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TAXFAMGENSPESUB
Checklist for Family Scarabaeidae Latreille, 1802 (Dung beetles)
and region Spain (Iberian Peninsula)


Class
 
Insecta
Linnaeus, 1758
Insects, True insects
679
9809
32121
Families
Genera
Species
 
Taxom
 

Subclass
 
Pterygota
Lang, 1888
Winged insects
675
9790
32064
Families
Genera
Species
 
Taxom
 

Infraclass
 
Neoptera
van der Wulp, 1890
Modern Wing-folding insects
642
9663
31803
Families
Genera
Species
 
Taxom
 

Superorder
 
Holometabola
Heider, 1889
Holometabolous Insects
416
7897
27351
Families
Genera
Species
 
Taxom
 

Order
 
Coleoptera
Linnaeus, 1758
Beetles, Coleopterans
118
2951
11883
Families
Genera
Species
 
Taxom
 

Suborder
 
Polyphaga
Emery, 1886
Polyphags
105
2579
9709
Families
Genera
Species
 
Taxom
 

Infraorder
 
Scarabaeiformia
Crowson, 1960
12
260
933
Families
Genera
Species
 
Taxom
 

Superfamily
 
Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles
12
260
933
Families
Genera
Species
 
Taxom
 
FamilyScarabaeidae
Latreille, 1802
Dung beetles, Flower beetles, Fruit chafers, Rain beetles
213
795
Genera
Species
 
Taxom
 


Genera for Spain
213 Genera with 795 Species
#ABCDEFGHILMNOPRSTUVX
Acanthobodilus Dellacasa, 17991SpeciesTaxom
Acrossus Mulsant, 17585SpeciesTaxom
Adoretus Laporte, 19141SpeciesTaxom
Aegialia Latreille, 17871SpeciesTaxom
Aethiessa Burmeister, 02SpeciesTaxom
Agoliinus Schmidt, 1913
0SpeciesTaxom
Agolius Mulsant & Rey, 19815SpeciesTaxom
Agrilinus Mulsant & Rey, 09SpeciesTaxom
Ahermodontus Baguena, 19302SpeciesTaxom
Alocoderus Schmidt, 17982SpeciesTaxom
Amadotrogus Amadotrogus, 18422SpeciesTaxom
Amaladera Reitter, 19973SpeciesTaxom
Amidorus Mulsant & Rey, 18006SpeciesTaxom
Ammoecius Mulsant, 18486SpeciesTaxom
Amphiceratodon Huchet, 20021SpeciesTaxom
Amphimallon Lepeletier &..., 1823European Chafers29SpeciesTaxom
Ancognatha Erichson, 19111SpeciesTaxom
Anisoplia Schönherr, 178710SpeciesTaxom
Anomala Samouelle, 1844Shining Leaf Chafer18SpeciesTaxom
Anomius Mulsant & Rey, 18033SpeciesTaxom
Anoxia Laporte, 18608SpeciesTaxom
Antemnacrassa Gomez Pallerola, 19791SpeciesTaxom
Anthoplia Medvedev, 18472SpeciesTaxom
Aphodius Hellwig, 197584SpeciesTaxom
Apotriodonta Baraud, 19621SpeciesTaxom
Archophileurus Kolbe, 19101SpeciesTaxom
Aschnarhyparus Makhan, 19341SpeciesTaxom
Aspidolea Bates, 18883SpeciesTaxom
Ataenius Harold, 18879SpeciesTaxom
Ateuchetus Bedel, 17875SpeciesTaxom
Ateuchus Weber, 18862SpeciesTaxom
Barybas Burmeister, 1855
0SpeciesTaxom
Bdelyropsis Pereira, Vul..., 19761SpeciesTaxom
Biralus Mulsant & Rey, 17892SpeciesTaxom
Blitopertha Reitter, 18882SpeciesTaxom
Bodiloides3SpeciesTaxom
Bodilopsis Adám, 05SpeciesTaxom
Bodilus Mulsant & Rey, 17994SpeciesTaxom
Brindalus Landin, 19803SpeciesTaxom
Bubas Mulsant, 18113SpeciesTaxom
Caccobius Thomson, 17672SpeciesTaxom
Calamosternus Motschulsky, 17895SpeciesTaxom
Calicnemis Castelnau, 07SpeciesTaxom
Callistethus Blanchard, 20152SpeciesTaxom
Canthidium Erichson, 19814SpeciesTaxom
Canthon Hoffmannsegg, 1887Tumblebug4SpeciesTaxom
Ceramida Baraud, 190213SpeciesTaxom
Ceraspis Serville, 1825
0SpeciesTaxom
Cetonia Fabricius, 17878SpeciesTaxom
Chasmatopterus Latreille, 192516SpeciesTaxom
Cheironitis van Lansberge, 17896SpeciesTaxom
Chilothorax Motschulsky, 180513SpeciesTaxom
Chironitis Lansberge, 17903SpeciesTaxom
Chlaenobia Blanchard, 19921SpeciesTaxom
Coelosis Hope, 17671SpeciesTaxom
Colobopterus Mulsant, 17581SpeciesTaxom
Coprimorphus Mulsant, 17891SpeciesTaxom
Copris Geoffroy, 1758Mexican dung beetle4SpeciesTaxom
Coprophanaeus Olsoufieff, 18473SpeciesTaxom
Cotinis Burmeister, 1833Green June Beetles2SpeciesTaxom
Cyclocephala Dejean, 1847Masked Chafers18SpeciesTaxom
Cyphonotus Bell, 01SpeciesTaxom
Deltochilum Eschscholtz, 18871SpeciesTaxom
Dendropaemon Perty, 1830
0SpeciesTaxom
Dialytes Harold, 18451SpeciesTaxom
Diastictus Mulsant, 18051SpeciesTaxom
Dichotomius Hope, 18676SpeciesTaxom
Diplognathoides Holm & Peris..., 2004
0SpeciesTaxom
Diplotaxis Kirby, 1889Wall-Rockets1SpeciesTaxom
Disphysema Harold, 18731SpeciesTaxom
Dynastes MacLeay, 1758Hercules Beetles1SpeciesTaxom
Dyscinetus Harold, 1847Rice Beetles3SpeciesTaxom
Elaphocera Géné, 190227SpeciesTaxom
Epirinus Reiche, 17751SpeciesTaxom
Eremazus Mulsant, 18511SpeciesTaxom
Erioscelis Burmeister, 19661SpeciesTaxom
Erytus Mulsant & Rey, 18922SpeciesTaxom
Esymus Mulsant & Rey, 18834SpeciesTaxom
Eudolus Mulsant & Rey, 17831SpeciesTaxom
Euetheola Bates, 18472SpeciesTaxom
Euheptaulacus Dellacasa, 17833SpeciesTaxom
Euoniticellus Janssens, 17814SpeciesTaxom
Euonthophagus Balthasar, 17907SpeciesTaxom
Euorodalus Dellacasa, 18485SpeciesTaxom
Euphoria Burmeister, 18333SpeciesTaxom
Eupleurus Mulsant, 17582SpeciesTaxom
Eupotosia Miksic, 18423SpeciesTaxom
Europtron Marseul, 18781SpeciesTaxom
Eurysternus Dalman, 18692SpeciesTaxom
Euserica Rietter, 19758SpeciesTaxom
Exomala Reitter, 18041SpeciesTaxom
Firminus1SpeciesTaxom
Genuchinus Westwood, 1874
0SpeciesTaxom
Gnorimus Le Peletier ..., 1758Chafer3SpeciesTaxom
Golofa Hope, 18371SpeciesTaxom
Gymnetis MacLeay, 18131SpeciesTaxom
Gymnopleurus Illiger, 18217SpeciesTaxom
Haplidia Hope, 19711SpeciesTaxom
Haroldiellus Gordon and S..., 18631SpeciesTaxom
Hemiphileurus Kolbe, 18471SpeciesTaxom
Heptaulacus Mulsant, 17754SpeciesTaxom
Heterogomphus Burmeister, 1847
0SpeciesTaxom
Heterorhina Westwood, 17811SpeciesTaxom
Heterorrhina Westwood, 17811SpeciesTaxom
Holcorobeus Nikritin, 19791SpeciesTaxom
Homaloplia Stephens, 17751SpeciesTaxom
Homophileurus Kolbe, 18061SpeciesTaxom
Hoplia Illiger, 1849Monkey Beetles16SpeciesTaxom
Hoplopyga Thomson, 17891SpeciesTaxom
Hybalus Brullé, 18527SpeciesTaxom
Hymenochelus Reitter, 18761SpeciesTaxom
Hymenoplia Eschscholtz, 180319SpeciesTaxom
Hypotrichia LeConte, 1862
Hypotrichs0SpeciesTaxom
Isonychus Mannerheim, 19181SpeciesTaxom
Labarrus Mulsant and Rey, 02SpeciesTaxom
Leucothyreus MacLeay, 18441SpeciesTaxom
Limarus Mulsant & Rey, 18131SpeciesTaxom
Liothorax Motschulsky, 17673SpeciesTaxom
Loraspis Mulsant & Rey, 18701SpeciesTaxom
Macraspis MacLeay, 17642SpeciesTaxom
Macrodactylus Dejean, 1887Rose Chafers1SpeciesTaxom
Maladera Mulsant, 19974SpeciesTaxom
Mecynodes Mulsant & Rey, 18352SpeciesTaxom
Megasoma Kirby, 1785Elephant beetles2SpeciesTaxom
Megatelus Reitter, 18451SpeciesTaxom
Melinopterus Mulsant, 197511SpeciesTaxom
Melolontha Fabricius, 1803Billy witch9SpeciesTaxom
Mendidaphodius Reitter, 19632SpeciesTaxom
Mendidius Harold, 18812SpeciesTaxom
Mimela Kirby, 18491SpeciesTaxom
Mimeoma Casey, 19021SpeciesTaxom
Monotropus Erichson, 18614SpeciesTaxom
Neagolius Kozhantschikov, 19385SpeciesTaxom
Netocia A. Costa, 18983SpeciesTaxom
Nialus Mulsant & Rey, 18051SpeciesTaxom
Nimbus Mulsant & Rey, 19847SpeciesTaxom
Nobiellus1SpeciesTaxom
Nobius Mulsant & Rey, 18921SpeciesTaxom
Odontolytes Koshantschikov, 18641SpeciesTaxom
Omaloplia Schönherr, 17753SpeciesTaxom
Oniticellus Serville, 18073SpeciesTaxom
Onitis Fabricius, 17894SpeciesTaxom
Onthophagus Latreille, 186451SpeciesTaxom
Ootoma Blanchard, 18381SpeciesTaxom
Oromus Mulsant & Rey, 17631SpeciesTaxom
Orphnus MacLeay, 18971SpeciesTaxom
Oryctes Hellwig, 1864Coconut rhinoceros beetles4SpeciesTaxom
Osmoderma Le Peletier ..., 17631SpeciesTaxom
Otophorus Mulsant, 17581SpeciesTaxom
Oxyomus Dejean, 17631SpeciesTaxom
Oxythyrea Mulsant, 17611SpeciesTaxom
Pachydema Laporte, 192714SpeciesTaxom
Paleira Reiche, 18031SpeciesTaxom
Parabodilus1SpeciesTaxom
Parammoecius Seidlitz, 18632SpeciesTaxom
Paratriodonta Baraud, 18901SpeciesTaxom
Passaliolla Balthasar, 18871SpeciesTaxom
Pelidnota MacLeay, 1840Precious Metal Scarabs6SpeciesTaxom
Pentodon Hope, 19026SpeciesTaxom
Phalacronothus Motschulsky, 17613SpeciesTaxom
Phanaeus MacLeay, 1887Burrowing Dung Beetles1SpeciesTaxom
Pharaphodius Reitter, 17811SpeciesTaxom
Phileurus Sieg, 17582SpeciesTaxom
Phyllognathus Eschscholtz, 17711SpeciesTaxom
Phyllopertha Stephens, 19132SpeciesTaxom
Phyllophaga Harris, 1888June beetles8SpeciesTaxom
Plagiogonus Mulsant, 18602SpeciesTaxom
Planolinoides Dellacasa an..., 18271SpeciesTaxom
Planolinus Mulsant and Rey, 17892SpeciesTaxom
Platytomus Mulsant, 17982SpeciesTaxom
Pleurophorus Mulsant, 19614SpeciesTaxom
Polyphylla Harris, 1871Lined June Beetles3SpeciesTaxom
Popillia Leach, 18381SpeciesTaxom
Potosia Mulsant & Rey, 17874SpeciesTaxom
Protaetia Burmeister, 199227SpeciesTaxom
Psammodius Fallen, 18926SpeciesTaxom
Pseudacrossus Reitter, 18741SpeciesTaxom
Pseudesymus Orbigny, 18451SpeciesTaxom
Pseudotrochalus Quedenfeldt, 18011SpeciesTaxom
Pubinus Mulsant & Rey, 17761SpeciesTaxom
Rhizotrogus Lepeletier &..., 195528SpeciesTaxom
Rhodaphodius1SpeciesTaxom
Rhyssemodes Reitter, 18741SpeciesTaxom
Rhyssemus Mulsant, 184212SpeciesTaxom
Scabrostomus Gordon and S..., 19911SpeciesTaxom
Scarabaeolus Balthasar, 18681SpeciesTaxom
Scarabaeus Linnaeus, 177811SpeciesTaxom
Serica MacLeay, 18953SpeciesTaxom
Sericesthis Hope, 19871SpeciesTaxom
Sigorus Mulsant & Rey, 17921SpeciesTaxom
Sinodrepanus Simonis, 18751SpeciesTaxom
Sisyphus Latreille, 17583SpeciesTaxom
Stenocrates Burmeister, 19851SpeciesTaxom
Strategus Kirby, 1847Ox Beetles2SpeciesTaxom
Strigoderma Burmeister, 18881SpeciesTaxom
Subrinus Mulsant & Rey, 18452SpeciesTaxom
Temnorhynchus Hope, 18561SpeciesTaxom
Temnorrhynchus Alluaud, 18561SpeciesTaxom
Teuchestes Mulsant, 17582SpeciesTaxom
Tomarus Erichson, 1888Carrot Beetles4SpeciesTaxom
Trichius Fabricius, 1829Bee beetles6SpeciesTaxom
Trichonotulus Bedel, 17871SpeciesTaxom
Trigonopeltastes Burmeister &..., 20162SpeciesTaxom
Triodontella Reitter, 19786SpeciesTaxom
Triodontus Westwood, 18681SpeciesTaxom
Trochalus Laporte, 19171SpeciesTaxom
Tropinota Mulsant, 20186SpeciesTaxom
Uroxys Westwood, 18871SpeciesTaxom
Valgus Scriba, 02SpeciesTaxom
Volinus Mulsant & Rey, 17981SpeciesTaxom
Xeropsamobeus Saylor, 19071SpeciesTaxom
Xylonychus Blanchard, 19241SpeciesTaxom
Xylotrupes Hope, 1920Elephant beetles2SpeciesTaxom


