Checklist for Family Scarabaeidae Latreille, 1802 (Dung beetles)
and region Japan (East Asia)
Class
| Insecta
Linnaeus, 1758
Insects, True insects | 696
12064
41018 | Families
Genera
Species |
Taxom
|
Subclass
| Pterygota
Lang, 1888
Winged insects | 693
12055
41003 | Families
Genera
Species |
Taxom
|
Infraclass
| Neoptera
van der Wulp, 1890
Modern Wing-folding insects | 664
11905
40649 | Families
Genera
Species |
Taxom
|
Superorder
| Holometabola
Heider, 1889
Holometabolous Insects | 434
9807
35125 | Families
Genera
Species |
Taxom
|
Order
| Coleoptera
Linnaeus, 1758
Beetles, Coleopterans | 130
3704
14154 | Families
Genera
Species |
Taxom
|
Suborder
| Polyphaga
Emery, 1886
Polyphags | 119
3331
11800 | Families
Genera
Species |
Taxom
|
Infraorder
| Scarabaeiformia
Crowson, 1960
| 12
301
1125 | Families
Genera
Species |
Taxom
|
Superfamily
| Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles | 12
301
1125 | Families
Genera
Species |
Taxom
| | Family | Scarabaeidae
Latreille, 1802
Dung beetles, Flower beetles, Fruit chafers, Rain beetles | 218
739 | Genera
Species |
Taxom
|
Genera for Japan
218 Genera with 739 Species #ABCDEGHLMNOPRSTVXY
| Acanthobodilus Dellacasa, 1922 | | 1 | Species | Taxom | | Acrossus Mulsant, 1960 | | 11 | Species | Taxom | | Adeloparius Schmidt, 1942 | | 1 | Species | Taxom | | Adoretus Laporte, 1875 | | 6 | Species | Taxom | | Aegialia Latreille, 1986 | | 7 | Species | Taxom | | Aganocrossus Reitter, 1862 | | 3 | Species | Taxom | | Agoliinus Schmidt, 1981 | | 6 | Species | Taxom | | Agrilinus Mulsant & Rey, 1875 | | 10 | Species | Taxom | Aleurostictus Kirby, 1827 | | 0 | Species | Taxom | | Alissonotum Arrow, 1847 | | 4 | Species | Taxom | | Alleurostictus | | 1 | Species | Taxom | | Allomyrina Arrow, 1771 | | 3 | Species | Taxom | | Amidorus Mulsant & Rey, 1911 | | 1 | Species | Taxom | | Ammoecius Mulsant, 1960 | | 1 | Species | Taxom | Anisorrhina Westwood, 1842 | | 0 | Species | Taxom | | Anomala Samouelle, 1959 | Shining Leaf Chafer | 52 | Species | Taxom | | Anoplognathus Leach, 1817 | Christmas beetle | 3 | Species | Taxom | | Anthracophora Burmeister, 1842 | | 1 | Species | Taxom | | Aparammoecius Petrovitz, 1875 | | 3 | Species | Taxom | | Aphodaulacus Kozhantschikov, 1875 | | 2 | Species | Taxom | | Aphodiellus Schmidt, 1875 | | 1 | Species | Taxom | | Aphodius Hellwig, 1951 | | 42 | Species | Taxom | | Apogonia Melichar, 1940 | | 17 | Species | Taxom | | Ataenius Harold, 1861 | | 5 | Species | Taxom | | Atys Reiche, 1859 | | 1 | Species | Taxom | | Augosoma Burmeister, 1775 | | 1 | Species | Taxom | | Aulacoserica Brenske, 1968 | | 1 | Species | Taxom | | Autoserica Brenske, 1927 | Garden beetle | 1 | Species | Taxom | | Blitopertha Reitter, 1878 | | 2 | Species | Taxom | | Bodiloides | | 1 | Species | Taxom | | Bodilopsis Adám, 0 | | 5 | Species | Taxom | | Brachiaphodius Koshantschikov, 1874 | | 2 | Species | Taxom | | Brahmina Blanchard, 1965 | | 3 | Species | Taxom | | Bunbunius Nomura, 1970 | | 1 | Species | Taxom | | Caccobius Thomson, 1936 | | 15 | Species | Taxom | | Caccocnemus Jekel, 1855 | | 1 | Species | Taxom | | Caccophilus Jekel, 1935 | | 1 | Species | Taxom | | Caelius Lewis, 1887 | | 2 | Species | Taxom | Caelorrhina Burmeister, 1843 | | 0 | Species | Taxom | | Calamosternus Motschulsky, 1875 | | 2 | Species | Taxom | | Calloodes White, 1912 | | 1 | Species | Taxom | | Callynomes Mohnike, 1973 | | 1 | Species | Taxom | | Carinaulus Tesar, 1935 | | 1 | Species | Taxom | | Cetonia Fabricius, 0 | | 8 | Species | Taxom | | Chaetadoretus Ohaus, 1982 | | 2 | Species | Taxom | | Chalcosoma Hope, 1789 | | 2 | Species | Taxom | | Charitovalgus Kolbe, 1913 | | 2 | Species | Taxom | | Cheirotonus Hope, 1989 | | 2 | Species | Taxom | | Chilothorax Motschulsky, 1875 | | 7 | Species | Taxom | Chlorocala Kirby & Spence, 1828 | | 0 | Species | Taxom | | Chrysina Kirby, 1990 | Jewel scarabs | 1 | Species | Taxom | | Clinterocera Motschulsky, 1853 | | 3 | Species | Taxom | | Coelodera Géné, 1972 | | 1 | Species | Taxom | | Coenochilus Burmeister &..., 1874 | | 1 | Species | Taxom | | Coilodera Hope, 1972 | | 2 | Species | Taxom | | Colobopterus Mulsant, 1847 | | 3 | Species | Taxom | | Copris Geoffroy, 1860 | Mexican dung beetle | 8 | Species | Taxom | | Coptochiroides Balthasar, 0 | | 2 | Species | Taxom | Coptomia Burmeister, 1842 | | 0 | Species | Taxom | | Cosmiomorpha Saunders, 1964 | | 4 | Species | Taxom | | Cotinis Burmeister, 1883 | Green June Beetles | 1 | Species | Taxom | | Craterocyphus Schmidt, 1847 | | 1 | Species | Taxom | | Cryptoscatomaseter Gordon and S..., 2001 | | 1 | Species | Taxom | | Dasylepida Moser, 1927 | | 1 | Species | Taxom | | Dasyvalgus Kolbe, 1941 | | 5 | Species | Taxom | | Dialytes Harold, 1909 | | 1 | Species | Taxom | Diasticus Mulsant, 1842 | | 0 | Species | Taxom | | Dicronocephalus Hope, 1914 | | 1 | Species | Taxom | | Dicronorrhina Burmeister, 1773 | | 1 | Species | Taxom | | Didactylia Orbigny, 1961 | | 1 | Species | Taxom | | Digitonthophagus Balthasar, 1787 | | 1 | Species | Taxom | | Dilochrosis Thomson, 1871 | | 1 | Species | Taxom | | Diplotaxis Kirby, 1932 | Wall-Rockets | 1 | Species | Taxom | | Ectinohoplia Redtenbacher, 1860 | | 3 | Species | Taxom | | Eophileurus Arrow, 1964 | | 4 | Species | Taxom | | Epitrichius Tagawa, 1897 | | 1 | Species | Taxom | | Erytus Mulsant & Rey, 1892 | | 2 | Species | Taxom | | Esymus Mulsant & Rey, 1789 | | 2 | Species | Taxom | | Eucetonia Schoch, 1880 | | 2 | Species | Taxom | | Eudicella White, 1836 | | 1 | Species | Taxom | | Eupoecila Burmeister, 1805 | | 1 | Species | Taxom | | Euselates J. Thomson, 1915 | | 1 | Species | Taxom | | Exomala Reitter, 1903 | | 5 | Species | Taxom | | Gametis Burmeister, 1835 | | 5 | Species | Taxom | | Gastromaladera | | 1 | Species | Taxom | | Gastroserica Brenske, 1895 | | 2 | Species | Taxom | | Gibbonthophagus Balthasar, 1875 | | 1 | Species | Taxom | | Gilletianus Balthasar, 1920 | | 2 | Species | Taxom | | Glycyphana Burmeister, 1914 | | 6 | Species | Taxom | | Glycyphaniola Miksic, 1942 | | 1 | Species | Taxom | | Gnorimus Le Peletier ..., 1887 | Chafer | 4 | Species | Taxom | | Granida Motschoulsky, 1950 | | 1 | Species | Taxom | Gymnetis MacLeay, 1819 | | 0 | Species | Taxom | | Heliocopris Hope, 1968 | | 1 | Species | Taxom | | Heptophylla Motschoulsky, 1963 | | 5 | Species | Taxom | | Heptophyllla Motschulsky, 1857 | | 1 | Species | Taxom | | Heterorrhina Westwood, 1873 | | 1 | Species | Taxom | | Heterosoma Schaum, 1901 | | 1 | Species | Taxom | | Hexataenius Fairmaire, 1891 | | 1 | Species | Taxom | | Hilyotrogus Fairmaire, 1970 | | 1 | Species | Taxom | | Holotrichia Hope, 1956 | | 18 | Species | Taxom | | Hoplia Illiger, 1959 | Monkey Beetles | 7 | Species | Taxom | | Hoplomaladera | | 1 | Species | Taxom | | Hybovalgus Kolbe, 1939 | | 1 | Species | Taxom | | Labarrus Mulsant and Rey, 1860 | | 1 | Species | Taxom | | Lachnosterna Hope, 1856 | | 1 | Species | Taxom | | Lasiotrichius Reitter, 2007 | | 3 | Species | Taxom | | Leiopsammodius Rakovic, 1878 | | 1 | Species | Taxom | | Lepidiota Kirby, 0 | | 1 | Species | Taxom | | Liatongus Reitter, 1937 | | 3 | Species | Taxom | | Liocola Thomson, 1792 | | 1 | Species | Taxom | | Lomaptera Gory & Perch..., 1899 | | 1 | Species | Taxom | | Maladera Mulsant, 2006 | | 42 | Species | Taxom | | Megaphonia Schürhoff, 1905 | | 1 | Species | Taxom | | Megasybacodes | | 1 | Species | Taxom | | Melinopterus Mulsant, 1898 | | 2 | Species | Taxom | | Melolontha Fabricius, 1964 | Billy witch | 11 | Species | Taxom | | Mesoserica Brenske, 1914 | | 1 | Species | Taxom | | Mimela Kirby, 2017 | | 23 | Species | Taxom | | Miridiba Reitter, 1913 | | 3 | Species | Taxom | | Mozartius Nomura & Nakane, 0 | | 11 | Species | Taxom | | Myrhessus Balthasar, 2006 | | 5 | Species | Taxom | | Neotrichiorhyssemus RakoviÄ & K..., 1943 | | 2 | Species | Taxom | | Neovalgus Miyake, 1941 | | 5 | Species | Taxom | | Nimbus Mulsant & Rey, 1891 | | 1 | Species | Taxom | | Nipponaphodius Nakane, 1951 | | 1 | Species | Taxom | | Nipponoagoliinus | | 1 | Species | Taxom | | Nipponoserica Nomura, 1895 | | 8 | Species | Taxom | | Nipponovalgus Sawada, 1941 | | 4 | Species | Taxom | | Octoplasia Brenske, 1996 | | 3 | Species | Taxom | | Odochilus Harold, 1971 | | 1 | Species | Taxom | Omaloplia Schönherr, 1817 | | 0 | Species | Taxom | | Oniticellus Serville, 1934 | | 1 | Species | Taxom | | Onthophagus Latreille, 1937 | | 71 | Species | Taxom | | Ophthalmoserica Brenske, 1959 | | 2 | Species | Taxom | | Oromus Mulsant & Rey, 1763 | | 1 | Species | Taxom | | Oryctes Hellwig, 1758 | Coconut rhinoceros beetles | 2 | Species | Taxom | | Osmoderma Le Peletier ..., 1887 | | 2 | Species | Taxom | | Otophorus Mulsant, 1758 | | 1 | Species | Taxom | | Oxycetonia Arrow, 1835 | | 1 | Species | Taxom | | Oxyomus Dejean, 1977 | | 1 | Species | Taxom | | Pachyserica Brenske, 1959 | | 1 | Species | Taxom | Pachystethus Blanchard, 1850 | | 0 | Species | Taxom | | Panelus Lewis, 1973 | | 4 | Species | Taxom | | Paragymnopleurus Shipp, 1875 | | 2 | Species | Taxom | | Paramaladera Nikolajev, 1974 | | 1 | Species | Taxom | | Parammoecius Seidlitz, 1960 | | 1 | Species | Taxom | | Paraphytus Harold, 1895 | | 1 | Species | Taxom | | Parascatonomus Paulian, 1823 | | 13 | Species | Taxom | | Paraschizognathus Ohaus, 1835 | | 1 | Species | Taxom | | Paraserica Reitter, 1866 | | 3 | Species | Taxom | | Parastasia Westwood, 1964 | | 8 | Species | Taxom | | Paratrichius Janson, 1864 | | 9 | Species | Taxom | | Parmaladera | | 1 | Species | Taxom | | Paulianellus Balthasar, 1938 | | 2 | Species | Taxom | | Pelidnota MacLeay, 1832 | Precious Metal Scarabs | 1 | Species | Taxom | | Pentodon Hope, 1902 | | 1 | Species | Taxom | Petrovitzius RakoviÄ, 1979 | | 0 | Species | Taxom | | Phaeaphodius Reitter, 1866 | | 1 | Species | Taxom | | Phaedimus Westwood, 1893 | | 2 | Species | Taxom | | Phalacronothus Motschulsky, 1945 | | 1 | Species | Taxom | | Pharaphodius Reitter, 1875 | | 4 | Species | Taxom | | Phyllopertha Stephens, 1943 | | 4 | Species | Taxom | | Phyllophaga Harris, 1888 | June beetles | 3 | Species | Taxom | | Plagiogonus Mulsant, 1789 | | 1 | Species | Taxom | | Planolinellus Dellacasa an..., 1825 | | 1 | Species | Taxom | | Planolinus Mulsant and Rey, 1951 | | 1 | Species | Taxom | | Platytomus Mulsant, 2006 | | 1 | Species | Taxom | | Pleuraphodius Schmidt, 1875 | | 1 | Species | Taxom | | Pleurophorus Mulsant, 1796 | | 1 | Species | Taxom | Poecilophila Kolbe, 1893 | | 0 | Species | Taxom | | Poecilophilides Kraatz, 1842 | | 1 | Species | Taxom | | Pollaplonyx Waterhouse, 1875 | | 1 | Species | Taxom | Polybathes Kirby, 1827 | | 0 | Species | Taxom | | Polyphylla Harris, 1950 | Lined June Beetles | 6 | Species | Taxom | | Popillia Leach, 1787 | | 7 | Species | Taxom | | Potosia Mulsant & Rey, 1939 | | 1 | Species | Taxom | | Proagopertha Reitter, 1875 | | 2 | Species | Taxom | | Protaetia Burmeister, 1941 | | 54 | Species | Taxom | | Psammodius Fallen, 2011 | | 5 | Species | Taxom | | Psammoporus Thomson, 1986 | | 7 | Species | Taxom | | Pseudochalcothea Ritsema, 1874 | | 1 | Species | Taxom | | Pseudosymmachia Dalle Torre, 1887 | | 1 | Species | Taxom | | Pseudotorynorrhina Miksic, 0 | | 2 | Species | Taxom | | Rakovicius Pittino, 2019 | | 4 | Species | Taxom | | Rhomborhina Hope, 0 | | 6 | Species | Taxom | | Rhomborrhina Burmeister, 1987 | | 2 | Species | Taxom | | Rhopaea Erichson, 1888 | | 1 | Species | Taxom | | Rhyparus Westwood, 1905 | | 7 | Species | Taxom | | Rhyssemus Mulsant, 1951 | | 2 | Species | Taxom | | Saprosites Redtenbacher, 2019 | | 3 | Species | Taxom | | Scabrostomus Gordon and S..., 1991 | | 1 | Species | Taxom | | Scarabaeus Linnaeus, 1773 | | 1 | Species | Taxom | | Serica MacLeay, 1937 | | 30 | Species | Taxom | | Sericania | | 47 | Species | Taxom | | Setylaides | | 3 | Species | Taxom | | Sigorus Mulsant & Rey, 1792 | | 1 | Species | Taxom | | Silluvia Landin, 1949 | | 1 | Species | Taxom | | Sinodiapterna Dellacasa, 1897 | | 1 | Species | Taxom | | Smaragdesthes Kraatz, 1836 | | 1 | Species | Taxom | | Sophrops Fairmaire, 1989 | | 7 | Species | Taxom | | Stenotothorax Schmidt, 0 | | 2 | Species | Taxom | | Subrinus Mulsant & Rey, 1870 | | 1 | Species | Taxom | | Taeniodera Burmeister, 1867 | | 1 | Species | Taxom | | Teuchestes Mulsant, 1874 | | 2 | Species | Taxom | | Torynorrhina Arrow, 1831 | | 1 | Species | Taxom | | Trematodes Falderman, 1871 | | 1 | Species | Taxom | | Trichiorhyssemus Clouët, 2007 | | 9 | Species | Taxom | | Trichius Fabricius, 1943 | Bee beetles | 4 | Species | Taxom | | Tricholontha | | 3 | Species | Taxom | | Trichoserica Reitter, 1971 | | 1 | Species | Taxom | | Trigonophorinus Pouillaude, 1897 | | 1 | Species | Taxom | | Trigonophorus Nordmann, 1897 | | 1 | Species | Taxom | | Trypoxylus Arrow, 1771 | | 6 | Species | Taxom | | Valgus Scriba, 1838 | | 1 | Species | Taxom | | Xylonychus Blanchard, 1924 | | 1 | Species | Taxom | | Xylotrupes Hope, 1767 | Elephant beetles | 1 | Species | Taxom | | Yanovalgus Nomura, 1952 | | 1 | Species | Taxom |
Species list for Japan
