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Checklist for Family Scarabaeidae Latreille, 1802 (Dung beetles)
and region Bhutan (South Asia)


Class
 
Insecta
Linnaeus, 1758
Insects, True insects
205
1532
2897
Families
Genera
Species
 
Taxom
 

Subclass
 
Pterygota
Lang, 1888
Winged insects
205
1532
2897
Families
Genera
Species
 
Taxom
 

Infraclass
 
Neoptera
van der Wulp, 1890
Modern Wing-folding insects
198
1516
2881
Families
Genera
Species
 
Taxom
 

Superorder
 
Holometabola
Heider, 1889
Holometabolous Insects
138
1363
2673
Families
Genera
Species
 
Taxom
 

Order
 
Coleoptera
Linnaeus, 1758
Beetles, Coleopterans
50
546
1154
Families
Genera
Species
 
Taxom
 

Suborder
 
Polyphaga
Emery, 1886
Polyphags
48
457
941
Families
Genera
Species
 
Taxom
 

Infraorder
 
Scarabaeiformia
Crowson, 1960
6
140
377
Families
Genera
Species
 
Taxom
 

Superfamily
 
Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles
6
140
377
Families
Genera
Species
 
Taxom
 
FamilyScarabaeidae
Latreille, 1802
Dung beetles, Flower beetles, Fruit chafers, Rain beetles
102
257
Genera
Species
 
Taxom
 


Genera for Bhutan
102 Genera with 257 Species
#ABCDEGHIJKLMNOPRSTX
Acrossus Mulsant, 19761SpeciesTaxom
Adoretosoma Blanchard, 18512SpeciesTaxom
Adoretus Laporte, 18313SpeciesTaxom
Aganocrossus Reitter, 18621SpeciesTaxom
Agoliinus Schmidt, 20111SpeciesTaxom
Amiserica1SpeciesTaxom
Anomala Samouelle, 1831Shining Leaf Chafer24SpeciesTaxom
Aphodius Hellwig, 18811SpeciesTaxom
Apogonia Melichar, 19753SpeciesTaxom
Asactopholis Brenske, 19722SpeciesTaxom
Bombodes Westwood, 18572SpeciesTaxom
Brachiaphodius Koshantschikov, 18741SpeciesTaxom
Brachyllus Brenske, 20051SpeciesTaxom
Brahmina Blanchard, 18441SpeciesTaxom
Callistethus Blanchard, 05SpeciesTaxom
Campsiura Hope, 18881SpeciesTaxom
Carinaulus Tesar, 19651SpeciesTaxom
Catharsius Hope, 18062SpeciesTaxom
Chaetadoretus Ohaus, 18312SpeciesTaxom
Cheirotonus Hope, 18403SpeciesTaxom
Clerota Burmeister, 18412SpeciesTaxom
Clinteria Burmeister, 19422SpeciesTaxom
Clypeolontha2SpeciesTaxom
Coelodera Géné, 1836
0SpeciesTaxom
Coenochilus Burmeister &..., 19101SpeciesTaxom
Dactylopopillia Arrow, 18311SpeciesTaxom
Dasyvalgus Kolbe, 18502SpeciesTaxom
Dicheros Gory & Perch..., 19092SpeciesTaxom
Dicronocephalus Hope, 18311SpeciesTaxom
Ectinohoplia Redtenbacher, 19971SpeciesTaxom
Eophileurus Arrow, 18351SpeciesTaxom
Eotrichia Medvedev, 18924SpeciesTaxom
Escarabaeus Zidek & Pokorny, 18401SpeciesTaxom
Eupatorus Burmeister, 18312SpeciesTaxom
Euselates J. Thomson, 18313SpeciesTaxom
Exolontha Reitter, 19751SpeciesTaxom
Gametis Burmeister, 17752SpeciesTaxom
Gilletianus Balthasar, 19702SpeciesTaxom
Glycosia Schoch, 02SpeciesTaxom
Glycyphana Burmeister, 19101SpeciesTaxom
Gynaecoserica Brenske, 20118SpeciesTaxom
Hemiheterorrhina Miksic, 18311SpeciesTaxom
Heterorhina Westwood, 18422SpeciesTaxom
Heterorrhina Westwood, 18411SpeciesTaxom
Hilyotrogus Fairmaire, 19753SpeciesTaxom
Holotrichia Hope, 19752SpeciesTaxom
Hoplia Illiger, 1983Monkey Beetles12SpeciesTaxom
Ischnopopillia Kraatz, 18923SpeciesTaxom
Jumnos Saunders, 02SpeciesTaxom
Koshantschikovius Schmidt, 19711SpeciesTaxom
Labarrus Mulsant and Rey, 19461SpeciesTaxom
Lasioserica Brenske, 20168SpeciesTaxom
Leuroserica Arrow, 19751SpeciesTaxom
Liatongus Reitter, 17984SpeciesTaxom
Macronotops Krikken, 18942SpeciesTaxom
Maladera Mulsant, 200414SpeciesTaxom
Melinopterus Mulsant, 19751SpeciesTaxom
Melolontha Fabricius, 1896Billy witch1SpeciesTaxom
Microserica Brenske, 19753SpeciesTaxom
Mimela Kirby, 185511SpeciesTaxom
Neocalaphodius Bordat, 18011SpeciesTaxom
Neoserica Brenske, 20041SpeciesTaxom
Neovalgus Miyake, 19281SpeciesTaxom
Nepaloserica Frey, 19962SpeciesTaxom
Onitis Fabricius, 17861SpeciesTaxom
Onthophagus Latreille, 190710SpeciesTaxom
Oreoderus Burmeister, 19101SpeciesTaxom
Oxyserica Brenske, 19754SpeciesTaxom
Pachyserica Brenske, 20041SpeciesTaxom
Panotrogus Reitter, 19131SpeciesTaxom
Paraphytus Harold, 19131SpeciesTaxom
Parastasia Westwood, 18413SpeciesTaxom
Paulianellus Balthasar, 19811SpeciesTaxom
Peltonotus Burmeister, 19102SpeciesTaxom
Peperonota Westwood, 02SpeciesTaxom
Pharaphodius Reitter, 17812SpeciesTaxom
Philistina MacLeay, 18811SpeciesTaxom
Polyphylla Harris, 0Lined June Beetles2SpeciesTaxom
Popillia Leach, 199413SpeciesTaxom
Propomacrus Newman, 20071SpeciesTaxom
Protaetia Burmeister, 01SpeciesTaxom
Pseudopanotrogus2SpeciesTaxom
Rhomborrhina Burmeister, 1842
0SpeciesTaxom
Rhyparus Westwood, 19051SpeciesTaxom
Rhyssemus Mulsant, 19841SpeciesTaxom
Saprosites Redtenbacher, 18751SpeciesTaxom
Serica MacLeay, 201812SpeciesTaxom
Silluvia Landin, 19771SpeciesTaxom
Sisyphus Latreille, 18331SpeciesTaxom
Sophrops Fairmaire, 19734SpeciesTaxom
Spinohoplia1SpeciesTaxom
Sybacodes Fairmaire, 02SpeciesTaxom
Synapsis Forster, 19312SpeciesTaxom
Taeniodera Burmeister, 19141SpeciesTaxom
Tetraserica1SpeciesTaxom
Teuchestes Mulsant, 17871SpeciesTaxom
Tibiodrepanus Krikken, 18231SpeciesTaxom
Torynorrhina Arrow, 18314SpeciesTaxom
Trigonophorus Nordmann, 18312SpeciesTaxom
Trioserica Moser, 18941SpeciesTaxom
Trypoxylus Arrow, 17711SpeciesTaxom
Xylotrupes Hope, 0Elephant beetles2SpeciesTaxom