Species list for Spain
795 Species and 96 Subspecies
Acanthobodilus immundus Creutzer, 1799
Acrossus atratus Waterhouse, 1875
Acrossus carpetanus Graells, 1846
Acrossus depressus Kugelann, 1792
Acrossus luridus Fabricius, 1775
Acrossus rufipes Linnaeus, 1758Night-Flying Dung Beetle
Adoretus (Adoretus) lithobius Ohaus, 1914
Aegialia (Aegialia) arenaria Fabricius, 1787Dune sand scarab beetle
Aethiessa floralis Fabricius, 1787
Aethiessa floralis floralis
Agolius abdominalis Bonelli, 1812
Agolius abdominalis abdominalis Bonelli, 1812
Agolius bonvouloiri Harold, 1860
Agolius bonvouloiri bonvouloiri
Agolius bonvouloiri cantabricus Pittino, 1981
Agrilinus aquilus Schmidt, 1916
Agrilinus ater De Geer, 1774
Agrilinus ater ascendens Reiche, 1863
Agrilinus ater lucasi Harold, 1859
Agrilinus bellus (Schmidt, 1916)⇒ Agrilinus aquilus Schmidt, 1916
Agrilinus constans Duftschmid, 1805
Agrilinus convexus Erichson, 1848
Agrilinus ibericus Harold, 1874
Agrilinus ibericus ibericus
Ahermodontus ambrosi Pardo Alcaide, 1936
Ahermodontus marini Baguena, 1930
Alocoderus carinifrons Reitter, 1892
Alocoderus hydrochaeris Fabricius, 1798
Amadotrogus patruelis Reiche, 1862
Amadotrogus vicinus Mulsant, 1842
Amaladera cobosi Baraud, 1964
Amaladera espagnoli Baraud, 1964
Amaladera longipennis Verdu & Mico..., 1997
Amidorus cribrarius Brullé, 1832
Amidorus cribricollis Lucas, 1846
Amidorus immaturus Mulsant, 1842
Amidorus obscurus Fabricius, 1792
Amidorus obscurus obscurus Fabricius, 1792
Amidorus thermicola Sturm, 1800
Ammoecius brevis Erichson, 1848
Ammoecius dentatus Schmidt, 1908
Ammoecius elevatus Olivier, 1789
Ammoecius frigidus Brisout, 1866
Ammoecius lugubris Boheman, 1857
Ammoecius lusitanicus Erichson, 1848
Amphiceratodon arrowi crypticus Huchet, 2002
Amphimallon (Amphimallon) assimile Herbst, 1790
Amphimallon (Amphimallon) atrum Herbst, 1790
Amphimallon atrum atrum Herbst, 1790
Amphimallon atrum nomadicum Reiche, 1862
Amphimallon (Amphimallon) majale Razoumowsky, 1789European Chafer
Amphimallon (Amphimallon) ochraceum (Knoch, 1801)⇒ Amphimallon fallenii Gyllenhall, 1817
Amphimallon (Amphimallon) pini Olivier, 1789
Amphimallon pini diluticolle Baraud & Tauzin, 1987
Amphimallon pini pini
Amphimallon (Amphimallon) solstitiale Linnaeus, 1758Cliff Beetle
Amphimallon solstitiale solstitiale Linnaeus, 1758
Amphimallon (Amphimallon) solstitialis (Linnaeus, 1758)⇒ Amphimallon volgense Fischer, 1823
Amphimallon cantabricum Heyden, 1870
Amphimallon fallenii Gyllenhall, 1817
Amphimallon fissiceps Fairmaire, 1860
Amphimallon ilustre López Colón ..., 2022
Amphimallon lusitanicum Gyllenhal, 1817
Amphimallon menori Baguena, 1955
Amphimallon naceyroi Mulsant, 1859
Amphimallon nigrum Waltl, 1835
Amphimallon nigrum ebeninum Baraud, 1975
Amphimallon nigrum nigrum
Amphimallon pygiale Mulsant, 1846
Amphimallon roris Baraud, 1981
Amphimallon sainzi Graells, 1852
Amphimallon seidlitzi Brenske, 1891
Amphimallon variolatus Fairmaire, 1880
Amphimallon vivesi Baraud, 1967
Amphimallon volgense Fischer, 1823
Ancognatha vulgaris Arrow, 1911
Anisoplia (Anisoplia) baetica Erichson, 1847
Anisoplia (Anisoplia) depressa Erichson, 1847
Anisoplia (Anisoplia) faldermanni Reitter, 1883
Anisoplia (Anisoplia) farraria Erichson, 1847
Anisoplia (Anisoplia) remota Reitter, 1889
Anisoplia (Anisoplia) tempestiva Erichson, 1847
Anisoplia (Anisoplia) villosa (Goeze, 1777)⇒ Anisoplia faldermanni Reitter, 1883
Anisoplia (Autanisoplia) austriaca (Herbst, 1783)⇒ Anisoplia tempestiva Erichson, 1847
Anisoplia arvicola Fabricius, 1781
Anisoplia floricola Fabricius, 1787
Anomala (Anomala) ausonia Erichson, 1847
Anomala (Anomala) dubia (Scopoli, 1763)⇒ Anomala undulata Melsheimer, 1845
Anomala dubia dubia
Anomala (Dichomala) devota Rossi, 1790
Anomala adscita Robinson, 1941
Anomala chiriquina Bates, 1888
Anomala clathrata Ohaus, 1930
Anomala gemella Say, 1835
Anomala hispidula Bates, 1888
Anomala longisacculata Filippini, M..., 2015
Anomala oxylabis Ohaus, 1916
Anomala pseudoeucoma Filippini, M..., 2013
Anomala quadripunctata Olivier, 1789
Anomala solisi Filippini, G..., 2014
Anomala sulana Zorn, 2006
Anomala testaceipennis (Blanchard, 1851)⇒ Anomala nitidula Blanchard, 1851
Anomala undulata Melsheimer, 1845Blossom anomala
Anomala valida (Burmeister, 1844)⇒ Anomala insularis Castelnau, 1840
Anomius annamariae Baraud, 1982
Anomius baeticus Mulsant & Rey, 1869
Anomius castaneus Illiger, 1803
Anoxia scutellaris scutellaris Mulsant, 1842
Anoxia (Anoxia) villosa Fabricius, 1781
Anoxia villosa villosa Fabricius, 1781
Anoxia (Mesanoxia) australis Gyllenhal, 1817
Anoxia matutinalis sardoa Motschulsky, 1861
Anoxia emarginata Coquerel, 1860
Anoxia rattoi Escalera, 1906
Anoxia sardoa (Motschulsky, 1860)⇒ Anoxia matutinalis sardoa Motschulsky, 1861
Antemnacrassa monreali J. E. Gomez ..., 1979
Anthoplia (Anthoplia) floricola Fabricius, 1787
Anthoplia floricola floricola Gistl, 1847
Aphodius (Acanthobodilus) immundus (Creutzer, 1799)⇒ Pubinus tomentosus O.F.Muller, 1776
Aphodius (Acrossus) carpetanus Graells, 1847
Aphodius (Acrossus) depressus (Kugelann, 1792)⇒ Acrossus depressus Kugelann, 1792
Aphodius (Acrossus) luridus (Fabricius, 1775)⇒ Acrossus luridus Fabricius, 1775
Aphodius (Acrossus) rufipes (Linnaeus, 1758)⇒ Acrossus rufipes Linnaeus, 1758
Aphodius (Agrilinus) ater (DeGeer, 1774)⇒ Agrilinus ater De Geer, 1774
Aphodius (Agrilinus) constans (Duftschmid, 1805)⇒ Agrilinus constans Duftschmid, 1805
Aphodius (Agrilinus) rufus Moll, 1782
Aphodius (Ahermodontus) marini Baguena, 1930
Aphodius (Amidorus) immaturus (Mulsant, 1842)⇒ Amidorus immaturus Mulsant, 1842
Aphodius (Amidorus) obscurus (Fabricius, 1792)⇒ Amidorus obscurus Fabricius, 1792
Aphodius (Ammoecius) brevis (Erichson, 1848)⇒ Ammoecius brevis Erichson, 1848
Aphodius (Ammoecius) elevatus (Olivier, 1789)⇒ Ammoecius elevatus Olivier, 1789
Aphodius (Anomius) castaneus (Illiger, 1803)⇒ Anomius castaneus Illiger, 1803
Aphodius (Aphodius) coniugatus Panzer, 1795
Aphodius (Aphodius) conjugatus (Mulsant, 1842)⇒ Aphodius coniugatus Panzer, 1795
Aphodius (Aphodius) fimetarius Linnaeus, 1758Common dung beetle
Aphodius (Aphodius) foetens (Fabricius, 1787)⇒ Rhodaphodius foetens Fabricius, 1787
Aphodius (Aphodius) foetidus Herbst, 1783
Aphodius (Biralus) satellitius (Herbst, 1789)⇒ Biralus satellitius Herbst, 1789
Aphodius (Bodilus) ictericus (Laicharting, 1781)⇒ Bodiloides ictericus Laicharting, 1781
Aphodius (Bodilus) longispina Küster, 1854
Aphodius (Bodilus) lugens (Creutzer, 1799)⇒ Bodilus lugens Creutzer, 1799
Aphodius (Bodilus) rufus (Moll, 1782)⇒ Bodilopsis rufa Moll, 1782
Aphodius (Bodilus) wollastoni Harold, 1862
Aphodius (Calamosternus) granarius (Linnaeus, 1767)⇒ Calamosternus granarius Linnaeus, 1767
Aphodius (Calamosternus) hyxos Petrovitz, 1962
Aphodius (Calamosternus) mayeri (Pilleri, 1953)⇒ Calamosternus mayeri Pilleri, 1953