739 Species and 176 Subspecies | Acanthobodilus languidulus Schmidt, 1922 | | | Acrossus atratus Waterhouse, 1875 | | | Acrossus igai Nakane, 1956 | | | Acrossus igai pseudoigai Nakane & Mas..., 1967 | | | Acrossus igai sakuraii Nakane & Mas..., 1967 | | | Acrossus igai tsurugianus Nakane & Mas..., 1967 | | | Acrossus japonicus Nomura & Nakane, 1951 | | | Acrossus luridus Fabricius, 1775 | | | Acrossus rufipes Linnaeus, 1758 | Night-Flying Dung Beetle | | Acrossus superatratus Nomura & Nakane, 1951 | | | Acrossus unifasciatus Nomura & Nakane, 1951 | | | Acrossus unifasciatus kiiensis Nakane, 1960 | | | Adeloparius formosus Paulian, 1942 | | | Adoretus (Adoretus) falciungulatus Nomura, 1965 | | | Adoretus (Adoretus) lithobius Ohaus, 1914 | | | Adoretus (Adoretus) variegatus Boheman, 1857 | | | Adoretus formosanus sakishimanus Kobayashi, 1982 | | | Adoretus sinicus Burmeister, 1855 | Chinese rose beetle | | Adoretus tenuimaculatus Waterhouse, 1875 | Brown chestnut chafer | | Aegialia comis Lewis, 1895 | | | Aegialia friebi (Balthasar, 1935) | ⇒ Psammoporus friebi Balthasar, 1935 | | Aegialia kamtschaticus Motschulsky, 1860 | | | Aegialia nakanei Masumoto, 1986 | | | Aegialia nakanei horii Ochi & Kawahara, 2012 | | | Aegialia nitida Waterhouse, 1875 | | | Aegialia tsukamotoi Masumoto, 1986 | | | Aganocrossus amoenus Boheman, 1857 | | | Aganocrossus postpilosus Reitter, 1895 | | | Aganocrossus urostigma Harold, 1862 | | | Agoliinus cruentatus LeConte, 1878 | | | Agoliinus kiuchii Masumoto, 1984 | | | Agoliinus morii Nakane, 1983 | | | Agoliinus setchan Masumoto, 1984 | | | Agoliinus shibatai Nakane, 1983 | | | Agoliinus tanakai Masumoto, 1981 | | | Agrilinus breviusculus Motschulsky, 1866 | | | Agrilinus hasegawai Nomura & Nakane, 1951 | | | Agrilinus hasegawai aka Nakane, 1960 | | | Agrilinus hasegawai akahane Nakane, 1960 | | | Agrilinus hayachinensis Nomura & Nakane, 1951 | | | Agrilinus ishidai Masumoto & K..., 1987 | | | Agrilinus madara Nakane, 1960 | | | Agrilinus ritsukoae Kawai, 2004 | | | Agrilinus sordidus (Fabricius, 1775) | ⇒ Bodilopsis sordida Fabricius, 1775 | | Agrilinus uniformis Waterhouse, 1875 | | | Alissonotum cribratellum Fairmaire, 1893 | | | Alissonotum impressicolle Arrow, 1908 | Cane white grub | | Alissonotum pauper Burmeister, 1847 | Sugarcane beetle | | Alissonotum pauperum Burmeister, 1847 | | | Alleurostictus subopacus (Tauzin, 2000) | ⇒ Gnorimus subopacus Motschulsky, 1860 | | Allomyrina dichotoma Linnaeus, 1771 | Japanese horned beetle | | Allomyrina dichotoma dichotoma | | | Allomyrina dichotomus Linnaeus, 1771 | | | Amidorus koshantschikovi Jacobson, 1911 | | | Ammoecius yamato Nakane, 1960 | | | Anomala dubia aenea De Geer, 1774 | | | Anomala albopilosa Hope, 1839 | Green chafer | | Anomala albopilosa albopilosa | | | Anomala albopilosa gracilis Schönfeldt, 1890 | | | Anomala albopilosa sakishimana Nomura, 1964 | | | Anomala albopilosa yashiroi Sawada, 1950 | | | Anomala albopilosa yonaguniana Nomura, 1964 | | | Anomala antiqua Gyllenhal, 1817 | | | Anomala aulacoides Ohaus, 1915 | | | Anomala castaneoventris Bates, 1866 | | | Anomala chamaeleon Fairmaire, 1887 | | | Anomala chloroderma Arrow, 1913 | | | Anomala corruscans Chevrolat, 1841 | | | Anomala costulata Fairmaire, 1887 | | | Anomala cpustulata Matsumura, 1940 | | | Anomala cuprea (Hope, 1839) | ⇒ Anomala corruscans Chevrolat, 1841 | | Anomala daimiana Harold, 1877 | | | Anomala difficilis (Waterhouse, 1875) | ⇒ Mimela flavilabris Waterhouse, 1875 | | Anomala edentula Ohaus, 1925 | | | Anomala edentula amamiana Nomura, 1962 | | | Anomala edentula okinawana Nomura, 1965 | | | Anomala edentula yaeyamana Nomura, 1965 | | | Anomala esakii Sawada, 1950 | | | Anomala geniculata (Motschulsky, 1866) | ⇒ Anomala triangularis Schönfeldt, 1890 | | Anomala japonica Arrow, 1913 | | | Anomala limbifera Ohaus, 1915 | | | Anomala lucens Ballion, 1871 | | | Anomala luculenta (Erichson, 1847) | ⇒ Anomala dubia aenea De Geer, 1774 | | Anomala matsumurai (Sawada, 1950) | ⇒ Anomala cpustulata Matsumura, 1940 | | Anomala miyakona Nomura, 1977 | | | Anomala mongolica Faldermann, 1835 | | | Anomala motschulskyi Harold, 1877 | | | Anomala multistriata Motschulsky, 1862 | | | Anomala niijimae Ohaus, 1925 | | | Anomala octiescostata Burmeister, 1844 | | | Anomala orientalis Waterhouse, 1875 | Oriental beetle | | Anomala osakana Sawada, 1942 | | | Anomala pleurimargo Reitter, 1903 | | | Anomala puncticollis (Harold, 1877) | ⇒ Anomala albopilosa Hope, 1839 | | Anomala rufocuprea (Motschulsky, 1860) | ⇒ Anomala lucens Ballion, 1871 | | Anomala sauteri yanoi Nomura, 1959 | | | Anomala schoenfeldti Ohaus, 1915 | | | Anomala shirakii Nomura, 1959 | | | Anomala sieversi (Heyden, 1887) | ⇒ Anomala octiescostata Burmeister, 1844 | | Anomala testaceipennis (Blanchard, 1851) | ⇒ Anomala nitidula Blanchard, 1851 | | Anomala testaceipes Motschulsky, 1860 | | | Anomala triangularis Schönfeldt, 1890 | | | Anomala undulata Melsheimer, 1845 | Blossom anomala | | Anomala varians (Olivier, 1789) | ⇒ Anomala undulata Melsheimer, 1845 | | Anomala viridana (Kolbe, 1886) | ⇒ Anomala mongolica Faldermann, 1835 | | Anomala xanthopleura Arrow, 1913 | | | Anomala yanoi (Nomura, 1959) | ⇒ Anomala sauteri yanoi Nomura, 1959 | | Anoplognathus aeneus Waterhouse, 1868 | | | Anoplognathus aureus Waterhouse, 1889 | | | Anoplognathus porosus Dalman, 1817 | | | Anthracophora rusticola (Burmeister, 1842) | ⇒ Poecilophilides rusticola Burmeister, 1842 | | Aparammoecius isaburoi Nakane, 1956 | | | Aparammoecius mizo Nakane, 1967 | | | Aparammoecius pallidiligonis Waterhouse, 1875 | | | Aphodaulacus nigrotessellatus Motschulsky, 1866 | | | Aphodaulacus variabilis Waterhouse, 1875 | | | Aphodiellus impunctatus Waterhouse, 1875 | | | Aphodius (Acrossus) rufipes (Linnaeus, 1758) | ⇒ Acrossus rufipes Linnaeus, 1758 | | Aphodius (Aganocrossus) urostigma Harold, 1862 | | | Aphodius (Ahermodontus) marini Baguena, 1930 | | | Aphodius (Aphodiellus) impunctatus (Waterhouse, 1875) | ⇒ Melinopterus consputus Creutzer, 1799 | | Aphodius (Aphodius) elegans Allibert, 1847 | | | Aphodius (Aphodius) variabilis (C. O. Waterh..., 1875) | ⇒ Aphodaulacus nigrotessellatus Motschulsky, 1866 | | Aphodius (Bodilus) sordidus (Fabricius, 1775) | ⇒ Bodilopsis rufus Moll, 1782 | | Aphodius (Calamosternus) sublimbatus Motschulsky, 1860 | | | Aphodius (Colobopteroides) latus A.Schmidt, 1911 | | | Aphodius (Diapterna) hamatus Say, 1824 | | | Aphodius (Euorodalus) pusillus (Herbst, 1789) | ⇒ Esymus pusillus Herbst, 1789 | | Aphodius (Nialus) sturmi (Harold, 1870) | ⇒ Subrinus sturmi Harold, 1870 | | Aphodius (Otophorus) haemorrhoidalis (Linnaeus, 1758) | ⇒ Otophorus haemorrhoidalis Linnaeus, 1758 | | Aphodius (Paremadus) isaburoi Nakane, 1967 | | | Aphodius (Paulianellus) maderi Balthasar, 1938 | | | Aphodius (Phaeaphodius) rectus Motschulsky, 1866 | | | Aphodius (Pharaphodius) marginellus (Fabricius, 1781) | ⇒ Ataenius luteomargo Chapin, 1940 | | Aphodius (Pleuraphodius) lewisi C. O. Waterh..., 1875 | | | Aphodius (Sinodiapterna) troitzkyi Jacobson, 1897 | | | Aphodius armatus (Roth, 1851) | ⇒ Craterocyphus rhinocerus Reiche, 1847 | | Aphodius bifasciatus (Dalla Torre, 1879) | ⇒ Chilothorax melanostictus Schmidt, 1840 | | Aphodius chokaiensis (Nomura & Nakane, 1951) | ⇒ Gilletianus comatus Schmidt, 1920 | | Aphodius ealaensis (Endrödi, 1956) | ⇒ Adeloparius formosus Paulian, 1942 | | Aphodius edai (Balthasar, 1956) | ⇒ Phaeaphodius rectus Motschulsky, 1866 | | Aphodius epotus (Dellacasa, 1983) | ⇒ Chilothorax conspurcatus Linnaeus, 1758 | | Aphodius haroldianus (Balthasar, 1932) | ⇒ Colobopterus quadratus Reiche, 1847 | | Aphodius hibernalis Nakane & Tsu..., 1956 | | | Aphodius igai Nakane, 1956 | | | Aphodius informis (Mulsant, 1842) | ⇒ Acrossus luridus Fabricius, 1775 | | Aphodius irakensis (Baraud, 1977) | ⇒ Erytus opacior Koshantschikov, 1894 | | Aphodius lapponicus (Villa, 1833) | ⇒ Oromus alpinus Scopoli, 1763 | | Aphodius madari (Tesar, 1945) | ⇒ Nimbus makowskyi Koshantschikov, 1891 | | Aphodius mara (Krikken & Kaas, 1984) | ⇒ Didactylia szunyoghyi Endrödi, 1961 | | Aphodius marginatus (LeConte, 1878) | ⇒ Cryptoscatomaseter depressiusculus Schmidt, 1907 | | Aphodius mizo Nakane, 1967 | | | Aphodius nigerrimus (Dallatorre, 1879) | ⇒ Oromus alpinus Scopoli, 1763 | | Aphodius nigrotessellatus (Nakane, 1972) | ⇒ Chilothorax punctatus Waterhouse, 1875 | | Aphodius obsoleteguttatus (Waterhouse, 1875) | ⇒ Aphodaulacus nigrotessellatus Motschulsky, 1866 | | Aphodius plasoni Kaufel, 1914 | | | Aphodius rugostriatus (Reitter, 1892) | ⇒ Pharaphodius rugosostriatus Waterhouse, 1875 | | Aphodius snowi (Brown, 1929) | ⇒ Agoliinus cruentatus LeConte, 1878 | | Aphodius unifasciatus Nomura & Nakane, 1951 | | | Apogonia amida Lewis, 1896 | | | Apogonia bicarinata Lewis, 1896 | | | Apogonia bicarinata okinawana (Nomura, 1964) | ⇒ Apogonia bicarinata Lewis, 1896 | | Apogonia bicarinata yaeyamana (Nomura, 1965) | ⇒ Apogonia bicarinata Lewis, 1896 | | Apogonia bicavata (Arrow, 1913) | ⇒ Apogonia major bicavata Arrow, 1913 | | Apogonia cupreoviridis Kolbe, 1886 | | | Apogonia ishiharai Sawada, 1940 | | | Apogonia ishiharai iriei (Miyake, 1986) | ⇒ Apogonia ishiharai Sawada, 1940 | | Apogonia ishiharai yakushimana (Nomura, 1959) | ⇒ Apogonia ishiharai Sawada, 1940 | | Apogonia kamiyai Sawada, 1940 | | | Apogonia laevicollis Lansberge, 1879 | | | Apogonia major Waterhouse, 1875 | | | Apogonia major bicavata Arrow, 1913 | | | Apogonia major major | | | Apogonia niponica Lewis, 1895 | | | Apogonia splendida (Boheman, 1858) | ⇒ Apogonia laevicollis Lansberge, 1879 | | Apogonia tanigawaensis Sawada, 1940 | | | Ataenius australasiae Boheman, 1858 | | | Ataenius luteomargo Chapin, 1940 | | | Ataenius okinawensis (Nakane, 1960) | ⇒ Ataenius australasiae Boheman, 1858 | | Ataenius peregrinator Harold, 1877 | Birthworts | | Ataenius picinus Harold, 1861 | Antarctic blennies | | Atys procellana (Gould, 1859) | ⇒ Liloa procellana Gould, 1859 | | Augosoma centaurus Fabricius, 1775 | Centaurus beetle | | Aulacoserica pusilla Frey, 1968 | | | Autoserica takaoana (Niijima & Ki..., 1927) | ⇒ Maladera secreta Brenske, 1897 | | Blitopertha (Exomala) orientalis Waterhouse, 1875 | | | Blitopertha conspurcata Harold, 1878 | | | Bodiloides ictericus Laicharting, 1781 | | | Bodilopsis rufa Moll, 1782 | | | Bodilopsis rufus Moll, 1782 | | | Bodilopsis sordida Fabricius, 1775 | | | Bodilopsis sordidus Fabricius, 1775 | | | Bodilopsis sordidus sordidus | | | Brachiaphodius eccoptus Bates, 1889 | | | Brachiaphodius pilosus Harold, 1874 | | | Brahmina mikado Itoh, 1996 | | | Brahmina miyako Itoh, 1996 | | | Brahmina sakishimana Nomura, 1965 | | | Bunbunius opacipennis Nomura, 1970 | | | Caccobius (Caccophilus) unicornis Fabricius, 1798 | | | Caccobius boucomonti Balthasar, 1935 | | | Caccobius brevis Waterhouse, 1875 | | | Caccobius castaneus (Klug, 1855) | ⇒ Caccocnemus castaneus Klug, 1855 | | Caccobius hirayamai (Matsumura, 1936) | ⇒ Caccobius jessoensis Harold, 1867 | | Caccobius jessoensis Harold, 1867 | | | Caccobius jononis (Matsumura, 1936) | ⇒ Gibbonthophagus viduus Harold, 1875 | | Caccobius kelleri Olsoufieff, 1907 | | | Caccobius nikkoensis Lewis, 1895 | | | Caccobius sappoensis Matsumura, 1936 | | | Caccobius sapporensis (Matsumura, 1936) | ⇒ Caccobius jessoensis Harold, 1867 | | Caccobius sordidus Harold, 1886 | | | Caccobius suzukii (Matsumura, 1936) | ⇒ Caccobius boucomonti Balthasar, 1935 | | Caccobius yamauchii (Matsumura, 1936) | ⇒ Caccobius unicornis Fabricius, 1798 | | Caccobius yubariensis (Matsumura, 1936) | ⇒ Caccobius jessoensis Harold, 1867 | | Caccocnemus castaneus Klug, 1855 | | | Caccophilus sibiricus (Balthasar, 1935) | ⇒ Caccobius kelleri Olsoufieff, 1907 | | Caelius denticollis (Lewis, 1895) | ⇒ Silluvia elongata Landin, 1949 | | Caelius rufescens Horn, 1887 | | | Calamosternus prusai Tesar, 1945 | | | Calamosternus uniplagiatus Waterhouse, 1875 | | | Calloodes frenchi (Ohaus, 1912) | ⇒ Anoplognathus aureus Waterhouse, 1889 | | Callynomes ishikawai (Kurosawa, 1973) | ⇒ Clinterocera ishikawai Kurosawa, 1973 | | Carinaulus inexpectatus Balthasar, 1935 | | | Cetonia pilifera Motschulsky, 1860 | | | Cetonia pilifera izuensis Nomura, 1969 | | | Cetonia pilifera pilifera | | | Cetonia roelofsi Harold, 1880 | Green flower chafer | | Cetonia roelofsi gotoana Nakane, 1964 | | | Cetonia roelofsi iijimai Sakai, 1996 | | | Cetonia roelofsi ijimai Sakai, 1996 | | | Cetonia roelofsi roelofsi | | | Chaetadoretus formosanus Ohaus, 1914 | | | Chaetadoretus formosanus sakishimanus Kobayashi, 1982 | | | Chalcosoma atlas atlas Linnaeus, 1758 | | | Chalcosoma chiron (Olivier, 1789) | ⇒ Chalcosoma caucasus Fabricius, 1801 | | Charitovalgus fumosus (Lewis, 1887) | ⇒ Neovalgus fumosus Lewis, 1887 | | Charitovalgus laetus Arrow, 1913 | | | Cheirotonus jambar Kurosawa, 1984 | Yanbaru Long-armed Scarab Beetle | | Cheirotonus otai ♰ K. Ueda, 1989 | | | Chilothorax conspurcatus Linnaeus, 1758 | | | Chilothorax melanostictus Schmidt, 1840 | | | Chilothorax ohishii Masumoto, 1975 | | | Chilothorax okadai Nakane, 1951 | | | Chilothorax okadai kiritachiensis Ochi, Kawaha..., 2012 | | | Chilothorax okadai okadai | | | Chilothorax punctatus Waterhouse, 1875 | | | Chrysina difficilis Morón, 1990 | | | Clinterocera ishikawai Kurosawa, 1973 | | | Clinterocera mandarina Westwood, 1874 | | | Clinterocera scabrosa Motschulsky, 1853 | | | Coelodera helleri (Miksic, 1972) | ⇒ Coilodera helleri Miksic, 1972 | | Coenochilus striatus Westwood, 1874 | | | Coilodera formosana Moser, 1910 | | | Coilodera helleri Miksic, 1972 | | | Colobopterus erraticus Linnaeus, 1758 | Erratic small dung beetle | | Colobopterus indagator Mannerheim, 1849 | | | Colobopterus quadratus Reiche, 1847 | | | Copris (Copris) tripartitus Waterhouse, 1875 | | | Copris (Sinocopris) pecuarius Lewis, 1884 | | | Copris acutidens Motschulsky, 1860 | | | Copris brachypterus Nomura, 1964 | | | Copris brachypterus brachypterus | | | Copris brachypterus toguchii Ochi & Kawahara, 2012 | | | Copris hispanus Linnaeus, 1764 | | | Copris ochus Motschulsky, 1860 | | | Coptochiroides subcostatus Kolbe, 1886 | | | Coptochiroides subcostatus subcostatus | | | Cosmiomorpha similis Fairmaire, 1899 | | | Cosmiomorpha similis miyakoana Nomura, 1964 | | | Cosmiomorpha similis nigra Niijima & Ki..., 1927 | | | Cosmiomorpha similis yonakuniana Nomura, 1964 | | | Cotinis (Cotinis) mutabilis Gory & Perch..., 1883 | Fig beetle | | Craterocyphus rhinocerus Reiche, 1847 | | | Cryptoscatomaseter gambrinus Skelley & Go..., 2001 | Warteye stargazer | | Dasylepida ishigakiensis Niijima & Ki..., 1927 | | | Dasyvalgus angusticollis Waterhouse, 1875 | | | Dasyvalgus angusticollis maedai Nakane, 1956 | | | Dasyvalgus inouei Sawada, 1939 | | | Dasyvalgus tuberculatus Lewis, 1887 | | | Dasyvalgus yonakuniensis Sawada, 1941 | | | Dialytes foveatus (Schmidt, 1909) | ⇒ Setylaides foveatus Schmidt, 1909 | | Dicronocephalus wallichi bourgoini Pouillaude, 1914 | | | Dicronorrhina micans Drury, 1773 | | | Didactylia szunyoghyi Endrödi, 1961 | | | Digitonthophagus gazella Fabricius, 1787 | Brown dung beetle | | Dilochrosis balteata Vollenhoven, 1871 | Red spot rosechafer | | Diplotaxis pinalica Fall, 1932 | | | Ectinohoplia gracilipes Lewis, 1895 | | | Ectinohoplia obducta Motschulsky, 1857 | | | Ectinohoplia rufipes Motschulsky, 1860 | | | Eophileurus chinensis Faldermann, 1835 | | | Eophileurus chinensis chinensis Faldermann, 1835 | | | Eophileurus chinensis irregularis Prell, 1913 | | | Eophileurus chinensis okinawanus Nomura, 1964 | | | Epitrichius lagopus Fairmaire, 1897 | | | Erytus opacior Koshantschikov, 1894 | | | Erytus pruinosus Reitter, 1892 | | | Esymus pusillus Herbst, 1789 | | | Esymus pusillus pusillus Herbst, 1789 | | | Eucetonia pilifera Motschulsky, 1860 | | | Eucetonia roelofsi Harold, 1880 | | | Eudicella smithi Buquet, 1836 | | | Eupoecila australasiae Donovan, 1805 | Fiddler beetle | | Euselates proxima Bourgoin, 1915 | | | Exomala conspurcata Harold, 1878 | | | Exomala ohdaiensis Sawada, 1941 | | | Exomala okiwaensis Ohaus, 1925 | | | Exomala orientalis C. O. Waterh..., 1875 | Asiatic beetle | | Exomala pallidipennis Reitter, 1903 | | | Gametis forticula Janson, 1881 | | | Gametis forticula forticula | | | Gametis forticula kotoensis Nomura, 1959 | | | Gametis forticula yonakuniana Nomura, 1959 | | | Gametis jucunda Faldermann, 1835 | Scarab | | Gastromaladera major Nomura, 1959 | | | Gastroserica brevicornis Lewis, 1895 | | | Gastroserica higonia Lewis, 1895 | | | Gibbonthophagus viduus Harold, 1875 | | | Gilletianus atsushii Ochi, 1986 | | | Gilletianus comatus Schmidt, 1920 | | | Glycyphana cincta Wallace, 1867 | | | Glycyphana fulvistemma Motschulsky, 1858 | | | Glycyphana gracilis (Sawada, 1942) | ⇒ Glycyphaniola gracilis Sawada, 1942 | | Glycyphana gracilis viridis Sawada, 1942 | | | Glycyphana hybrida Miksic, 1970 | | | Glycyphana tonkinensis Moser, 1914 | | | Glycyphaniola gracilis Sawada, 1942 | | | Gnorimus otsukae (Niijima & Ki..., 1923) | ⇒ Paratrichius doenitzi Harold, 1879 | | Gnorimus subopacus Motschulsky, 1860 | | | Gnorimus subopacus subopacus | | | Gnorimus subopacus viridiopacus Lewis, 1887 | | | Granida albolineata (Sawada, 1950) | ⇒ Polyphylla albolineata Motschulsky, 1861 | | Heliocopris antiquus ♰ I. Fujiyama, 1968 | | | Heptophylla picea (Motschulsky, 1857) | ⇒ Heptophyllla picea Motschulsky, 1857 | | Heptophylla picea iriei Kusui, 1971 | | | Heptophylla picea maenamii Nomura, 1969 | | | Heptophylla picea picea | | | Heptophylla tosana Miyatake, 1963 | | | Heptophyllla picea Motschulsky, 1857 | | | Heterorrhina paupera Mohnike, 1873 | | | Heterosoma exasperata Fairmaire, 1901 | | | Hexataenius protensus Fairmaire, 1891 | | | Hilyotrogus yasuii Nomura, 1970 | | | Holotrichia amamiana Nomura, 1964 | | | Holotrichia aritai Nomura, 1964 | | | Holotrichia convexopyga Moser, 1912 | | | Holotrichia danjoensis Miyake & Ima..., 1982 | | | Holotrichia diomphalia Bates, 1888 | | | Holotrichia ernesti Reitter, 1902 | | | Holotrichia kiotoensis (Niijima & Ki..., 1923) | ⇒ Holotrichia kiotonensis Brenske, 1894 | | Holotrichia kiotonensis Brenske, 1894 | | | Holotrichia koraiensis Murayama, 1937 | | | Holotrichia lata Brenske, 1892 | | | Holotrichia loochooana Sawada, 1950 | | | Holotrichia loochooana loochooana | | | Holotrichia loochooana okinawana Nomura, 1964 | | | Holotrichia motschulskyi Saylor, 1937 | | | Holotrichia niponensis Lewis, 1895 | | | Holotrichia parallela (Motschulsky, 1854) | ⇒ Holotrichia kiotonensis Brenske, 1894 | | Holotrichia picea Waterhouse, 1875 | | | Holotrichia tokara Nakane, 1956 | | | Hoplia aureola Pallas, 1781 | | | Hoplia communis Waterhouse, 1875 | Long-legged chafer | | Hoplia hakonensis Sawada, 1938 | | | Hoplia moerens Waterhouse, 1875 | | | Hoplia parvula Krynicki, 1832 | | | Hoplia reinii Heyden, 1878 | | | Hoplia shirakii Nomura, 1959 | | | Hoplomaladera saitoi Kobayashi, 1975 | | | Hybovalgus major Sawada, 1939 | | | Labarrus sublimbatus Motschulsky, 1860 | | | Lachnosterna sororia (LeConte, 1856) | ⇒ Phyllophaga micans Knoch, 1801 | | Lasiotrichius succinctus Pallas, 1781 | | | Lasiotrichius succinctus hanaoi Yawata, 1943 | | | Lasiotrichius succinctus tokushimus Krajcik, 2007 | | | Leiopsammodius japonicus Harold, 1878 | | | Lepidiota trichosternus | | | Liatongus minutus Motschulsky, 1860 | | | Liatongus phanaeoides Westwood, 1839 | | | Liatongus phanaeoides fumiroi (Yohena, 1937) | ⇒ Liatongus minutus Motschulsky, 1860 | | Liocola marmorata Fabricius, 1792 | | | Lomaptera pseudorufa Heller, 1899 | | | Maladera (Aserica) japonica Motschulsky, 1857 | Velvety chafer | | Maladera japonica japonica | | | Maladera japonica yaeyamana (Nomura, 1963) | ⇒ Maladera yaeyamana Nomura, 1963 | | Maladera (Cephaloserica) castanea Arrow, 1913 | Asian garden beetle | | Maladera (Eumaladera) nitididorsis Nomura, 1967 | | | Maladera nitididorsis nitididorsis | | | Maladera nitididorsis ootsuboi Kobayashi, 2009 | | | Maladera amamiana Nomura, 1959 | | | Maladera cariniceps Moser, 1915 | | | Maladera ejimai Miyake & Ima..., 1987 | | | Maladera formosae Brenske, 1898 | Asiatic Garden Beetle | | Maladera hiemalis Kobayashi, 2014 | | | Maladera hiranoi Miyake, 1986 | | | Maladera horii Kobayashi, 2010 | | | Maladera imasakai Miyake & Yamaya, 1995 | | | Maladera impressithorax Nomura, 1973 | | | Maladera impressithorax okinoerabuana (Kobayashi, 1978) | ⇒ Maladera okinoerabuana Kobayashi, 1978 | | Maladera inadai Kobayashi, 2010 | | | Maladera kamiyai Sawada, 1937 | | | Maladera kawaharai Kobayashi, 2010 | | | Maladera kawaii Kobayashi, 2010 | | | Maladera kunigami Kobayashi, K..., 2006 | | | Maladera kusuii Miyake, 1986 | | | Maladera nitidiceps Nomura, 1967 | | | Maladera nitidipes Moser, 1915 | | | Maladera okinawaensis Kobayashi, 1978 | | | Maladera okinoerabuana Kobayashi, 1978 | | | Maladera opima Nomura, 1967 | | | Maladera orientalis Motschulsky, 1857 | Smaller velvety chafer | | Maladera oshimana Nomura, 1962 | | | Maladera ovatula Fairmaire, 1891 | | | Maladera renardi Ballion, 1870 | | | Maladera satoi Niijima & Ki..., 1923 | | | Maladera secreta Brenske, 1897 | | | Maladera secreta secreta | | | Maladera shimogana Ahrens & Fab..., 2016 | | | Maladera spissigrada Brenske, 1897 | | | Maladera taiheii Kobayashi, 2016 | | | Maladera tokunoshimana Kobayashi, K..., 2006 | | | Maladera yaeyamana Nomura, 1963 | | | Maladera yakushimana Kobayashi, K..., 2006 | | | Maladera yonaguniensis Kobayashi, K..., 2006 | | | Megaphonia pratti Janson, 1905 | | | Megasybacodes brevitarsis Kakizoe, Mar..., 2019 | | | Melinopterus consputus Creutzer, 1799 | | | Melinopterus scuticollis Semenov, 1898 | | | Melolontha (Melolontha) melolontha Linnaeus, 1758 | Cockchafer | | Melolontha frater Arrow, 1913 | | | Melolontha frater frater | | | Melolontha incana Motschulsky, 1853 | | | Melolontha insulana Burmeister, 1939 | | | Melolontha japonica Burmeister, 1855 | | | Melolontha masafumii Nomura, 1952 | | | Melolontha satsumaensis Niijima & Ki..., 1923 | | | Melolontha satsumaensis satsumaensis | | | Melolontha satsumaensis shikokuana Nomura, 1977 | | | Melolontha tamina Nomura, 1964 | | | Mesoserica (Mesoserica) sachalinensis Matsumura, 1914 | | | Mimela chinensis Kirby, 1825 | | | Mimela confucius Hope, 1836 | | | Mimela confucius ishigakiensis Sawada, 1950 | | | Mimela costata Hope, 1839 | | | Mimela difficilis C. O. Waterh..., 1875 | | | Mimela flavilabris Waterhouse, 1875 | | | Mimela heterochropus hopei Burmeister, 1855 | | | Mimela holosericea Kirby, 1825 | | | Mimela holosericea japonica (Machatschke, 1952) | ⇒ Mimela holosericea Kirby, 1825 | | Mimela horsfieldi Hope, 1836 | | | Mimela ignicauda Bates, 1866 | | | Mimela inoei Nomura, 1967 | | | Mimela peckinensis Heyden, 1886 | | | Mimela sakishimana Nomura, 1973 | | | Mimela splendens (Gyllenhal, 1817) | ⇒ Mimela heterochropus hopei Burmeister, 1855 | | Mimela takemurai Sawada, 1942 | | | Mimela testacea Lin, 1990 | | | Mimela testaceipes Motschulsky, 1860 | Striated chafer | | Mimela testaceipes testaceipes | | | Mimela testaceipes yuasai Nomura, 1969 | | | Mimela testaceoviridis Blanchard, 1851 | | | Mimela yonaguniensis Nomura, 1965 | | | Mimela yotakemurai Kobayashi, 2017 | | | Miridiba castanea Waterhouse, 1875 | | | Miridiba hirsuta Itoh, 2001 | | | Miridiba tuberculipennis Moser, 1913 | | | Mozartius jugosus Lewis, 1895 | | | Mozartius jugosus horii Ochi, Kawaha..., 2006 | | | Mozartius jugosus jugosus | | | Mozartius jugosus shingoi Ochi, Kawaha..., 2006 | | | Mozartius kyushuensis Ochi & Kawah..., 2002 | | | Mozartius testaceus Nomura & Nakane, 1951 | | | Mozartius testaceus shikokuensis Masumoto, 1984 | | | Mozartius testaceus testaceus | | | Mozartius uenoi Masumoto, 1984 | | | Mozartius uenoi hadai Kawai, 2003 | | | Mozartius uenoi uenoi | | | Myrhessus kubotai Ochi, Kawaha..., 2012 | | | Myrhessus nishinoi Ochi, Kawaha..., 2012 | | | Myrhessus samurai Balthasar, 1941 | | | Myrhessus sunadai Ochi, Kawaha..., 2019 | | | Myrhessus yorikoae Ochi, Kawaha..., 2006 | | | Neotrichiorhyssemus boninensis Nakane, 1960 | | | Neotrichiorhyssemus esakii Nomura, 1943 | | | Neovalgus fumosus Lewis, 1887 | | | Neovalgus laetus Arrow, 1913 | | | Neovalgus laetus laetus | | | Neovalgus laetus ootsuboi Kobayashi, 2011 | | | Neovalgus sikokuensis Sawada, 1941 | | | Nimbus makowskyi Koshantschikov, 1891 | | | Nipponaphodius gotoi Nomura & Nakane, 1951 | | | Nipponoagoliinus yasutakai Ochi & Kawahara, 2001 | | | Nipponoserica daisenensis Sawada, 1937 | | | Nipponoserica daisensis Sawada, 1937 | | | Nipponoserica gomadana Nomura, 1976 | | | Nipponoserica kunitachiana Nomura, 1976 | | | Nipponoserica peregrina Chapin, 1938 | Picture rockskipper | | Nipponoserica pubiventris Nomura, 1976 | | | Nipponoserica setiventris Nomura, 1976 | | | Nipponoserica similis Lewis, 1895 | Beetle | | Nipponovalgus angusticollis C. O. Waterh..., 1875 | | | Nipponovalgus angusticollis angusticollis | | | Nipponovalgus angusticollis maedai Nakane, 1956 | | | Nipponovalgus yonakuniensis (Sawada, 1941) | ⇒ Dasyvalgus yonakuniensis Sawada, 1941 | | Octoplasia basilanensis Itoh, 1996 | | | Octoplasia prolixa Arrow, 1906 | | | Octoplasia pseudogigantea (Itoh, 1996) | ⇒ Octoplasia prolixa Arrow, 1906 | | Odochilus convexus Nomura, 1971 | | | Oniticellus yohenai (Matsumura, 1934) | ⇒ Liatongus phanaeoides Westwood, 1839 | | Onthophagus (Euonthophagus) carbonarius Klug, 1855 | | | Onthophagus (Gibbonthophagus) apicetinctus (d´Orbigny, 1900) | ⇒ Onthophagus atripennis Waterhouse, 1875 | | Onthophagus (Gibbonthophagus) atripennis Waterhouse, 1875 | | | Onthophagus (Gibbonthophagus) viduus (Harold, 1875) | ⇒ Onthophagus orbus Boucomont, 1924 | | Onthophagus (Indachorius) suginoi Ochi, 1984 | | | Onthophagus (Matashia) yubarinus (Matsumura, 1937) | ⇒ Onthophagus fodiens Waterhouse, 1875 | | Onthophagus (Onthophagus) acuticollis (Gillet, 1927) | ⇒ Onthophagus amphioxus Arrow, 1933 | | Onthophagus (Onthophagus) lutosopictus Fairmaire, 1897 | | | Onthophagus (Onthophagus) sauteri Gillet, 1924 | | | Onthophagus (Onthophagus) trituber Wiedemann, 1823 | | | Onthophagus (Palaeonthophagus) gibbulus Pallas, 1781 | | | Onthophagus gibbulus gibbulus | | | Onthophagus (Palaeonthophagus) olsoufieffi Boucomont, 1924 | | | Onthophagus (Paraphanaeomorphus) argyropygus Gillet, 1927 | | | Onthophagus (Parascatonomus) koichii (Ochi & Araya, 1994) | ⇒ Caccobius jessoensis Harold, 1867 | | Onthophagus (Parascatonomus) miyakei Ochi & Araya, 1992 | | | Onthophagus (Parascatonomus) murasakianus Nomura, 1976 | | | Onthophagus (Phanaeomorphus) ater Waterhouse, 1875 | | | Onthophagus (Phanaeomorphus) lenzi Harold, 1875 | | | Onthophagus (Strandius) japonicus Harold, 1875 | | | Onthophagus aequiperus Matsumura & ..., 1937 | | | Onthophagus akirai (Matsumura & ..., 1937) | ⇒ Onthophagus atripennis Waterhouse, 1875 | | Onthophagus amamiensis Nomura, 1965 | | | Onthophagus amamiensis amamiensis | | | Onthophagus amamiensis matsumurai Ochi, Kawaha..., 2019 | | | Onthophagus amphioxus Arrow, 1933 | | | Onthophagus basicruentatus Goidanich, 1926 | | | Onthophagus bivertex Heyden, 1887 | | | Onthophagus carpophilus Pereira & Ha..., 1961 | | | Onthophagus chibanus Matsumura & ..., 1937 | | | Onthophagus chuzenjianus (Matsumura & ..., 1937) | ⇒ Onthophagus ater Waterhouse, 1875 | | Onthophagus fodiens Waterhouse, 1875 | | | Onthophagus hikosanus (Matsumura & ..., 1937) | ⇒ Onthophagus fodiens Waterhouse, 