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TAXFAMGENSPESUB
Checklist for Family Scarabaeidae Latreille, 1802 (Dung beetles)
and region Cambodia (Indochinese Peninsula)


Class
 
Insecta
Linnaeus, 1758
Insects, True insects
231
1540
2878
Families
Genera
Species
 
Taxom
 

Subclass
 
Pterygota
Lang, 1888
Winged insects
231
1540
2878
Families
Genera
Species
 
Taxom
 

Infraclass
 
Neoptera
van der Wulp, 1890
Modern Wing-folding insects
216
1447
2709
Families
Genera
Species
 
Taxom
 

Superorder
 
Holometabola
Heider, 1889
Holometabolous Insects
146
1189
2362
Families
Genera
Species
 
Taxom
 

Order
 
Coleoptera
Linnaeus, 1758
Beetles, Coleopterans
43
569
1195
Families
Genera
Species
 
Taxom
 

Suborder
 
Polyphaga
Emery, 1886
Polyphags
42
475
933
Families
Genera
Species
 
Taxom
 

Infraorder
 
Scarabaeiformia
Crowson, 1960
10
137
337
Families
Genera
Species
 
Taxom
 

Superfamily
 
Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles
10
137
337
Families
Genera
Species
 
Taxom
 
FamilyScarabaeidae
Latreille, 1802
Dung beetles, Flower beetles, Fruit chafers, Rain beetles
101
266
Genera
Species
 