Aphodius (Calamosternus) unicolor (Lucas, 1846)⇒ Anomius castaneus Illiger, 1803
Aphodius (Chilothorax) cervorum (Fairmaire, 1871)⇒ Chilothorax cervorum Fairmaire, 1871
Aphodius (Chilothorax) distinctus O.F. Müller, 1776Maculated Dung Beetle
Aphodius (Chilothorax) hieroglyphicus (Klug, 1845)⇒ Chilothorax hieroglyphicus Klug, 1845
Aphodius (Chilothorax) lineolatus (Illiger, 1803)⇒ Chilothorax lineolatus Illiger, 1803
Aphodius (Chilothorax) paykulli Bedel, 1907
Aphodius (Chilothorax) pictus (D. Koshantsc..., 1894)⇒ Chilothorax pictus Sturm, 1805
Aphodius (Chilothorax) sticticus (Panzer, 1798)⇒ Volinus sticticus Panzer, 1798
Aphodius (Colobopterus) erraticus (Linnaeus, 1758)⇒ Colobopterus erraticus Linnaeus, 1758
Aphodius (Coprimorphus) scrutator (Herbst, 1789)⇒ Coprimorphus scrutator Herbst, 1789
Aphodius (Esymus) merdarius (Fabricius, 1775)⇒ Esymus merdarius Fabricius, 1775
Aphodius (Eudolus) quadriguttatus (Herbst, 1783)⇒ Eudolus quadriguttatus Herbst, 1783
Aphodius (Euorodalus) coenosus (Panzer, 1798)⇒ Euorodalus coenosus Panzer, 1798
Aphodius (Euorodalus) paracoenosus (Balthasar & ..., 1960)⇒ Euorodalus paracoenosus Balthasar & ..., 1960
Aphodius (Euorodalus) pusillus (Herbst, 1789)⇒ Esymus pusillus Herbst, 1789
Aphodius (Euorodalus) tersus (Rosenhauer, 1856)⇒ Euorodalus tersus Erichson, 1848
Aphodius (Eupleurus) subterraneus (Linnaeus, 1758)⇒ Eupleurus subterraneus Linnaeus, 1758
Aphodius (Lorapsis) frater (Mulsant & Rey, 1869)⇒ Loraspis frater Mulsant & Rey, 1870
Aphodius (Melinopterus) consputus (Creutzer, 1799)⇒ Melinopterus consputus Creutzer, 1799
Aphodius (Melinopterus) prodromus (Braham, 1790)⇒ Melinopterus prodromus Brahm, 1790
Aphodius (Melinopterus) reyi (Reitter, 1892)⇒ Melinopterus reyi Reitter, 1892
Aphodius (Melinopterus) sphacelatus (Panzer, 1798)⇒ Melinopterus sphacelatus Panzer, 1798
Aphodius (Melinopterus) tingens (Reitter, 1892)⇒ Melinopterus tingens Reitter, 1892
Aphodius (Neagolius) heydeni Harold, 1871
Aphodius (Nialus) lividus (Olivier, 1789)⇒ Acrossus luridus Fabricius, 1775
Aphodius (Nialus) sturmi (Harold, 1870)⇒ Subrinus sturmi Harold, 1870
Aphodius (Nialus) varians (Duftschmid, 1805)⇒ Nialus varians Duftschmid, 1805
Aphodius (Nimbus) affinis (Lucas, 1846)⇒ Agrilinus ater De Geer, 1774
Aphodius (Nimbus) contaminatus (Herbst, 1783)⇒ Nimbus contaminatus Herbst, 1783
Aphodius (Nimbus) obliteratus (Panzer, 1823)⇒ Nimbus obliteratus Panzer, 1823
Aphodius (Nobius) bonnairei (Reitter, 1892)⇒ Nobiellus bonnairei Reitter, 1892
Aphodius (Oromus) alpinus (Scopoli, 1763)⇒ Euheptaulacus carinatus carinatus Germar, 1824
Aphodius (Otophorus) haemorrhoidalis (Linnaeus, 1758)⇒ Otophorus haemorrhoidalis Linnaeus, 1758
Aphodius (Parammoecius) corvinus (Haldeman, 1848)⇒ Parammoecius corvinus Erichson, 1848
Aphodius (Phalocronothus) biguttatus (Dallatorre, 1879)⇒ Phalacronothus biguttatus Germar, 1824
Aphodius (Phalocronothus) quadrimaculatus (Linnaeus, 1761)⇒ Teuchestes brachysomus Solsky, 1874
Aphodius (Planolinoides) borealis (Gyllenhal, 1827)⇒ Planolinoides borealis Gyllenhal, 1827
Aphodius (Planolinus) uliginosus (Hardy, 1847)⇒ Planolinus fasciatus Olivier, 1789
Aphodius (Pseudacrossus) thermicola (Sturm, 1800)⇒ Amidorus thermicola Sturm, 1800
Aphodius (Pseudocalamosternus) rigidus Balthasar, 1936
Aphodius (Sigorus) porcus (Fabricius, 1792)⇒ Sigorus porcus Fabricius, 1792
Aphodius (Teuchestes) fossor (Linnaeus, 1758)⇒ Teuchestes fossor Linnaeus, 1758
Aphodius (Trichonotulus) scrofa (Fabricius, 1787)⇒ Trichonotulus scrofa Fabricius, 1787
Aphodius abeillei Sietti, 1903
Aphodius annamariae Baraud, 1982
Aphodius asellus Schmidt, 1907
Aphodius baeticus Mulsant & Rey, 1870
Aphodius baraudi (Villarreal, 1975)⇒ Euorodalus boiteli Théry, 1918
Aphodius bonvouloiri (Harold, 1860)⇒ Agolius bonvouloiri Harold, 1860
Aphodius cardinalis (Reitter, 1892)⇒ Aphodius fimetarius Linnaeus, 1758
Aphodius cylindricus (Dejean, 1833)⇒ Mendidaphodius linearis Reiche & Saulcy, 1856
Aphodius nanus (Horn, 1887)⇒ Aphodius asellus Schmidt, 1907
Aphodius palmetincolus (Karsch, 1881)⇒ Mendidius palmetincola Karsch, 1881
Aphodius scalaris (Dellacasa, 1983)⇒ Chilothorax conspurcatus Linnaeus, 1758
Aphodius sharpi Harold, 1874
Aphodius villarreali Baraud, 1975
Apotriodonta hispanica Baraud, 1962
Archophileurus chaconus Kolbe, 1910
Aschnarhyparus peregrinus Hinton, 1934
Aspidolea fuliginea Burmeister, 1847
Aspidolea notaticollis Höhne, 1922
Aspidolea singularis Bates, 1888
Ataenius brevicollis Wollaston, 1854
Ataenius complicatus Harold, 1869
Ataenius glabriventris Schmidt, 1911
Ataenius gracilis Melsheimer, 1845
Ataenius languidus Schmidt, 1911Antarctic dragonfishes
Ataenius picinus Harold, 1861Antarctic blennies
Ataenius platensis Blanchard, 1846
Ataenius pseudousingeri Galante, Ste..., 2003
Ataenius scalptifrons Bates, 1887
Ateuchetus cicatricosus Lucas, 1846
Ateuchetus laticollis Linnaeus, 1767
Ateuchetus puncticollis Latreille, 1819
Ateuchetus semipunctatus Fabricius, 1792
Ateuchetus variolosus Fabricius, 1787
Ateuchus aeneomicans Harold, 1868
Ateuchus rodriguezi Preudhomme d..., 1886
Bdelyropsis venezuelensis Howden, 1976
Biralus mahunkaorum Adám, 1983
Biralus satellitius Herbst, 1789
Blitopertha (Blitopertha) lineata (Fabricius, 1798)⇒ Blitopertha nigripennis Reitter, 1888
Blitopertha nigripennis Reitter, 1888
Bodiloides ictericus Laicharting, 1781
Bodiloides ictericus ghardimaouensis Balthasar, 1929
Bodiloides ictericus ictericus Laicharting, 1781
Bodilopsis aquila Schmidt, 1916
Bodilopsis aquilus Schmidt, 1916
Bodilopsis rufa Moll, 1782
Bodilopsis rufus Moll, 1782
Bodilopsis sordidus sordidus
Bodilus barbarus Fairmaire, 1860
Bodilus ictericus (Laicharting, 1781)⇒ Bodiloides ictericus Laicharting, 1781
Bodilus longispina Küster, 1854
Bodilus lugens Creutzer, 1799
Brindalus porcicollis Illiger, 1803
Brindalus rotundipennis Reitter, 1892
Brindalus schatzmayri Pittino, 1980
Bubas bison Linnaeus, 1767
Bubas bubaloides Janssens, 1938
Bubas bubalus Olivier, 1811
Caccobius histeroides Ménétriés, 1832
Caccobius schreberi Linnaeus, 1767
Calamosternus granarius Linnaeus, 1767Granular Small Dung Beetle
Calamosternus hyxos Petrovitz, 1962
Calamosternus mayeri Pilleri, 1953
Calamosternus trucidatus Harold, 1863
Calamosternus unicolor Olivier, 1789
Calicnemis atlantica (Mosconi, 1996)⇒ Calicnemis obesa Erichson, 1841
Calicnemis bahilloi Lopez-colon, 2003
Calicnemis bercedoi Lopez-colon, 2004
Calicnemis latreillei Laporte, 1832
Calicnemis obesa Erichson, 1841
Calicnemis truncatifrons Laporte, 1832
Calicnemis truncatifrons truncatifrons
Callistethus granulipygus Bates, 1888
Callistethus lativittis Filippini, G..., 2015
Canthidium ardens Bates, 1887
Canthidium hespenheidei Howden & Young, 1981
Canthidium perceptibile Howden & Young, 1981
Canthidium subdopuncticolle Howden & Young, 1981
Canthon (Glaphyrocanthon) femoralis Chevrolat, 1834
Canthon (Glaphyrocanthon) viridis (Palisot de B..., 1805)⇒ Epilissus viridis Klug, 1832