1875 | | Onthophagus ibonus (Matsumura & ..., 1937) | ⇒ Onthophagus atripennis Waterhouse, 1875 | | Onthophagus itoi Nomura, 1976 | | | Onthophagus jedensis Matsumura, 1937 | | | Onthophagus kandai (Matsumura, 1937) | ⇒ Onthophagus olsoufieffi Boucomont, 1924 | | Onthophagus kawarinus (Matsumura & ..., 1937) | ⇒ Onthophagus ater Waterhouse, 1875 | | Onthophagus kogatanus (Matsumura & ..., 1937) | ⇒ Onthophagus atripennis Waterhouse, 1875 | | Onthophagus komabellus Matsumura & ..., 1937 | | | Onthophagus kozunonis (Matsumura, 1937) | ⇒ Onthophagus bivertex Heyden, 1887 | | Onthophagus lenzii Harold, 1874 | | | Onthophagus malthinus Gillet, 1930 | | | Onthophagus matsuokai (Matsumura & ..., 1937) | ⇒ Onthophagus sauteri Gillet, 1924 | | Onthophagus minoi Matsumura & ..., 1937 | | | Onthophagus minokuchianus (Matsumura, 1934) | ⇒ Onthophagus bivertex Heyden, 1887 | | Onthophagus misujianus Matsumura & ..., 1937 | | | Onthophagus miyazakianus Matsumura & ..., 1937 | | | Onthophagus nitidulus Klug, 1845 | | | Onthophagus nitidus Waterhouse, 1875 | | | Onthophagus ocellatopunctatus Waterhouse, 1875 | | | Onthophagus ohbayashii Nomura, 1939 | | | Onthophagus oishii Matsumura & ..., 1937 | | | Onthophagus okushiranus Matsumura & ..., 1937 | | | Onthophagus okushirianus Matsumura, 1937 | | | Onthophagus oniellus (Matsumura, 1937) | ⇒ Onthophagus bivertex Heyden, 1887 | | Onthophagus orbus Boucomont, 1924 | | | Onthophagus oshimanus Nakane, 1960 | | | Onthophagus shibatai Nakane, 1960 | | | Onthophagus shigeo (Matsumura & ..., 1937) | ⇒ Onthophagus atripennis Waterhouse, 1875 | | Onthophagus shigeoi Matsumura, 1937 | | | Onthophagus shinanensis (Matsumura, 1934) | ⇒ Onthophagus bivertex Heyden, 1887 | | Onthophagus shirakii Nakane, 1960 | | | Onthophagus shurianus Matsumura & ..., 1937 | | | Onthophagus sobosanus (Matsumura & ..., 1937) | ⇒ Onthophagus fodiens Waterhouse, 1875 | | Onthophagus takabayashii Matsumura, 1937 | | | Onthophagus uedanus (Matsumura, 1937) | ⇒ Onthophagus olsoufieffi Boucomont, 1924 | | Onthophagus ushiodai (Matsumura & ..., 1937) | ⇒ Onthophagus fodiens Waterhouse, 1875 | | Onthophagus yakuinsulanus Nakane, 1984 | | | Onthophagus yugianus (Matsumura & ..., 1937) | ⇒ Onthophagus fodiens Waterhouse, 1875 | | Onthophagus yumigatanus Matsumura, 1937 | | | Ophthalmoserica elongata (Nomura, 1959) | ⇒ Serica nitididorsis opacidorsis Nomura, 1972 | | Ophthalmoserica sawadai (Nomura, 1959) | ⇒ Serica takagii Sawada, 1937 | | Oromus alpinus Scopoli, 1763 | | | Oryctes hisamatsui Nagai, 2002 | | | Oryctes rhinoceros Linnaeus, 1758 | Asiatic rhinoceros beetle | | Osmoderma opicum Lewis, 1887 | | | Osmoderma opicum opicum Lewis, 1887 | | | Otophorus haemorrhoidalis Linnaeus, 1758 | Bloody small dung beetle | | Oxycetonia jucunda (Falderman, 1835) | ⇒ Gametis jucunda Faldermann, 1835 | | Oxyomus ishidai Nakane, 1977 | | | Pachyserica yanoi Nomura, 1959 | | | Panelus kubotai Kawahara, In..., 2007 | | | Panelus ovatus Nomura, 1973 | | | Panelus parvulus Waterhouse, 1874 | | | Panelus rufulus Nomura, 1973 | | | Paragymnopleurus ambiguus Janssens, 1945 | | | Paragymnopleurus stipes Sharp, 1875 | | | Paramaladera major Nomura, 1974 | | | Parammoecius yamato (Nakane, 1960) | ⇒ Ammoecius yamato Nakane, 1960 | | Paraphytus dentifrons Lewis, 1895 | | | Parascatonomus acuticollis sakishimanus Nomura, 1976 | | | Parascatonomus aokii Nomura, 1976 | | | Parascatonomus itoi Nomura, 1976 | | | Parascatonomus itoi itoi | | | Parascatonomus itoi kumejimensis Ochi, Kawaha..., 2019 | | | Parascatonomus miyakei Ochi & Araya, 1992 | | | Parascatonomus murasakianus Nomura, 1976 | | | Parascatonomus murasakianus carnarius Nomura, 1976 | | | Parascatonomus murasakianus miyakoinsularis Ochi, Miyake..., 1999 | | | Parascatonomus murasakianus murasakianus | | | Parascatonomus nitidus (Nomura, 1976) | ⇒ Onthophagus nitidus Waterhouse, 1875 | | Parascatonomus shibatai Nakane, 1960 | | | Parascatonomus tricornis Wiedemann, 1823 | | | Paraschizognathus prasinus Boisduval, 1835 | | | Paraserica gricea (Fujioka, 2001) | ⇒ Parmaladera grisea Motschulsky, 1866 | | Paraserica griesea Motschulsky, 1866 | | | Paraserica grisea Motschulsky, 1866 | | | Parastasia ferrieri Nonfried, 1895 | | | Parastasia ferrieri boninensis Nakane, 1983 | | | Parastasia ferrieri ferrieri | | | Parastasia ferrieri ichikawai Nakane, 1983 | | | Parastasia ferrieri sakishimana Nomura, 1965 | | | Parastasia ferrieri tokarana Nomura, 1964 | | | Parastasia oberthueri Ohaus, 1900 | | | Parastasia oberthueri ishigakiana Nomura, 1964 | | | Paratrichius doenitzi Harold, 1879 | | | Paratrichius duplicatus Lewis, 1895 | | | Paratrichius duplicatus duplicatus | | | Paratrichius duplicatus okinawanus Nomura, 1959 | | | Paratrichius geibiensis Miyake, 1997 | | | Paratrichius itoi Tagawa, 1938 | | | Paratrichius kyushuensis Miyake, 1990 | | | Paratrichius osimanus Chujo, 1941 | | | Paratrichius septemdecimguttatus Snellen van ..., 1864 | | | Parmaladera grisea Motschulsky, 1866 | | | Paulianellus asahinai Nakane, 1951 | | | Paulianellus maderi Balthasar, 1938 | | | Pelidnota (Chalcoplethis) kirbyi Gray, 1832 | | | Pentodon idiota memnon Jakovlev, 1902 | | | Phaeaphodius rectus Motschulsky, 1866 | | | Phaedimus cumingi (Waterhouse, 1841) | ⇒ Phaedimus mohnikei Kraatz, 1893 | | Phaedimus mohnikei Kraatz, 1893 | | | Phalacronothus botulus Balthasar, 1945 | | | Pharaphodius marginellus Fabricius, 1781 | | | Pharaphodius orientalis Harold, 1862 | | | Pharaphodius priscus Motschulsky, 1858 | | | Pharaphodius rugosostriatus Waterhouse, 1875 | | | Phyllopertha diversa Waterhouse, 1875 | | | Phyllopertha intermixta Arrow, 1913 | | | Phyllopertha irregularis (Waterhouse, 1875) | ⇒ Phyllopertha intermixta Arrow, 1913 | | Phyllopertha suzukii Sawada, 1943 | | | Phyllophaga (Phyllophaga) diffinis Blanchard, 1851 | | | Phyllophaga (Phyllophaga) panorpa Sanderson, 1950 | | | Phyllophaga polyphylla Bates, 1888 | | | Plagiogonus arenarius Olivier, 1789 | | | Planolinellus vittatus Say, 1825 | | | Planolinus pratensis Nomura & Nakane, 1951 | | | Platytomus yadai Ochi, Kawaha..., 2006 | | | Pleuraphodius lewisi Waterhouse, 1875 | | | Pleurophorus caesus Creutzer, 1796 | Cut small dung beetle | | Poecilophilides rusticola Burmeister, 1842 | | | Pollaplonyx (Polloplonyx) flavidus Waterhouse, 1875 | | | Polyphylla (Granida) albolineata Motschulsky, 1861 | | | Polyphylla (Gynexophylla) laticollis Lewis, 1887 | | | Polyphylla laticollis laticollis | | | Polyphylla davidis Fairmaire, 1888 | | | Polyphylla schoenfeldti Brenske, 1890 | | | Polyphylla taiwana Sawada, 1950 | | | Popillia comma Niijima & Ma..., 1923 | | | Popillia dilutipennis Fairmaire, 1888 | | | Popillia insularis Lewis, 1895 | | | Popillia japonica Newman, 1838 | Japanese