Species list for Bhutan
257 Species and 26 Subspecies
Acrossus emodus Petrovitz, 1976
Adoretosoma bruschii Sabatinelli, 1997
Adoretosoma fulviventre Blanchard, 1851
Adoretus bhutanensis Frey, 1975
Adoretus lasiopygus Burmeister, 1855Grapevine chafer
Adoretus testaceus (Hope, 1831)⇒ Chaetadoretus capicola Burmeister, 1844
Aganocrossus urostigma Harold, 1862
Agoliinus durrelli Rakovic & Mencl, 2011
Amiserica argentata Frey, 1975
Anomala agilis Arrow, 1917
Anomala angusta Arrow, 1912
Anomala anthracina Arrow, 1912
Anomala bengalensis Blanchard, 1851
Anomala bilunata Fairmaire, 1888
Anomala dimidiata dimidiata
Anomala dorsalis Fabricius, 1775
Anomala flaviventris Arrow, 1912
Anomala flavofasciata Arrow, 1912
Anomala flavonotata Arrow, 1912
Anomala flavopicta Arrow, 1912
Anomala lineatopennis Blanchard, 1851
Anomala marginipennis Arrow, 1912
Anomala perplexa Hope, 1839
Anomala platypyga Fairmaire, 1893
Anomala propinqua Arrow, 1912
Anomala rufiventris Kollar & Red..., 1848
Anomala rugosa Arrow, 1899
Anomala spiloptera Burmeister, 1855
Anomala stenodera Arrow, 1917
Anomala trivirgata Fairmaire, 1888
Anomala trochanterica Arrow, 1917
Anomala varicolor Gyllenhal, 1817
Anomala variegata Hope, 1831
Aphodius fasciger Harold, 1881
Apogonia laevicollis Lansberge, 1879
Apogonia splendida (Boheman, 1858)⇒ Apogonia laevicollis Lansberge, 1879
Apogonia wittmeri Frey, 1975
Asactopholis bicolor bicolor
Asactopholis microsquamosa Frey, 1972
Bombodes ursus dejeani Pouillaude, 1914
Bombodes westwoodi Thomson, 1857
Brachiaphodius pilosus Harold, 1874
Brachyllus bhutanensis Keith, 2005
Brahmina cribricollis Redtenbacher, 1844
Callistethus auronitens Hope, 1835
Callistethus pterygophorus Ohaus, 1903
Callistethus pyroscelis Hope, 1841
Callistethus stoliczkae Sharp, 1878
Callistethus tumidicauda tumidicauda
Campsiura oberthuri Janson, 1888
Carinaulus sikkimensis Balthasar, 1965
Catharsius birmanensis Lansberge, 1874
Catharsius sagax Quenstedt, 1806
Chaetadoretus lasiopygus Burmeister, 1855
Chaetadoretus testaceus Hope, 1831
Cheirotonus gestroi Pouillaude, 1913
Cheirotonus gestroi gestroi
Cheirotonus macleayi Hope, 1840
Clerota shiva Valck Lucassen, 1931
Clerota vittigera (Hope, 1841)⇒ Clerota shiva Valck Lucassen, 1931
Clinteria confinis confinis
Clinteria confinis pseudoconfinis Schürhoff, 1942
Clypeolontha alboplagiata Brenske, 1898
Clypeolontha alboplagiata alboplagiata
Coenochilus solidus Arrow, 1910
Dactylopopillia virescens Hope, 1831
Dasyvalgus fulvicauda Arrow, 1910
Dasyvalgus penicillatus Blanchard, 1850
Dicheros bimacula Wiedemann, 1823
Dicheros bimacula gracilis Janson, 1909
Dicronocephalus wallichi Hope, 1831
Ectinohoplia ahrensis Sabatinelli, 1997
Eophileurus chinensis chinensis Faldermann, 1835
Eotrichia braeti Brenske, 1896
Eotrichia glabricollis Brenske, 1896
Eotrichia longipennis Blanchard, 1851
Eotrichia sikkimensis Brenske, 1892
Escarabaeus brahminus Castelnau, 1840
Eupatorus hardwickei Hope, 1831
Eupatorus hardwickii Hope, 1831
Euselates antennata Wallace, 1868
Euselates jansoni Arrow, 1910
Euselates quadrilineata Hope, 1831
Exolontha bhutanensis Frey, 1975
Gametis jucunda Faldermann, 1835Scarab
Gametis versicolor Fabricius, 1775
Gilletianus costatulus Schmidt, 1908
Gilletianus rangoonensis Petrovitz, 1970
Glycosia luctifera dureli Pouillaude, 1914