Canthon deyrollei Harold, 1868
Canthon euryscelis Bates, 1887
Ceramida adusta Kraatz, 1882
Ceramida baraudi (Branco, 1981)⇒ Ceramida longitarsis Illiger, 1803
Ceramida bedeaui Erichson, 1840
Ceramida cobosi Baguena, 1955
Ceramida jiennensis Sánchez Garc..., 2010
Ceramida longitarsis Illiger, 1803
Ceramida luisae (López-Colón ..., 1990)⇒ Ceramida zuzartei Branco, 1981
Ceramida luna Sánchez Garc..., 2010
Ceramida malaceensis Rambur, 1843
Ceramida malacensis Rambur, 1843
Ceramida mauritanica Rambur, 1843
Ceramida moelleri Flach, 1906
Ceramida ohausi Reitter, 1902
Cetonia aurata Linnaeus, 1758Flower beetle
Cetonia aurata aurata Linnaeus, 1761
Cetonia aurata pisana Heer, 1841
Cetonia carthami Gory & Perch..., 1833
Cetonia carthami aurataeformis Curti, 1913
Cetonia cuprea (Gory & Perch..., 1833)⇒ Heterorhina elegans Fabricius, 1781
Cetonia funeraria Gory & Perch..., 1833
Cetonia opaca (Fabricius, 1787)⇒ Potosia opaca opaca Fabricius, 1787
Chasmatopterus almeriensis Baraud, 1965
Chasmatopterus canosus Baguena, 1955
Chasmatopterus cobosi Baraud, 1965
Chasmatopterus cobosi barrancoi Ruiz & Avila, 1994
Chasmatopterus cobosi cobosi
Chasmatopterus esuris Baraud, 1965
Chasmatopterus hirtulus Illiger, 1803
Chasmatopterus hirtulus hirtulus
Chasmatopterus hirtus Blanchard, 1850
Chasmatopterus illigeri Perris, 1855
Chasmatopterus parvulus Graells, 1851
Chasmatopterus pilosulus Illiger, 1803
Chasmatopterus uxorum Baraud & Branco, 1985
Chasmatopterus villosulus Illiger, 1803
Chasmatopterus villosulus villosulus Illiger, 1803
Chasmatopterus zonatus Escalera, 1925
Cheironitis furcifer Rossi, 1792
Cheironitis hungaricus Herbst, 1789
Cheironitis irroratus Rossi, 1790
Cheironitis ungaricus Herbst, 1789
Cheironitis ungaricus irroratus (Rossi, 1790)⇒ Cheironitis irroratus Rossi, 1790
Cheironitis ungaricus ungaricus Herbst, 1789
Chilothorax aljibei Blanco, 1986
Chilothorax brancoi Baraud, 1981
Chilothorax cervorum Fairmaire, 1871
Chilothorax clathratus Reitter, 1892
Chilothorax conspurcatus Linnaeus, 1758
Chilothorax distinctus (Müller, 1776)⇒ Aphodius distinctus O.F. Müller, 1776
Chilothorax exclamationis Motschulsky, 1849
Chilothorax hieroglyphicus Klug, 1845
Chilothorax lineolatus Illiger, 1803
Chilothorax melanostictus Schmidt, 1840
Chilothorax melanostictus melanostictus W. Schmidt, 1840
Chilothorax paykulli Bedel, 1907
Chilothorax pictus Sturm, 1805
Chironitis furcifer Rossi, 1792
Chironitis hungaricus (Herbst, 1789)⇒ Cheironitis ungaricus Herbst, 1789
Chironitis irroratus Rossi, 1790
Chlaenobia halffteriana Morón, 1992
Coelosis biloba Linnaeus, 1767
Colobopterus erraticus Linnaeus, 1758Erratic small dung beetle
Coprimorphus scrutator Herbst, 1789
Copris hispanus Linnaeus, 1764
Copris hispanus hispanus Linné, 1764
Copris lugubris Boheman, 1858
Copris lunaris Linnaeus, 1758Horned Dung Beetle
Coprophanaeus boucardi Nevinson, 1891
Coprophanaeus corythus Harold, 1863
Coprophanaeus telamon Erichson, 1847
Cotinis (Cotinis) mutabilis Gory & Perch..., 1883Fig beetle
Cotinis lebasi (Gory & Perch..., 1833)⇒ Cotinorrhina lebasi Gory & Perch..., 1833
Cyclocephala almitana Dechambre, 1992
Cyclocephala amazona Linnaeus, 1767
Cyclocephala amblyopsis Hurley, 1959
Cyclocephala carbonaria Milne Edwards, 1848
Cyclocephala complanata Burmeister, 1847
Cyclocephala discolor Herbst, 1792
Cyclocephala erotylina Arrow, 1914
Cyclocephala gravis Bates, 1888
Cyclocephala guttata Bates, 1888
Cyclocephala ligyrina Bates, 1888
Cyclocephala lunulata Burmeister, 1847Twister
Cyclocephala macrophylla Cohen, 1998
Cyclocephala mafaffa Burmeister, 1847Twister
Cyclocephala melanocephala (Fabricius, 1775)⇒ Cyclocephala olivieri Arrow, 1911
Cyclocephala multiplex (Casey, 1915)⇒ Cyclocephala amazona Linnaeus, 1767
Cyclocephala ovulum (Bates, 1888)⇒ Cyclocephala testacea Burmeister, 1847
Cyclocephala stictica Burmeister, 1847
Cyclocephala testacea Burmeister, 1847Twister
Cyphonotus incertus
Deltochilum lobipes Bates, 1887
Dialytes truncatus Melsheimer, 1845
Diastictus vulneratus Sturm, 1805
Dichotomius (Luederwaldtinia) centralis Harold, 1869
Dichotomius (Luederwaldtinia) yucatanus Bates, 1887
Dichotomius annae Linnaeus, 1767
Dichotomius carolinus Linnaeus, 1767Carolina copris
Dichotomius colonicus (Say, 1835)⇒ Dichotomius carolinus Linnaeus, 1767
Dichotomius satanas Harold, 1867
Diplotaxis denigrata Bates, 1889
Disphysema candezei Harold, 1873
Dynastes hercules Linnaeus, 1758Hercules beetle
Dyscinetus dubius Olivier, 1789
Dyscinetus laevipunctatus Bates, 1888
Dyscinetus picipes Burmeister, 1847Spotted kelpfish
Elaphocera affinis Schaufuss, 1874
Elaphocera alonsoi Lopez-colon, 1992
Elaphocera alonsoi alonsoi
Elaphocera alonsoi semperei López-Colón, 1992
Elaphocera alonsoi semperi Lopez-colon, 1992
Elaphocera ampla Baguena, 1955
Elaphocera angusta Kraatz, 1882
Elaphocera autumnalis Motschulsky, 1859
Elaphocera baguenae Mancini, 1925
Elaphocera cacerensis Lopez Colon ..., 1986
Elaphocera capdeboni Schaufuss, 1879
Elaphocera capdeboui (Baguena, 1967)⇒ Elaphocera capdeboni Schaufuss, 1879
Elaphocera carteiensis Rambur, 1843
Elaphocera christina Baraud, 1966
Elaphocera churianensis Rambur, 1843
Elaphocera ferreri Lopez Colon ..., 1989
Elaphocera heydeni Kraatz, 1882
Elaphocera hirticollis Kraatz, 1882
Elaphocera hispalensis Rambur, 1843
Elaphocera ibicensis Escalera, 1926
Elaphocera lajonquierei Baraud, 1966
Elaphocera martini Lopez-colon, 1993
Elaphocera martorellii Fairmaire, 1879
Elaphocera nupcialis Ruiz & Avila, 1995
Elaphocera perezlopezi Ruiz, 1996
Elaphocera segurensis Escalera, 1923
Elaphocera tethys Reitter, 1902
Epirinus flagellatus Fabricius, 1775
Eremazus unistriatus Mulsant, 1851
Erioscelis columbica Endrödi, 1966
Erytus cognatus Fairmaire, 1860
Erytus pruinosus Reitter, 1892
Esymus merdarius Fabricius, 1775
Esymus pusillus Herbst, 1789
Esymus pusillus pusillus Herbst, 1789
Esymus sesquivittatus Fairmaire, 1883
Eudolus quadriguttatus Herbst, 1783
Euetheola bidentata Burmeister, 1847
Euetheola humilis Burmeister, 1847Spotted kelpfish
Euheptaulacus carinatus Germar, 1824
Euheptaulacus carinatus carinatus Germar, 1824
Euheptaulacus sus Herbst, 1783
Euoniticellus fulvus Goeze, 1777
Euoniticellus inaequalis Reiche, 1850
Euoniticellus pallens Olivier, 1789
Euoniticellus pallipes Fabricius, 1781Southern sandy dung beetle
Euonthophagus amyntas Olivier, 1789
Euonthophagus amyntas amyntas Olivier, 1789
Euonthophagus crocatus Mulsant & Go..., 1870
Euonthophagus crocatus crocatus
Euonthophagus dorbignyi Olsoufieff, 1900
Euonthophagus dorbignyi neutto Olsoufieff, 1900
Euonthophagus gibbosus Scriba, 1790
Euorodalus baraudi (Villarreal, 1975)⇒ Euorodalus boiteli Théry, 1918
Euorodalus boiteli Théry, 1918
Euorodalus coenosus Panzer, 1798
Euorodalus paracoenosus Balthasar & ..., 1960
Euorodalus tersus Erichson, 1848
Euphoria geminata Chevrolat, 1835