beetle | | Popillia lewisi Arrow, 1913 | | | Popillia psilopyga Ohaus, 1897 | | | Popillia quadriguttata Fabricius, 1787 | Four spotted beetle | | Potosia aerosa (Hepp, 1939) | ⇒ Protaetia metallica bourgini Ruter, 1967 | | Proagopertha ohbayashii Nomura, 1965 | | | Proagopertha pubicollis Waterhouse, 1875 | | | Protaetia (Calopotosia) descarpentriesi Ruter, 1978 | | | Protaetia (Calopotosia) inquinata Arrow, 1913 | | | Protaetia (Calopotosia) lewisi Janson, 1888 | | | Protaetia lewisi hisatoi Ochi, Toguch..., 2022 | | | Protaetia lewisi leachi Kurosawa, 1959 | | | Protaetia lewisi lewisi | | | Protaetia (Calopotosia) orientalis Gory & Perch..., 1833 | Asian flower beetle | | Protaetia orientalis orientalis Gory & Perch..., 1833 | | | Protaetia orientalis sakaii Kobayashi, 1994 | | | Protaetia orientalis tokorana Nomura, 1964 | | | Protaetia (Chalcoprotaetia) philippensis Fabricius, 1775 | | | Protaetia (Heteroprotaetia) fusca Herbst, 1790 | Asian Mango Flower Beetle | | Protaetia (Liocola) brevitarsis Lewis, 1879 | | | Protaetia brevitarsis brevitarsis | | | Protaetia (Liocola) cataphracta Arrow, 1913 | | | Protaetia (Liocola) formosana Moser, 1910 | | | Protaetia formosana nozatoi Miyake, 1986 | | | Protaetia (Liocola) insperata Lewis, 1879 | | | Protaetia (Liocola) lugubris (Herbst, 1786) | ⇒ Liocola marmorata Fabricius, 1792 | | Protaetia (Liocola) miyakoensis Niijima & Ki..., 1923 | | | Protaetia orientalis submarmorea Burmeister, 1842 | | | Protaetia (Niponoprotaetia) exasperata Fairmaire, 1898 | | | Protaetia exasperata akitai Masumoto & S..., 1988 | | | Protaetia exasperata erabuana Nomura, 1964 | | | Protaetia exasperata exasperata | | | Protaetia exasperata iheyana Sakai, 1989 | | | Protaetia exasperata nakana Nakane, 1956 | | | Protaetia exasperata nomurai Sakai, 1998 | | | Protaetia exasperata satoi Nakane, 1956 | | | Protaetia exasperata suwanoseana Nomura, 1964 | | | Protaetia exasperata uenoi Nomura, 1960 | | | Protaetia (Niponoprotaetia) lenzi Harold, 1878 | | | Protaetia (Pachyprotaetia) ciliata Olivier, 1785 | | | Protaetia metallica bourgini Ruter, 1967 | | | Protaetia (Protaetia) siamensis Nonfried, 1891 | | | Protaetia (Pseudocalopotosia) ishigakia Kurosawa, 1959 | | | Protaetia ishigakia iriomoteana Nomura, 1964 | | | Protaetia ishigakia ishigakia | | | Protaetia ishigakia kikaiana Nakane, 1960 | | | Protaetia ishigakia miyakoana Kurosawa, 1959 | | | Protaetia ishigakia miyakona Kurosawa, 1959 | | | Protaetia ishigakia okinawana Kurosawa, 1959 | | | Protaetia ishigakia yonakuniana Nomura, 1964 | | | Protaetia (Pyropotosia) oschimana Nonfried, 1895 | | | Protaetia (Pyropotosia) pryeri Janson, 1888 | Midway emerald beetle | | Protaetia pryeri akusekiana Nomura, 1964 | | | Protaetia pryeri esakii Nakane, 1956 | | | Protaetia pryeri nitidicosta Yawata, 1941 | | | Protaetia pryeri pryeri | | | Protaetia pryeri tsutsuii Nakane, 1956 | | | Protaetia (Tomentoprotaetia) culta Waterhouse, 1879 | | | Protaetia culta yaeyamana Nomura, 1965 | | | Protaetia acuminata Fabricius, 1775 | | | Protaetia nitidicosta (Yawata, 1941) | ⇒ Protaetia pryeri nitidicosta Yawata, 1941 | | Psammodius convexus Waterhouse, 1875 | | | Psammodius japonicus (Harold, 1878) | ⇒ Leiopsammodius japonicus Harold, 1878 | | Psammodius kobayashii Nomura, 1973 | | | Psammodius kondoi (Masumoto, 1984) | ⇒ Psammodius kobayashii Nomura, 1973 | | Psammodius maruyamai Ochi, Kawaha..., 2011 | | | Psammoporus comis Lewis, 1895 | | | Psammoporus friebi Balthasar, 1935 | | | Psammoporus kamtschaticus Motschulsky, 1860 | | | Psammoporus nakanei (Masumoto, 1986) | ⇒ Aegialia kamtschaticus Motschulsky, 1860 | | Psammoporus nakanei horii Ochi & Kawahara, 2012 | | | Psammoporus nakanei nakanei | | | Psammoporus tsukamotoi Masumoto, 1986 | | | Pseudochalcothea auripes Westwood, 1874 | | | Pseudosymmachia impressifrons Fairmaire, 1887 | | | Pseudotorynorrhina japonica Hope, 1841 | | | Pseudotorynorrhina japonica japonica | | | Rakovicius ainu Lewis, 1895 | | | Rakovicius coreanus Kim, 1980 | | | Rakovicius kyotoensis Ochi, Kawaha..., 2019 | | | Rakovicius takashii Ochi, Kawaha..., 2019 | | | Rhomborhina hamai Nomura, 1964 | | | Rhomborhina japonica Hope, 1841 | | | Rhomborhina polita Waterhouse, 1875 | | | Rhomborhina unicolor Motschulsky, 1861 | | | Rhomborhina unicolor fukueana Sakai, 2000 | | | Rhomborhina unicolor unicolor | | | Rhomborrhina japonica (Hope, 1841) | ⇒ Pseudotorynorrhina japonica Hope, 1841 | | Rhomborrhina kurosawai Masumoto & S..., 1987 | | | Rhopaea magnicornis Blackburn, 1888 | Brown Cockchafer | | Rhyparus amamianus (Nakane, 1956) | ⇒ Rhyparus helophoroides Fairmaire, 1893 | | Rhyparus azumai Nakane, 1956 | | | Rhyparus helophoroides Fairmaire, 1893 | | | Rhyparus kitanoi Miyake, 1982 | | | Rhyparus kitanoi kitanoi | | | Rhyparus loebli (Paulian, 1983) | ⇒ Rhyparus azumai loebli Paulian, 1983 | | Rhyparus peninsularis Arrow, 1905 | | | Rhyssemus samurai (Balthasar, 1941) | ⇒ Myrhessus samurai Balthasar, 1941 | | Rhyssemus tuberculatus (Nomura & Nakane, 1951) | ⇒ Myrhessus samurai Balthasar, 1941 | | Saprosites japonicus Waterhouse, 1875 | | | Saprosites narae Lewis, 1895 | | | Saprosites yasutakai Ochi & Kawahara, 2019 | | | Scabrostomus tanytarsus Skelley & Wo..., 1991 | Masked stargazer | | Scarabaeus aegaeon Drury, 1773 | | | Serica (Serica) boops Waterhouse, 1875 | | | Serica (Serica) brevitarsis Nomura, 1972 | | | Serica brevitarsis brevitarsis | | | Serica brevitarsis rectipes Nomura, 1972 | | | Serica (Serica) brunnea (Linnaeus, 1758) | ⇒ Serica similis Lewis, 1895 | | Serica (Serica) echigoana Nakane & Baba, 1960 | | | Serica (Serica) foobowana Sawada, 1937 | | | Serica (Serica) incurvata Nomura, 1971 | | | Serica (Serica) inexspectata Kontkanen, 1956 | | | Serica (Serica) karafutoensis Niijima & Ki..., 1923 | | | Serica karafutoensis honshuensis Nomura, 1972 | | | Serica karafutoensis karafutoensis | | | Serica (Serica) nigrovariata Lewis, 1895 | | | Serica (Serica) nipponica Nomura, 1959 | | | Serica (Serica) nitididorsis Nomura, 1971 | | | Serica nitididorsis nitididorsis | | | Serica nitididorsis opacidorsis Nomura, 1972 | | | Serica (Serica) ovata Nomura, 1974 | | | Serica (Serica) pilosa Nomura, 1971 | | | Serica (Serica) planifrons Nomura, 1972 | | | Serica (Serica) rosinae Pic, 1904 | | | Serica rosinae kurosawai Nomura, 1959 | | | Serica rosinae rosinae | | | Serica (Serica) tokejii Nomura, 1959 | | | Serica (Serica) trichofemorata Nomura, 1959 | | | Serica (Serica) yoshidai Nomura, 1959 | | | Serica niijima Kontkanen, 1956 | | | Serica similis Lewis, 1895 | | | Serica simillima Kobayashi, 1983 | | | Serica takagii Sawada, 1937 | | | Sericania alternata Sawada, 1938 | | | Sericania angulata Lewis, 1895 | | | Sericania awana Nomura, 1976 | | | