935 | | Trichillum pauliani Balthasar, 1939 | | | Trichillum pseudoarrowi Vaz-de-Mello..., 2005 | | | Trichiopsammobius brasiliensis Petrovitz, 1963 | | | Trigonopeltastes triangulus Kirby, 1818 | | | Trioplus cylindricus Mannerheim, 1829 | | | Trizogeniates goyanus Ohaus, 1917 | | | Trizogeniates vittatus Lucas, 1857 | | | Uroxys corporaali Balthasar, 1940 | | | Uroxys histeroides Arrow, 1933 | | | Uroxys thoracalis Balthasar, 1940 | | | Zonocopris gibbicollis Harold, 1868 | | | Zonocopris machadoi Vaz-de-Mello, 2007 | |
Subnodes of Scarabaeidae with occurrences in Paraguay | Subfamily | Aphodiinae
Leach, 1815
Aphodiine Dung Beetles, Small dung beetles | 24
67 | Genera
Species |
Taxom
| | Subfamily | Cetoniinae
Leach, 1815
Flower beetles, Flower chafers, Flower scarabs, Fruit and flower chafers | 15
46 | Genera
Species |
Taxom
| | Subfamily | Dynastinae
MacLeay, 1819
Horn beetles, Rhinoceros beetles | 28
109 | Genera
Species |
Taxom
| | Subfamily | Melolonthinae
Samouelle, 1819
May Beetles and Junebugs, White grubs | 17
56 | Genera
Species |
Taxom
| | Subfamily | Rutelinae
MacLeay, 1819
Shining Flower Chafers, Shining Leaf Chafers | 19
47 | Genera
Species |
Taxom
| | Subfamily | Scarabaeinae
Latreille, 1802
Dung Beetles, True dung beetles | 41
210 | Genera
Species |
Taxom
|
 | | Scarabaeidae (Latreille 1802) |
Links, sources and further information
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