Euphoria iridescens Schaum, 1841
Euphoria leucographa (Gory & Perch..., 1833)⇒ Euphoria sepulcralis rufina Gory & Perch..., 1833
Eupleurus subterraneus Linnaeus, 1758Subterranean small dung beetle
Eupleurus subterraneus subterraneus Linnaeus, 1758
Eupotosia affinis (Fievet, 1995)⇒ Protaetia affinis Andersch, 1797
Eupotosia affinis tyrrenica (fannyae Fievet, 1995)⇒ Protaetia affinis tyrrenica Miksic, 1957
Eupotosia mirifica Mulsant, 1842
Europtron confusum Marseul, 1878
Eurysternus caribaeus Herbst, 1789
Eurysternus mexicanus Harold, 1869
Euserica (Euserica) mutata Gyllenhal, 1817
Euserica cambeloi Ruiz & Avila, 1993
Euserica lucipeta Baraud, 1965
Euserica mulsanti Brenske, 1902
Euserica paenibaeticae Galante, 1987
Euserica pauliani Lopez Colon, 1988
Euserica segurana Brenske, 1897
Euserica villarreali Baraud, 1975
Exomala (Neoblitopertha) campestris Latreille, 1804
Firminus alsasuanus Reitter, 1902
Gnorimus nobilis Linnaeus, 1758Noble Chafer
Gnorimus nobilis nobilis Linnaeus, 1758
Gnorimus variabilis Linnaeus, 1758Variable Chafer
Golofa pizarro Hope, 1837
Gymnetis stellata Latreille, 1813
Gymnopleurus coriarius (Herbst, 1789)⇒ Gymnopleurus flagellatus Fabricius, 1787
Gymnopleurus flagellatus Fabricius, 1787
Gymnopleurus flagellatus flagellatus Fabricius, 1787
Gymnopleurus mopsus Pallas, 1781
Gymnopleurus mopsus mopsus
Gymnopleurus sturmi (MacLeay, 1821)⇒ Gymnopleurus sturmii MacLeay, 1821
Gymnopleurus sturmii MacLeay, 1821
Haplidia akbesiana Petrovitz, 1971
Haroldiellus sallei Harold, 1863
Hemiphileurus variolosus Burmeister, 1847
Heptaulacus algarbiensis Branco & Baraud, 1984
Heptaulacus brancoi Baraud, 1976
Heptaulacus gadetinus Baraud, 1973
Heptaulacus testudinarius Fabricius, 1775
Heterorhina elegans Fabricius, 1781
Heterorrhina elegans Fabricius, 1781
Holcorobeus monreali Gomez Pallerola, 1979
Homaloplia (Homaloplia) ruricola Fabricius, 1775
Homophileurus quadrituberculatus Palisot de B..., 1806
Hoplia (Hoplia) callipyge LeConte, 1856Aquatic oligochaete worm
Hoplia argentea Poda, 1761
Hoplia aulica Linnaeus, 1767
Hoplia bilineata Fabricius, 1801
Hoplia chlorophana (Erichson, 1847)⇒ Hoplia bilineata Fabricius, 1801
Hoplia coerulea Drury, 1773
Hoplia farinosa ramburi Heyden, 1870
Hoplia freyi Baraud, 1967
Hoplia hungarica Burmeister, 1844
Hoplia korbi Petrovitz, 1958
Hoplia peroni Blanchard, 1850
Hoplia peronii C. É. Blanch..., 1850
Hoplia philanthus Füessly, 1775Welsh Chafer
Hoplia philanthus gagates Bedel, 1911
Hoplia philanthus philanthus Fuesslin, 1775
Hoplia pubicollis (Küster, 1849)⇒ Hoplia callipyge LeConte, 1856
Hoplopyga liturata Olivier, 1789
Hybalus alonsoi Lopez-Colon ..., 2014
Hybalus ameliae Lopez Colon, 1986
Hybalus ameliae baguenae Lopez Colon, 1986
Hybalus baguenae Lopez-colon, 1986
Hybalus dorcas (Germar, 1838)⇒ Hybalus graecus Sturm, 1843
Hybalus saezi Lopez-colon, 1992
Hybalus tingitanus Fairmaire, 1852
Hymenochelus distinctus Uhagon, 1876
Hymenoplia arragonica Escalera, 1927
Hymenoplia castiliana Reitter, 1890
Hymenoplia castilianissima Baguena, 1954
Hymenoplia chevrolati Mulsant, 1842
Hymenoplia cineracens Rosenhauer, 1856
Hymenoplia cinerascens Rosenhauer, 1856
Hymenoplia clypealis Reitter, 1902
Hymenoplia escalerai Baguena, 1954
Hymenoplia fulvipennis (Blanchard, 1850)⇒ Hymenoplia miegii Graells, 1858
Hymenoplia galaica Baguena, 1954
Hymenoplia jayenensis Baguena, 1954
Hymenoplia lata Heyden, 1870
Hymenoplia lineolata Blanchard, 1850
Hymenoplia miegii Graells, 1858
Hymenoplia minuta Baguena, 1955
Hymenoplia pardoalcaidei Lopez-Colon ..., 2014
Hymenoplia pseudocinerascens Baguena, 1954
Hymenoplia rugulosa Mulsant, 1842
Hymenoplia strigosa (Illiger, 1803)⇒ Hymenoplia chevrolati Mulsant, 1842
Isonychus vicinus Moser, 1918
Labarrus lividus Olivier, 1789Scarab beetle
Labarrus lividus lividus
Leucothyreus femoratus Burmeister, 1844
Limarus zenkeri Germar, 1813
Liothorax muscorum (Adám, 1994)⇒ Liothorax niger Illiger, 1798
Liothorax niger Illiger, 1798Beaulieu dung beetle
Liothorax plagiatus Linnaeus, 1767
Loraspis frater Mulsant & Rey, 1870
Macraspis aterrima Waterhouse, 1881
Macraspis chrysis Linnaeus, 1764
Macrodactylus sericinus Bates, 1887
Maladera (Maladera) holosericea Scopoli, 1772
Maladera cobosi Baraud, 1964
Maladera espagnoli Baraud, 1964
Maladera longipennis Verdú, Mico ..., 1997
Mecynodes leucopterus Klug, 1845
Mecynodes striatulus Waltl, 1835
Megasoma elephas Fabricius, 1775Elephant beetle
Megasoma gyas Herbst, 1785
Megatelus contractus Klug, 1845
Melinopterus abeillei Sietti, 1903
Melinopterus consputus Creutzer, 1799
Melinopterus dellacasai Avila, 1986
Melinopterus prodromus Brahm, 1790Spring small dung beetle
Melinopterus pubescens Sturm, 1800
Melinopterus punctatosulcatus Sturm, 1805
Melinopterus punctatosulcatus punctatosulcatus Sturm, 1805
Melinopterus reyi Reitter, 1892
Melinopterus sphacelatus Panzer, 1798
Melinopterus tingens Reitter, 1892
Melinopterus villarreali Baraud, 1975
Melolontha (Melolontha) melolontha Linnaeus, 1758Cockchafer
Melolontha baetica Hillert, Rös..., 2019
Melolontha gonzalezi López-Colón ..., 2020
Melolontha hidalgoi Hillert, Rös..., 2019
Melolontha hippocastani Fabricius, 1801Chestnut cockchafer
Melolontha hippocastani hippocastani Fabricius, 1801
Melolontha hybrida (T. Charpentier, 1825)⇒ Melolontha papposa Illiger, 1803
Melolontha linaresi Hillert, Rös..., 2019
Melolontha papposa Illiger, 1803
Mendidaphodius linearis Reiche & Saulcy, 1856
Mendidaphodius paganettii Petrovitz, 1963
Mendidius palmetincola Karsch, 1881
Mendidius palmetincolus Karsch, 1881
Mimela rugatipennis Graells, 1849
Mimeoma acuta Arrow, 1902
Monotropus brancoi Baraud, 1979
Monotropus laticollis Pérez-Arcas, 1874
Monotropus marquezi (Lopez-colon, 1986)⇒ Monotropus brancoi Baraud, 1979
Monotropus staudingeri Schaufuss, 1861
Neagolius heydeni Harold, 1871
Neagolius montanus Erichson, 1848
Neagolius schlumbergeri Seidlitz, 1891
Neagolius schlumbergeri schlumbergeri Seidlitz, 1888
Neagolius schlumbergeri temperei Hoffmann, 1938
Netocia morio Fabricius, 1781
Netocia morio morio Fabricius, 1781
Netocia oblonga (Reitter, 1898)⇒ Protaetia oblonga Gory & Perch..., 1833
Nialus varians Duftschmid, 1805
Nimbus affinis Panzer, 1823
Nimbus affinis affinis
Nimbus contaminatus Herbst, 1783
Nimbus marianii Pittino, 1978
Nimbus obliteratus Panzer, 1823
Nimbus proximus Adám, 1994
Nimbus richardi Veiga, 1984
Nobiellus bonnairei Reitter, 1892
Nobius bonnairei (Reitter, 1892)⇒ Nobiellus bonnairei Reitter, 1892
Odontolytes denominatus Chevrolat, 1864
Omaloplia (Omaloplia) alternata Küster, 1849
Omaloplia (Omaloplia) ruricola (Fabricius, 1775)⇒ Omaloplia alternata Küster, 1849
Omaloplia ruricola ruricola Fabricius, 1775
Oniticellus falsus (Sharp, 1875)⇒ Sinodrepanus falsus Sharp, 1875
Oniticellus pallipes (Fairmaire, 1887)⇒ Euoniticellus inaequalis Reiche, 1850
Oniticellus pictus Hausmann, 1807
Onitis alexis (Klug, 1835)⇒ Cheironitis ungaricus ungaricus Herbst, 1789