Taxom
 


Genera for Cambodia
101 Genera with 266 Species
#ABCDEGHILMNOPRSTX
Adoretosoma Blanchard, 19552SpeciesTaxom
Adoretus Laporte, 191422SpeciesTaxom
Agnorimus Miyake & Iwase, 19011SpeciesTaxom
Alissonotum Arrow, 20133SpeciesTaxom
Anoctus Sharp, 19421SpeciesTaxom
Anomala Samouelle, 1775Shining Leaf Chafer22SpeciesTaxom
Aphodius Hellwig, 18471SpeciesTaxom
Apogonia Melichar, 20104SpeciesTaxom
Autoserica Brenske, 1972Garden beetle1SpeciesTaxom
Bivalgus Paulian, 19611SpeciesTaxom
Bombodes Westwood, 18481SpeciesTaxom
Caccobius Thomson, 17981SpeciesTaxom
Campsiura Hope, 18911SpeciesTaxom
Cassolus Sharp, 18751SpeciesTaxom
Catharsius Hope, 17583SpeciesTaxom
Centrognathus Guérin-Ménev..., 19131SpeciesTaxom
Chaetadoretus Ohaus, 19142SpeciesTaxom
Chalcosoma Hope, 18344SpeciesTaxom
Cheirotonus Hope, 18483SpeciesTaxom
Clinteria Burmeister, 02SpeciesTaxom
Coenochilus Burmeister &..., 20113SpeciesTaxom
Coilodera Hope, 18311SpeciesTaxom
Copris Geoffroy, 1921Mexican dung beetle13SpeciesTaxom
Cyphochilus Waterhouse, 18671SpeciesTaxom
Dasylepida Moser, 01SpeciesTaxom
Dasyvalgus Kolbe, 19611SpeciesTaxom
Dedalopterus2SpeciesTaxom
Dicaulocephalus Gestro, 19011SpeciesTaxom
Dicronocephalus Hope, 18311SpeciesTaxom
Didrepanephorus Wood-Mason, 20041SpeciesTaxom
Digitonthophagus Balthasar, 17751SpeciesTaxom
Emadiellus Schmidt, 03SpeciesTaxom
Eocorythoderus Maruyama, 20121SpeciesTaxom
Eophileurus Arrow, 19142SpeciesTaxom
Eotrichia Medvedev, 18921SpeciesTaxom
Epitrichius Tagawa, 18571SpeciesTaxom
Eupatorus Burmeister, 01SpeciesTaxom
Euselates J. Thomson, 04SpeciesTaxom
Exolontha Reitter, 02SpeciesTaxom
Gametis Burmeister, 18352SpeciesTaxom
Gilletianus Balthasar, 19632SpeciesTaxom
Glycyphana Burmeister, 02SpeciesTaxom
Goliathopsis Janson, 19912SpeciesTaxom
Heliocopris Hope, 18682SpeciesTaxom
Holotrichia Hope, 18924SpeciesTaxom
Hoplolontha Fairmaire, 19041SpeciesTaxom
Ischiopsopha Gestro, 18671SpeciesTaxom
Ixorida Thomson, 18682SpeciesTaxom
Leiopsammodius Rakovic, 19001SpeciesTaxom
Lepidiota Kirby, 19082SpeciesTaxom
Liatongus Reitter, 17985SpeciesTaxom
Macronota Hoffmansegg, 18861SpeciesTaxom
Maladera Mulsant, 18988SpeciesTaxom
Mecinonota Kraatz, 18571SpeciesTaxom
Megatelus Reitter, 18791SpeciesTaxom
Megistophylla Burmeister, 18911SpeciesTaxom
Melolontha Fabricius, 1896Billy witch1SpeciesTaxom
Microcopris Balthasar, 19212SpeciesTaxom
Microryctes Arrow, 18932SpeciesTaxom
Microserica Brenske, 18931SpeciesTaxom
Mimela Kirby, 19026SpeciesTaxom
Neocalaphodius Bordat, 18011SpeciesTaxom
Neoserica Brenske, 18915SpeciesTaxom
Oniticellus Serville, 18792SpeciesTaxom
Onitis Fabricius, 19313SpeciesTaxom
Onthophagus Latreille, 191422SpeciesTaxom
Oreoderus Burmeister, 20002SpeciesTaxom
Orphnus MacLeay, 18781SpeciesTaxom
Oryctes Hellwig, 1758Coconut rhinoceros beetles1SpeciesTaxom
Pachnessa Brenske, 20092SpeciesTaxom
Parachorius Harold, 18732SpeciesTaxom
Paragymnopleurus Shipp, 18753SpeciesTaxom
Parascatonomus Paulian, 18793SpeciesTaxom
Paratrichius Janson, 19156SpeciesTaxom
Peltonotus Burmeister, 19101SpeciesTaxom
Pentodon Hope, 17781SpeciesTaxom
Periphanesthes Kraatz, 18571SpeciesTaxom
Pharaphodius Reitter, 18581SpeciesTaxom
Philistina MacLeay, 19032SpeciesTaxom
Phyllopertha Stephens, 19051SpeciesTaxom
Platynocephalus Westwood, 02SpeciesTaxom
Polyphylla Harris, 2006Lined June Beetles1SpeciesTaxom
Popillia Leach, 19965SpeciesTaxom
Proagoderus Lansberge, 18751SpeciesTaxom
Protaetia Burmeister, 20206SpeciesTaxom
Pseudosinghala Heller, 18551SpeciesTaxom
Pseudotorynorrhina Miksic, 19652SpeciesTaxom
Rhinyptia Burmeister, 19141SpeciesTaxom
Rhomborhina Hope, 02SpeciesTaxom
Sisyphus Latreille, 18332SpeciesTaxom
Spinanomala Ohaus, 19321SpeciesTaxom
Synapsis Forster, 18811SpeciesTaxom
Taeniodera Burmeister, 19245SpeciesTaxom
Termitotrox Reichensperger, 20152SpeciesTaxom
Tetraserica12SpeciesTaxom
Thaumastopeus Kraatz, 18861SpeciesTaxom
Tibiodrepanus Krikken, 18681SpeciesTaxom
Tocama Reitter, 19132SpeciesTaxom
Trichanomala Arrow, 19081SpeciesTaxom
Trichogomphus Burmeister, 19081SpeciesTaxom
Xylotrupes Hope, 1920Elephant beetles5SpeciesTaxom