Glycosia luctifera luctifera
Glycyphana aurocincta Arrow, 1910
Gynaecoserica cymosa Brenske, 1896
Gynaecoserica gogonaica Ahrens, 1999
Gynaecoserica latesquamosa Frey, 1975
Gynaecoserica marginipes Brenske, 1896
Gynaecoserica pellecta Brenske, 1896
Gynaecoserica stemmleri Frey, 1975
Gynaecoserica variipennis wuermlii Frey, 1975
Gynaecoserica victori Ahrens & Fab..., 2011
Hemiheterorrhina mutabilis Hope, 1831
Heterorhina amoena Hope, 1841
Heterorhina tibialis Westwood, 1842
Heterorrhina amoena Hope, 1841
Hilyotrogus kolbei Brenske, 1892
Hilyotrogus nitens Frey, 1975
Hilyotrogus wittmeri Frey, 1975
Holotrichia bhutanensis Frey, 1975
Holotrichia kulzeri Frey, 1975
Hoplia coeruleosignata Moser, 1916
Hoplia coluzzi Sabatinelli, 1983
Hoplia coluzzii Sabatinelli, 1983
Hoplia freudei bhutanica Sabatinelli, 1983
Hoplia hirsuta Moser, 1912
Hoplia laetitiae Sabatinelli, 1983
Hoplia mahayana Sabatinelli, 1983
Hoplia sabraechatilae Sabatinelli, 1983
Hoplia tesari Sabatinelli, 1983
Hoplia virginioi Sabatinelli, 1983
Hoplia viridissima Brenske, 1894
Hoplia viridula gibbosa Sabatinelli, 1983
Ischnopopillia brancuccii Sabatinelli, 1997
Ischnopopillia lateralis Hope, 1831
Ischnopopillia moorei Kraatz, 1892
Jumnos roylei Hope, 1839
Jumnos ruckeri ruckeri
Koshantschikovius peculator Balthasar, 1971
Labarrus hoabinhensis Balthasar, 1946
Lasioserica breviclypeata Ahrens, 1999
Lasioserica bumthangana Ahrens, 1999
Lasioserica hamifer Ahrens & Fab..., 2011
Lasioserica imminuta Ahrens & Fab..., 2011
Lasioserica maculata bhutanica Ahrens, 1996
Lasioserica pacholatkoi Ahrens, 2000
Lasioserica pilosella Brenske, 1894
Lasioserica verschraegheni Ahrens & Fab..., 2016
Leuroserica stemmleri Frey, 1975
Liatongus clypeocornis Scheuern, 1988
Liatongus gagatinus Hope, 1831
Liatongus mergacerus Hope, 1831
Liatongus vertagus Fabricius, 1798
Macronotops sexmaculata Kraatz, 1894
Macronotops sexmaculatus Kraatz, 1894
Maladera bhutanensis Frey, 1975
Maladera cardoni Brenske, 1896
Maladera freyi Ahrens & Fab..., 2016
Maladera gardneri Ahrens, 2004
Maladera himalayica thimphuensis Ahrens, 2004
Maladera opaca Frey, 1975
Maladera paraquinquidens Ahrens, 2004
Maladera phuntsholingensis Ahrens, 2004
Maladera prenai Ahrens, 2004
Maladera sericella Brenske, 1898
Maladera siniaevi Ahrens, 2004
Maladera sprecherae Ahrens, 2004
Maladera thimphuensis Ahrens, 2004
Maladera trichotibialis Ahrens, 2004
Melinopterus wittmeri Petrovitz, 1975
Melolontha virescens Brenske, 1896
Microserica bhutanensis Frey, 1975
Microserica elegans Frey, 1975
Microserica hispidula Frey, 1975
Mimela dehaani Hope, 1839
Mimela fulgidivittata Blanchard, 1851
Mimela heterochropus Blanchard, 1851
Mimela heterochropus heterochropus
Mimela laevigata Arrow, 1908
Mimela marginalis Arrow, 1908
Mimela passerinii passerinii
Mimela pectoralis Blanchard, 1851
Mimela princeps Hope, 1841
Mimela vernicata Fairmaire, 1896
Mimela vittaticollis Burmeister, 1855
Neocalaphodius moestus Fabricius, 1801
Neoserica changrae Ahrens, 2004
Neovalgus laterufus Pic, 1928
Nepaloserica rufobrunnea Ahrens, 1999
Nepaloserica thimphui Ahrens & Sab..., 1996
Onitis falcatus Wulfen, 1786
Onthophagus (Colobonthophagus) dama Fabricius, 1798
Onthophagus (Macronthophagus) rubricollis Hope, 1831
Onthophagus (Matashia) anguliceps (Boucomont, 1914)⇒ Onthophagus avocetta Arrow, 1933