Onitis alexis septentrionalis Balthasar, 1942
Onitis belial Fabricius, 1798
Onitis ion Olivier, 1789
Onthophagus (Amphionthophagus) melitaeus Fabricius, 1798
Onthophagus (Furconthophagus) furcatus (Fabricius, 1781)⇒ Onthophagus furcaticeps Masters, 1886
Onthophagus (Micronthophagus) quaestus Sharp, 1875
Onthophagus (Onthophagus) batesi Howden and C..., 1963
Onthophagus (Onthophagus) illyricus Scopoli, 1763
Onthophagus (Onthophagus) landolti (Harold, 1880)⇒ Onthophagus schaefferi Howden and C..., 1963
Onthophagus (Onthophagus) taurus Schreber, 1759Bull Headed Dung Beetle
Onthophagus (Palaeonthophagus) albarracinus Baraud, 1979
Onthophagus (Palaeonthophagus) coenobita Herbst, 1783
Onthophagus (Palaeonthophagus) fracticornis (Preyssler, 1790)⇒ Onthophagus similis Scriba, 1790
Onthophagus (Palaeonthophagus) grossepunctatus Reitter, 1905
Onthophagus (Palaeonthophagus) joannae Goljan, 1953
Onthophagus (Palaeonthophagus) latigena d´Orbigny, 1897
Onthophagus (Palaeonthophagus) lemur Fabricius, 1781
Onthophagus (Palaeonthophagus) marginalis Gebler, 1817
Onthophagus (Palaeonthophagus) medius (Kugelann, 1792)⇒ Onthophagus vacca Linnaeus, 1767
Onthophagus (Palaeonthophagus) merdarius Chevrolat, 1865
Onthophagus (Palaeonthophagus) nuchicornis Linnaeus, 1758Mottled dung beetle
Onthophagus (Palaeonthophagus) opacicollis Reitter, 1893
Onthophagus (Palaeonthophagus) ovatus Linnaeus, 1767
Onthophagus (Palaeonthophagus) ruficapillus Brullé, 1832
Onthophagus ruficapillus ruficapillus Brullé, 1832
Onthophagus (Palaeonthophagus) semicornis Panzer, 1798
Onthophagus (Palaeonthophagus) similis (Scriba, 1790)⇒ Onthophagus fracticornis Preyssler, 1790
Onthophagus (Palaeonthophagus) stylocerus Graells, 1851
Onthophagus (Palaeonthophagus) trigibber Reitter, 1893
Onthophagus (Palaeonthophagus) vacca Linnaeus, 1767
Onthophagus (Palaeonthophagus) verticicornis (Laicharting, 1781)⇒ Onthophagus nuchicornis Linnaeus, 1758
Onthophagus (Paleonthophagus) andalusicus Waltl, 1835
Onthophagus andalusicus andalusicus
Onthophagus (Paleonthophagus) fracticornis Preyssler, 1790
Onthophagus (Paleonthophagus) nuchicornis Linnaeus, 1758
Onthophagus (Paleonthophagus) similis Scriba, 1790
Onthophagus (Paleonthophagus) vacca Linnaeus, 1767
Onthophagus (Parentius) emarginatus Mulsant, 1842
Onthophagus (Parentius) hispanicus Baraud, 1963
Onthophagus brucei cyanopterus (d´Orbigny, 1905)⇒ Proagoderus cyanopterus d´Orbigny, 1905
Onthophagus (Relictonthophagus) emarginatus (Mulsant, 1842)⇒ Onthophagus rubrimaculatus MacLeay, 1864
Onthophagus (Relictonthophagus) nigellus (Illiger, 1803)⇒ Caccobius histeroides Ménétriés, 1832
Onthophagus (Relictonthophagus) punctatus (Illiger, 1803)⇒ Onthophagus emarginatus Mulsant, 1842
Onthophagus (Trichonthophagus) hirtus Illiger, 1803
Onthophagus (Trichonthophagus) maki Illiger, 1803
Onthophagus carpophilus Pereira & Ha..., 1961
Onthophagus championi Bates, 1887
Onthophagus furcaticeps Masters, 1886
Onthophagus guatemalensis (Bates, 1887)⇒ Onthophagus medorensis Brown, 1929
Onthophagus lebasi Boucomont, 1932
Onthophagus malthinus Gillet, 1930
Onthophagus marginicollis Harold, 1880
Onthophagus praecellens Bates, 1887
Onthophagus rubrimaculatus MacLeay, 1864
Ootoma bipartita Gaspard Augu..., 1838
Oromus alpinus Scopoli, 1763
Orphnus distinctus (Fairmaire, 1897)⇒ Triodontus hova Fairmaire, 1868
Oryctes (Oryctes) nasicornis Linnaeus, 1758European rhinoceros beetle
Oryctes nasicornis ameliae López-Colón ..., 2020
Oryctes nasicornis grypus Illiger, 1803
Oryctes (Oryctes) prolixus Wollaston, 1864
Osmoderma eremita Scopoli, 1763Hermit Beetle
Otophorus haemorrhoidalis Linnaeus, 1758Bloody small dung beetle
Oxyomus sylvestris Scopoli, 1763Forest small dung beetle
Oxythyrea funesta Poda, 1761
Pachydema bipartita (Brullé, 1839)⇒ Ootoma bipartita Gaspard Augu..., 1838
Pachydema castanea Brullé, 1839
Pachydema fortunatorum Baraud, 1985
Pachydema fuscipennis Brullé, 1839
Pachydema gomerae Lopez-colon, 1999
Pachydema integra Wollaston, 1864
Pachydema lesnei Peyerimhoff, 1927
Pachydema megalops Mico & Galante, 2009
Pachydema menieri Baraud, 1985
Pachydema navatteae Baraud, 1985
Pachydema obscura Brullé, 1839
Pachydema obscurella Wollaston, 1864
Pachydema tinerfensis Galante & St..., 1992
Pachydema wollastoni Peyerimhoff, 1927
Paleira femorata Illiger, 1803
Parabodilus wollastoni Harold, 1862
Parammoecius corvinus Erichson, 1848
Parammoecius pyrenaeus Jacquelin du..., 1863
Paratriodonta alicantina Reitter, 1890
Passaliolla cossonoides Bates, 1887
Pelidnota belti Sharp, 1877
Pelidnota glabra Ohaus, 1922
Pelidnota notata (Blanchard, 1851)⇒ Pelidnota punctatus Linnaeus, 1758
Pelidnota prolixa Sharp, 1877
Pelidnota punctulata Bates, 1888
Pelidnota strigosa Laporte, 1840
Pentodon algerinum Herbst, 1789
Pentodon algerinus (Fuessly, 1778)⇒ Pentodon algerinum Herbst, 1789
Pentodon algerinus algerinus (Fuessly, 1778)⇒ Pentodon algerinum algerinum Herbst, 1789
Pentodon bidens punctatum Villers, 1789
Pentodon bidens punctatus (Villers, 1789)⇒ Pentodon bidens punctatum Villers, 1789
Pentodon idiota memnon Jakovlev, 1902
Phalacronothus biguttatus Germar, 1824
Phalacronothus putoni Reitter, 1894
Phalacronothus quadrimaculatus Linnaeus, 1761
Phanaeus (Notiophanaeus) pyrois Bates, 1887
Pharaphodius marginellus Fabricius, 1781
Phileurus didymus Linnaeus, 1758
Phileurus valgus Linnaeus, 1758
Phyllognathus excavatus Forster, 1771
Phyllopertha horticola (Linnaeus, 1758)⇒ Phyllopertha intermixta Arrow, 1913
Phyllopertha intermixta Arrow, 1913
Phyllophaga (Phyllophaga) panorpa Sanderson, 1950
Phyllophaga (Phyllophaga) setifera Burmeister, 1855Highfin blenny
Phyllophaga (Phytalus) obsoleta Blanchard, 1851
Phyllophaga caraga Saylor, 1943
Phyllophaga costaricensis Moser, 1918
Phyllophaga densata Moser, 1918
Phyllophaga elenans Saylor, 1938
Phyllophaga prolixa Bates, 1888
Plagiogonus arenarius Olivier, 1789
Plagiogonus nanus Fairmaire, 1860
Planolinoides borealis Gyllenhal, 1827
Planolinus borealis (Gyllenhal, 1827)⇒ Planolinoides borealis Gyllenhal, 1827
Planolinus fasciatus Olivier, 1789
Platytomus laevistriatus Perris, 1869
Platytomus tibialis Fabricius, 1798
Pleurophorus caesus Creutzer, 1796Cut small dung beetle
Pleurophorus maghrebinicus Pittino & Ma..., 1986
Pleurophorus mediterranicus Pittino & Ma..., 1986
Pleurophorus pannonicus Petrovitz, 1961
Polyphylla (Polyphylla) fullo (Linnaeus, 1758)⇒ Polyphylla olivieri Castelnau, 1840
Polyphylla (Polyphylla) olivieri Castelnau, 1840
Polyphylla gracilicornis Blanchard, 1871
Popillia japonica Newman, 1838Japanese beetle
Potosia cuprea (Mulsant, 1842)⇒ Protaetia brevitarsis Lewis, 1879
Potosia cuprea olivacea (Mulsant, 1842)⇒ Protaetia metallica bourgini Ruter, 1967
Potosia opaca Fabricius, 1787
Potosia opaca opaca Fabricius, 1787
Protaetia (Cetonischema) aeruginosa (Drury, 1770)⇒ Cetonischema speciossissima Scopoli, 1786