Species list for Cambodia
266 Species and 23 Subspecies
Adoretosoma fairmairei Arrow, 1899
Adoretosoma tonkinense Machatschke, 1955
Adoretus (Adoretus) epipleuralis Arrow, 1914
Adoretus (Adoretus) lajoyi Ohaus, 1914
Adoretus (Adoretus) laosensis Frey, 1970
Adoretus (Adoretus) nitens Frey, 1970
Adoretus (Adoretus) runcinatus Lin, 1974
Adoretus (Adoretus) saigonensis Ohaus, 1914
Adoretus (Adoretus) vietnamensis Frey, 1970
Adoretus arrowi Frey, 1970
Adoretus bacrii Limbourg, Co..., 2022
Adoretus cambodiensis Limbourg, Co..., 2022
Adoretus cirroseriatus Ohaus, 1914
Adoretus compressus Weber, 1801Brown cockchafer
Adoretus cribratus (White, 1844)⇒ Adoretus cribrosus Harold, 1869
Adoretus husemanni Limbourg, Co..., 2022
Adoretus khmerus Limbourg, Co..., 2022
Adoretus ohmomoi Kobayashi, 2007
Adoretus pauliani Frey, 1970
Adoretus phauki Limbourg, Co..., 2022
Adoretus sinicus Burmeister, 1855Chinese rose beetle
Adoretus sougnezi Limbourg, Co..., 2022
Adoretus villicollis Ohaus, 1914
Adoretus vitticauda Arrow, 1914
Agnorimus pictus Moser, 1901
Alissonotum cribratellum Fairmaire, 1893
Alissonotum suzukii Yamaya, 2016
Alissonotum yamayai Drumont, 2013
Anoctus siamensis Balthasar, 1942
Anomala amychodes Ohaus, 1914
Anomala anguliceps Arrow, 1917
Anomala antiqua Gyllenhal, 1817
Anomala aspera Ohaus, 1914
Anomala atrovirens Lin, 2000
Anomala aulax Wiedemann, 1823
Anomala bilunata Fairmaire, 1888
Anomala brevidens Ohaus, 1914
Anomala castelnaui Ohaus, 1910
Anomala controversa Hope, 1845
Anomala cupripes Hope, 1839Large green chafer beetle
Anomala curator Benderitter, 1929
Anomala densa Arrow, 1917
Anomala grandis Hope, 1839
Anomala limbourgi Zorn, 2011
Anomala obsoleta Blanchard, 1851
Anomala pallida Fabricius, 1775Cockchafer
Anomala russiventris Fairmaire, 1893
Anomala smetsi Limbourg & Zorn, 2010
Anomala transversa Lin, 1999
Anomala varicolor Gyllenhal, 1817
Anomala viridis (Fabricius, 1775)⇒ Anomala yunnana Fairmaire, 1886
Aphodius (Aphodius) elegans Allibert, 1847
Apogonia cambodjensis Moser, 1914
Apogonia cinerascens Fairmaire, 1893
Apogonia cribricollis Burmeister, 1855Chafer beetle
Apogonia illiberalis Kobayashi, 2010
Autoserica muscula Frey, 1972
Bivalgus harmandi Paulian, 1961
Bombodes ursus Westwood, 1848
Caccobius (Caccophilus) unicornis Fabricius, 1798
Campsiura insignis Gestro, 1891
Cassolus nudus Sharp, 1875
Catharsius birmanensis Lansberge, 1874
Catharsius javanus Lansberge, 1886
Catharsius molossus Linnaeus, 1758
Centrognathus bimaculatus Moser, 1913
Chaetadoretus cirroseriatus Ohaus, 1914
Chaetadoretus villicollis Ohaus, 1914
Chalcosoma atlas Linnaeus, 1758Atlas beetle
Chalcosoma atlas mantetsu Nagai, 2004
Chalcosoma chiron (Olivier, 1789)⇒ Chalcosoma caucasus Fabricius, 1801
Chalcosoma chiron belangeri Guérin-Ménev..., 1834
Cheirotonus gestroi Pouillaude, 1913
Cheirotonus gestroi gestroi
Cheirotonus parryi Gray, 1848
Clinteria ducalis White, 1856
Clinteria ducalis ducalis
Coenochilus apicalis Westwood, 1874
Coenochilus armiger Westwood, 1874
Coenochilus thailandicus Maruyama, Ue..., 2011
Coilodera penicillata Hope, 1831
Copris (Copris) cambodiensis Ochi, Kon & ..., 2008
Copris cambodiensis cambodiensis
Copris confucius Harold, 1877
Copris corpulentus Gillet, 1910
Copris kiriromensis Ochi & Kon, 2021
Copris marcus Gillet, 1921
Copris minutus Drury, 1773Small Black Dung Beetle
Copris nevinsoni Waterhouse, 1891Warton meshweaver
Copris punctulatus Wiedemann, 1823
Copris punctulatus punctulatus
Copris reflexus (Fabricius, 1787)⇒ Copris minutus Drury, 1773
Copris sinicus Hope, 1842
Copris vitalisi Gillet, 1921
Cyphochilus tricolor Waterhouse, 1867
Dasylepida rudepunctata rudepunctata
Dasyvalgus cambodjensis Paulian, 1961
Dedalopterus bezdekorum Zidek & Krajcik, 2007