Onthophagus (Onthophagus) taurus Schreber, 1759Bull Headed Dung Beetle
Onthophagus (Serrophorus) rectecornutus Lansberge, 1883
Onthophagus cupreiceps Arrow, 1907
Onthophagus furcicollis (Arrow, 1931)⇒ Onthophagus hecate Panzer, 1794
Onthophagus gagates Hope, 1831
Onthophagus piffli Petrovitz, 1961
Onthophagus rubripennis (Arrow, 1907)⇒ Onthophagus taurus Schreber, 1759
Oreoderus bhutanus Arrow, 1910
Oxyserica bhutanensis Frey, 1975
Oxyserica elegans Frey, 1975
Oxyserica hispidula Frey, 1975
Oxyserica varia Frey, 1975
Pachyserica himalayensis Ahrens, 2004
Panotrogus hirsutus Moser, 1913
Paraphytus hindu Arrow, 1913
Parastasia alternata Arrow, 1899
Parastasia indica Ohaus, 1898
Parastasia rufopicta Westwood, 1841
Paulianellus khandui Stebnicka, 1981
Peltonotus morio (Burmeister, 1847)⇒ Peltonotus nasutus Arrow, 1910
Peltonotus nasutus Arrow, 1910
Peperonota harringtoni Westwood, 1847
Peperonota harringtoni harringtoni
Pharaphodius jodhpurensis Petrovitz, 1958
Pharaphodius marginellus Fabricius, 1781
Philistina microphylla Wood-Mason, 1881
Polyphylla lerestifi Guerlach, 2012
Polyphylla sikkimensis sikkimensis
Popillia bhutanensis Frey, 1975
Popillia bhutanica Sabatinelli, 1993
Popillia brancucci Sabatinelli, 1993
Popillia gemma (Newman, 1839)⇒ Popillia pulchra Arrow, 1913
Popillia japonica Newman, 1838Japanese beetle
Popillia laevicollis Kraatz, 1892
Popillia laevistriata Arrow, 1913
Popillia petrarcai Sabatinelli, 1994
Popillia pulchra Arrow, 1913
Popillia sikkimensis Lin, 1987
Popillia vignai Sabatinelli, 1994
Popillia wittmeri Frey, 1975
Popillia zerchei Sabatinelli, 1994
Propomacrus muramotoae Fujioka, 2007
Protaetia caudata caudata
Pseudopanotrogus pilifera Moser, 1915
Pseudopanotrogus rosettae Frey, 1971
Rhyparus peninsularis Arrow, 1905
Rhyssemus sinuaticollis Pittino, 1984
Saprosites japonicus Waterhouse, 1875
Serica (Serica) angustatotibialis Ahrens, 1999
Serica (Serica) chasilakhae Ahrens, 1999
Serica (Serica) eberti Frey, 1965
Serica (Serica) filitarsata Ahrens, 1999
Serica (Serica) incognita Ahrens, 1999
Serica (Serica) khajiaris Mittal, 1988
Serica (Serica) thibetana Brenske, 1897
Serica exhausta Ahrens & Fab..., 2011
Serica pelelaensis Ahrens & Fab..., 2011
Serica rectidens Ahrens & Fab..., 2009
Serica sticta Ahrens & Fab..., 2009
Serica zianii Sreedevi, Sp..., 2018
Silluvia gogona Stebnicka, 1977
Sisyphus neglectus Gory, 1833
Sophrops burmanicus Moser, 1913
Sophrops cotesi Brenske, 1892
Sophrops setosus Brenske, 1892
Sophrops umbilicatus Frey, 1973
Spinohoplia ahrensis Sabatinelli, 1997
Sybacodes simplicicollis aureopilosus Schmidt, 1909
Sybacodes simplicicollis simplicicollis
Synapsis brahminus Hope, 1831
Synapsis gilleti Arrow, 1931
Taeniodera batillifera Bourgoin, 1914
Tetraserica brahmaputrae Ahrens, 2004
Teuchestes analis Fabricius, 1787
Tibiodrepanus setosus Wiedemann, 1823
Torynorrhina apicalis Westwood, 1842
Torynorrhina distincta Hope, 1841
Torynorrhina hyacinthina Hope, 1841
Torynorrhina opalina Hope, 1831
Trigonophorus gracilipes Westwood, 1845
Trigonophorus nepalensis Hope, 1831
Trioserica tarsata Brenske, 1894
Trypoxylus (Allomyrina) dichotomus (Linnaeus, 1771)⇒ Allomyrina dichotoma Linnaeus, 1771
Xylotrupes mniszechi Thomson, 1859
Xylotrupes mniszechi mniszechi