Protaetia (Eupotosia) affinis Andersch, 1797
Protaetia affinis affinis (Andersch, 1797)⇒ Eupotosia affinis affinis Andersch, 1797
Protaetia affinis tyrrenica Miksic, 1957
Protaetia (Eupotosia) mirifica (Mulsant, 1842)⇒ Eupotosia mirifica Mulsant, 1842
Protaetia mirifica mirifica (Mulsant, 1842)⇒ Eupotosia mirifica mirifica Mulsant, 1842
Protaetia (Liocola) brevitarsis Lewis, 1879
Protaetia (Macroprotaetia) distincta Antoine & Pa..., 1994
Protaetia (Macroprotaetia) gestroi Miksic, 1970
Protaetia (Netocia) cuprea (Fabricius, 1775)⇒ Protaetia distincta Antoine & Pa..., 1994
Protaetia cuprea brancoi Baraud, 1992
Protaetia cuprea ferreriesensis Compte-Sart ..., 2013
Protaetia cuprea hesperica Motschulsky, 1849
Protaetia cuprea incerta Costa, 1852
Protaetia cuprea olivacea Mulsant, 1842
Protaetia (Netocia) morio (Fabricius, 1781)⇒ Netocia morio Fabricius, 1781
Protaetia morio morio (Fabricius, 1781)⇒ Netocia morio morio Fabricius, 1781
Protaetia (Netocia) oblonga Gory & Perch..., 1833
Protaetia oblonga oblonga
Protaetia oblonga raffrayi Desbrochers ..., 1869
Protaetia (Netocia) opaca (Fabricius, 1787)⇒ Potosia opaca Fabricius, 1787
Protaetia opaca opaca Fabricius, 1787
Protaetia (Netocia) vidua Gory & Perch..., 1833
Protaetia cuprea viridiaurata Fuente, 1897
Protaetia (Potosia) metallica Herbst, 1782
Protaetia metallica bourgini Ruter, 1967
Protaetia metallica brancoi Baraud, 1992
Psammodius basalis Mulsant & Rey, 1870Crested blenny
Psammodius laevipennis Costa, 1844Awkward-winged small dung beetle
Psammodius pierottii Pittino, 1979Mud blenny
Psammodius plicicollis Erichson, 1848
Psammodius porcicollis (Fairmaire, 1870)⇒ Platytomus laevistriatus Perris, 1869
Psammodius rotundipennis (Reitter, 1892)⇒ Brindalus rotundipennis Reitter, 1892
Pseudacrossus sharpi Harold, 1874
Pseudesymus lucidus Klug, 1845
Pseudotrochalus quadrilineatus Fabricius, 1801
Pubinus tomentosus O.F.Muller, 1776
Rhizotrogus (Amphimallon) solstitialis (Linnaeus, 1758)⇒ Amphimallon volgense Fischer, 1823
Rhizotrogus aestivus A. G. Olivier, 1789
Rhizotrogus alicantinus Dieck, 1870
Rhizotrogus almeriensis Baraud, 1970
Rhizotrogus angelesae Galante, 1981
Rhizotrogus bolivari (Martinez y Saez, 1873)⇒ Amadotrogus patruelis Reiche, 1862
Rhizotrogus brunneus Fairmaire, 1870
Rhizotrogus camerosensis Baguena-Corella, 1955
Rhizotrogus chevrolati Graells, 1858
Rhizotrogus cicatricosus Mulsant, 1842
Rhizotrogus corduensis (Baguena, 1955)⇒ Rhizotrogus granatensis Baguena, 1955
Rhizotrogus flavicans Blanchard, 1850
Rhizotrogus granatensis Baguena, 1955
Rhizotrogus iglesiasi Baguena-Corella, 1955
Rhizotrogus lepidus (Ludovici & S..., 1869)⇒ Rhizotrogus pallidipennis Blanchard, 1850
Rhizotrogus maculicollis Villa, 1833
Rhizotrogus marginipes Mulsant, 1842
Rhizotrogus mascarauxi Desbrochers, 1895
Rhizotrogus monticola Blanchard, 1850
Rhizotrogus neglectus Perez Arcas, 1865
Rhizotrogus nevadensis Reitter, 1902
Rhizotrogus pallidipennis Blanchard, 1850
Rhizotrogus parvulus Rosenhauer, 1858
Rhizotrogus ribbei Reitter, 1908
Rhizotrogus rosalesi Fairmaire, 1862
Rhizotrogus subemarginatus Reiche, 1862
Rhizotrogus subsinuatus Baguena, 1955
Rhizotrogus toletanus (Baguena, 1955)⇒ Rhizotrogus parvulus Rosenhauer, 1858
Rhodaphodius foetens Fabricius, 1787
Rhyssemodes orientalis Mulsant & Go..., 1874
Rhyssemus algiricus Lucas, 1846
Rhyssemus cantabricus (Balthasar, 1961)⇒ Rhyssemus germanus Linnaeus, 1767
Rhyssemus convexus Benard, 1911
Rhyssemus germanus Linnaeus, 1767Germany small dung beetle
Rhyssemus marqueti Reiche, 1863
Rhyssemus mayeti Clouët, 1901
Rhyssemus parallelus Reitter, 1892
Rhyssemus plicatus Germar, 1817
Rhyssemus psammobiiformis Petrovitz, 1963
Rhyssemus purkynei Balthasar, 1961
Rhyssemus sulcatus Olivier, 1789
Rhyssemus verrucosus Mulsant, 1842
Scabrostomus tanytarsus Skelley & Wo..., 1991Masked stargazer
Scarabaeolus bohemani Harold, 1868
Scarabaeus (Ateuchetus) cicatricosus (Lucas, 1846)⇒ Scarabaeus palemo Olivier, 1789
Scarabaeus (Ateuchetus) laticollis Linnaeus, 1767
Scarabaeus (Ateuchetus) puncticollis (Latreille, 1819)⇒ Geotrupes spiniger Marsham, 1802
Scarabaeus (Ateuchetus) semipunctatus Fabricius, 1792
Scarabaeus (Ateuchetus) variolosus (Fabricius, 1787)⇒ Ateuchetus semipunctatus Fabricius, 1792
Scarabaeus (Scarabaeus) pius Illiger, 1803
Scarabaeus (Scarabaeus) sacer (Linnaeus, 1758)⇒ Scarabaeus pius Illiger, 1803
Scarabaeus (Scarabaeus) typhon (Fischer von ..., 1823)⇒ Megasoma gyas Herbst, 1785
Scarabaeus indus (Linnaeus, 1758)⇒ Euphoria inda Linnaeus, 1758
Scarabaeus palemo Olivier, 1789
Scarabaeus rufus (DeGeer, 1778)⇒ Rhysothorax rufus Fabricius, 1792
Serica (Serica) brunnea (Linnaeus, 1758)⇒ Serica similis Lewis, 1895
Serica brunna Linnaeus, 1758
Serica similis Lewis, 1895
Sericesthis accola Britton, 1987
Sigorus porcus Fabricius, 1792Mahogany Ridgeback scarab
Sinodrepanus falsus Sharp, 1875
Sisyphus crispatus Gory, 1833
Sisyphus schaefferi Linnaeus, 1758
Sisyphus schaefferi schaefferi Linnaeus, 1758
Stenocrates hardyi Dechambre, 1985
Strategus aloeus Linnaeus, 1758Coconut cockle
Strategus jugurtha Burmeister, 1847
Strigoderma rutelina Bates, 1888
Subrinus sturmi Harold, 1870
Subrinus vitellinus Klug, 1845
Temnorhynchus (Temnorhynchus) baal Reiche & Saulcy, 1856
Temnorrhynchus baal Reiche & Saulcy, 1856
Teuchestes brachysomus Solsky, 1874
Teuchestes fossor Linnaeus, 1758Digger small dung beetle
Tomarus bituberculatus Palisot de B..., 1805
Tomarus gyas Erichson, 1848
Tomarus nasutus Burmeister, 1847Spotted kelpfish
Tomarus sallaei Bates, 1888
Trichius fasciatus Linnaeus, 1758Bee Beetle
Trichius gallicus (Dejean, 1821)⇒ Coilodera penicillata Hope, 1831
Trichius gallicus gallicus Dejean, 1821
Trichius rosaceus Voet, 1769
Trichius sexualis Bedel, 1906
Trichius zonatus (Germar, 1829)⇒ Trichius rosaceus Voet, 1769
Trichonotulus scrofa Fabricius, 1787Sow small dung beetle
Trigonopeltastes geometricus Schaum, 1841
Trigonopeltastes mombachoensis Smith, 2016
Triodontella aquila Laporte, 1840
Triodontella castiliana Baraud, 1961
Triodontella castillana Baraud, 1961
Triodontella lajonquierei Baraud, 1961
Triodontella ruizi Pérez-López ..., 1994
Triodontella zuzartei Branco, 1978
Triodontus hova Fairmaire, 1868
Trochalus integriceps Moser, 1917
Tropinota (Epicometis) hirta Poda, 1761Blossom feeder
Tropinota hirta hirta Poda, 1761
Tropinota (Tropinota) squalida Scopoli, 1783
Tropinota squalida canariensis Lindberg, 1950
Tropinota squalida squalida Scopoli, 1783
Tropinota iberica Rössner, Blo..., 2018
Uroxys micros Bates, 1887
Valgus hemipterus Linnaeus, 1758Bug flower chafer
Valgus hemipterus hemipterus
Volinus sticticus Panzer, 1798
Xeropsamobeus asellus Schmidt, 1907Spotted weever
Xylonychus eucalypti pentaphyllus Arrow, 1924
Xylotrupes gideon Linnaeus, 1767Brown rhinoceros beetle
Xylotrupes gideon lamachus (Minck, 1920)⇒ Xylotrupes macleayi lamachus Minck, 1920