Dedalopterus itohi Sabatinelli ..., 1998
Dicaulocephalus fruhstorferi Felsche, 1901
Dicronocephalus wallichi Hope, 1831
Didrepanephorus ohbayashii Nagai, 2004
Digitonthophagus bonasus Fabricius, 1775
Emadiellus rufopustulatus Wiedemann, 1823
Emadiellus rufopustulatus pistor Balthasar, 1945
Emadiellus rufopustulatus rufopustulatus
Eocorythoderus incredibilis Maruyama, 2012
Eophileurus confinis Prell, 1913
Eophileurus quadrigeminatus Arrow, 1914
Eotrichia scrobiculata Brenske, 1892
Epitrichius bowringi Thomson, 1857
Eupatorus siamensis siamensis
Euselates magna Thomson, 1880
Euselates ornata Saunders, 1852
Euselates perraudieri taivanica Mi..., 1974
Euselates perraudierii perraudierii
Exolontha chiangmaiensis Keith & Saba..., 2012
Exolontha chiangmaiensis chiangmaiensis
Gametis bealiae Gory & Perch..., 1833
Gametis jucunda Faldermann, 1835Scarab
Gilletianus commatoides Balthasar, 1961
Gilletianus therondi Balthasar, 1963
Glycyphana horsfieldi Hope, 1831
Glycyphana horsfieldi horsfieldi
Goliathopsis despectus Westwood, 1874
Goliathopsis duponti Antoine, 1991
Heliocopris bucephalus Fabricius, 1775
Heliocopris dominus Bates, 1868
Holotrichia cochinchina Nonfried, 1891
Holotrichia costulata Frey, 1970
Holotrichia pilipyga Zhang, 1965
Holotrichia scrobiculata Brenske, 1892
Hoplolontha pavi Fairmaire, 1904
Ischiopsopha cambodiensis Wallace, 1867
Ixorida venerea siamensis Antoine, 1986
Ixorida mouhoti Wallace, 1868
Leiopsammodius gestroi Clouët, 1900
Lepidiota acuminata Moser, 1913
Lepidiota rudepunctata Moser, 1908
Liatongus affinis Arrow, 1908
Liatongus gagatinus Hope, 1831
Liatongus phanaeoides Westwood, 1839
Liatongus tridentatus Boucomont, 1919
Liatongus vertagus Fabricius, 1798
Macronota shangaicus Neervoort Va..., 1886
Maladera atavana Brenske, 1899
Maladera cambodjana Frey, 1972
Maladera eclogaria Brenske, 1899
Maladera eluctabilis Brenske, 1899
Maladera futschauana Brenske, 1897
Maladera muscula Frey, 1972
Maladera rufoplagiata Fairmaire, 1893
Maladera tibialis Brenske, 1898
Mecinonota venerea Thomson, 1857
Megatelus brahminus Harold, 1879
Megistophylla grandicornis Fairmaire, 1891
Melolontha carinata Brenske, 1896
Microcopris reflexus Fabricius, 1787
Microcopris vitalisi Gillet, 1921
Microryctes hainanensis Zhang, 2002
Microryctes monodon Fairmaire, 1893
Microserica malaccensis Brenske, 1893
Mimela chinensis Kirby, 1825
Mimela dehaani Hope, 1839
Mimela nigritarsis Lin, 1990
Mimela nigrosellata Ohaus, 1913
Mimela ohausi Arrow, 1908
Mimela signaticollis (Ohaus, 1902)⇒ Mimela junii Duftschmid, 1805
Neocalaphodius moestus Fabricius, 1801
Neoserica cardamomensis Ahrens, Liu,..., 2014
Neoserica inclinata Brenske, 1898
Neoserica insubida Brenske, 1898
Neoserica pavicana Brenske, 1899
Neoserica picea Nonfried, 1891
Oniticellus cinctus Fabricius, 1775Bordered dung beetle
Oniticellus tessellatus Harold, 1879
Onitis corydon Boisduval, 1835
Onitis falcatus Wulfen, 1786
Onitis subopacus Arrow, 1931
Onthophagus (Colobonthophagus) tragus Fabricius, 1792
Onthophagus (Furconthophagus) papulatus Boucomont, 1914
Onthophagus (Gibbonthophagus) duporti Boucomont, 1914
Onthophagus (Gibbonthophagus) limbatus Herbst, 1789
Onthophagus (Gibbonthophagus) luridipennis Boheman, 1858
Onthophagus (Gibbonthophagus) taurinus White, 1844
Onthophagus (Indachorius) phuquoci Paulian, 1945
Onthophagus (Micronthophagus) vigilans Boucomont, 1921
Onthophagus (Onthophagus) orientalis (Harold, 1868)⇒ Onthophagus fissinasus Fairmaire, 1895
Onthophagus (Onthophagus) proletarius Harold, 1875
Onthophagus (Onthophagus) trituber Wiedemann, 1823
Onthophagus (Serrophorus) rectecornutus Lansberge, 1883
Onthophagus (Serrophorus) sagittarius Fabricius, 1775Sagittarius dung beetle
Onthophagus armatus Blanchard, 1853