Subnodes of Scarabaeidae with occurrences in Bhutan
SubfamilyAphodiinae
Leach, 1815
Aphodiine Dung Beetles, Small dung beetles
19
21
Genera
Species

Taxom

SubfamilyCetoniinae
Leach, 1815
Flower beetles, Flower chafers, Flower scarabs, Fruit and flower chafers
26
37
Genera
Species

Taxom

SubfamilyDynastinae
MacLeay, 1819
Horn beetles, Rhinoceros beetles
4
5
Genera
Species

Taxom

SubfamilyEuchirinae
Hope, 1840
2
3
Genera
Species

Taxom

SubfamilyMelolonthinae
Samouelle, 1819
May Beetles and Junebugs, White grubs
29
95
Genera
Species

Taxom

SubfamilyRutelinae
MacLeay, 1819
Shining Flower Chafers, Shining Leaf Chafers
11
67
Genera
Species

Taxom

SubfamilyScarabaeinae
Latreille, 1802
Dung Beetles, True dung beetles
8
22
Genera
Species

Taxom



Scarabaeidae (Latreille 1802)
Scarabaeidae (Latreille 1802)


Links, sources and further information
Scarabaeidae at artfakta.se
Scarabaeidae at dyntaxa.se
Scarabaeidae at gbif.org
Scarabaeidae at irmng.org
Scarabaeidae at itis.gov