Subnodes of Scarabaeidae with occurrences in Spain
SubfamilyAclopinae
Blanchard, 1850
2
2
Genera
Species

Taxom

SubfamilyAphodiinae
Leach, 1815
Aphodiine Dung Beetles, Small dung beetles
76
273
Genera
Species

Taxom

SubfamilyCetoniinae
Leach, 1815
Flower beetles, Flower chafers, Flower scarabs, Fruit and flower chafers
21
50
Genera
Species

Taxom

SubfamilyChironinae
Blanchard, 1845
1
1
Genus
Species

Taxom

SubfamilyDynastinae
MacLeay, 1819
Horn beetles, Rhinoceros beetles
26
62
Genera
Species

Taxom

SubfamilyMelolonthinae
Samouelle, 1819
May Beetles and Junebugs, White grubs
40
221
Genera
Species

Taxom

SubfamilyOrphninae
Erichson, 1847
3
8
Genera
Species

Taxom

SubfamilyRutelinae
MacLeay, 1819
Shining Flower Chafers, Shining Leaf Chafers
14
48
Genera
Species

Taxom

SubfamilyScarabaeinae
Latreille, 1802
Dung Beetles, True dung beetles
27
124
Genera
Species

Taxom



Scarabaeidae (Latreille 1802)
Scarabaeidae (Latreille 1802)


Links, sources and further information
Scarabaeidae at artfakta.se
Scarabaeidae at dyntaxa.se
Scarabaeidae at gbif.org
Scarabaeidae at irmng.org
Scarabaeidae at itis.gov