Onthophagus baolocensis Ochi & Kon, 2015
Onthophagus crassicollis Boucomont, 1913
Onthophagus lunatus Harold, 1868
Onthophagus pacificus Lansberge, 1885
Onthophagus senex Boucomont, 1914
Onthophagus seniculus (Fabricius, 1781)⇒ Onthophagus senescens Péringuey, 1908
Onthophagus tonkineus Gillet, 1921
Onthophagus tragoides Boucomont, 1914
Oreoderus gracilicornis Paulian, 1961
Oreoderus siamensis Ricchiardi, 2000
Orphnus cribratellus Fairmaire, 1878
Oryctes rhinoceros Linnaeus, 1758Asiatic rhinoceros beetle
Pachnessa drumonti Keith, 2009
Pachnessa smetsi Keith, 2009
Parachorius nudus Sharp, 1875
Parachorius thomsoni Harold, 1873
Paragymnopleurus melanarius Harold, 1867
Paragymnopleurus sinuatus abax Sharp, 1875
Paragymnopleurus sinuatus productus Sharp, 1875
Parascatonomus alexeevi Tarasov, Kri..., 2010
Parascatonomus discedens Sharp, 1875
Parascatonomus penicillatus Harold, 1879
Paratrichius discolor Jordan, 1895
Paratrichius flavipes Moser, 1901
Paratrichius hajeki Krajcik, 2010
Paratrichius nicoudi Bourgoin, 1920
Paratrichius rotundatus Ma, 1990
Paratrichius vicinus Bourgoin, 1915
Peltonotus nasutus Arrow, 1910
Pentodon algerinus (Fuessly, 1778)⇒ Pentodon algerinum Herbst, 1789
Periphanesthes aurora Motschulsky, 1857
Pharaphodius priscus Motschulsky, 1858
Philistina tibetana Janson, 1917
Philistina tonkinensis Moser, 1903
Phyllopertha carinicollis Ohaus, 1905
Platynocephalus miyashitai Delpont, 1995
Platynocephalus miyashitai miyashitai
Polyphylla phongsali Zidek, 2006
Popillia bitacta Kraatz, 1892
Popillia flavosellata (Fairmaire, 1886)⇒ Popillia bitacta Kraatz, 1892
Popillia histeroidea Gyllenhal, 1817
Popillia subquadrata Kraatz, 1892
Popillia uhligi Sabatinelli, 1996
Proagoderus mouhoti Harold, 1875
Protaetia (Acanthoprotaetia) niveoguttata Janson, 1876
Protaetia (Heteroprotaetia) fusca Herbst, 1790Asian Mango Flower Beetle
Protaetia (Pachyprotaetia) sakaiana Antoine, 2001
Protaetia (Pseudocetonischema) ceylanica Schoch, 1894
Protaetia (Pseudocetonischema) pretiosa (Nonfried, 1891)⇒ Protaetia ceylanica Schoch, 1894
Protaetia cambodia Jákl, 2020
Pseudosinghala transversa Burmeister, 1855
Pseudotorynorrhina japonica tonkiniana Ruter, 1965
Pseudotorynorrhina tonkiniana Ruter, 1965
Rhinyptia parabolica Ohaus, 1914
Rhomborhina mellyi Gory & Perch..., 1833
Rhomborhina mellyi mellyi
Sisyphus denticrus (Fairmaire, 1886)⇒ Sisyphus neglectus Gory, 1833
Sisyphus neglectus Gory, 1833
Spinanomala benderitteri Burgeon, 1932
Synapsis tridens Sharp, 1881
Taeniodera garnieri Bourgoin, 1917
Taeniodera idolica Janson, 1909
Taeniodera miksiciana Krajcik, 2010
Taeniodera murzini Jakl, 2012
Taeniodera salvazai Bourgoin, 1924
Termitotrox cupido Maruyama, 2012
Termitotrox venus Kakizoe & Ma..., 2015
Tetraserica angkorthomensis Fabrizi, Dal..., 2019
Tetraserica angkorwatensis Fabrizi, Dal..., 2019
Tetraserica auriculata Fabrizi, Dal..., 2019
Tetraserica constanti Fabrizi, Dal..., 2019
Tetraserica kiriromensis Fabrizi, Dal..., 2019
Tetraserica nussi Fabrizi, Dal..., 2019
Tetraserica pailinensis Fabrizi, Dal..., 2019
Tetraserica quadrifurcata Fabrizi, Dal..., 2019
Tetraserica semiruiliensis Fabrizi, Dal..., 2019
Tetraserica smetsi Fabrizi, Dal..., 2019
Tetraserica subrotundata Fabrizi, Dal..., 2019
Tetraserica vari Fabrizi, Dal..., 2019
Thaumastopeus shangaicus Neervoort Va..., 1886
Tibiodrepanus sinicus Harold, 1868
Tocama laevipennis Blanchard, 1851
Tocama tonkinensis Moser, 1913
Trichanomala tonkinensis Ohaus, 1908
Trichogomphus mongol Arrow, 1908
Xylotrupes socrates Schauffus, 1864Siamese rhinoceros beetle
Xylotrupes socrates bourgini Paulian, 1945
Xylotrupes socrates siamensis Minck, 1920
Xylotrupes socrates socrates
Xylotrupes socrates tonkinensis Minck, 1920

Subnodes of Scarabaeidae with occurrences in Cambodia
SubfamilyAphodiinae
Leach, 1815
Aphodiine Dung Beetles, Small dung beetles
9
11
Genera
Species

Taxom

SubfamilyCetoniinae
Leach, 1815
Flower beetles, Flower chafers, Flower scarabs, Fruit and flower chafers
29
54
Genera
Species

Taxom

SubfamilyDynastinae
MacLeay, 1819
Horn beetles, Rhinoceros beetles
9
14
Genera
Species

Taxom

SubfamilyEuchirinae
Hope, 1840
1
2
Genus
Species

Taxom

SubfamilyMelolonthinae
Samouelle, 1819
May Beetles and Junebugs, White grubs
18
50
Genera
Species

Taxom

SubfamilyOrphninae
Erichson, 1847
1
1
Genus
Species

Taxom

SubfamilyRutelinae
MacLeay, 1819
Shining Flower Chafers, Shining Leaf Chafers
13
65
Genera
Species

Taxom

SubfamilyScarabaeinae
Latreille, 1802
Dung Beetles, True dung beetles
17
61
Genera
Species

Taxom



Scarabaeidae (Latreille 1802)
Scarabaeidae (Latreille 1802)


Links, sources and further information
Scarabaeidae at artfakta.se
Scarabaeidae at dyntaxa.se
Scarabaeidae at gbif.org
Scarabaeidae at irmng.org
Scarabaeidae at itis.gov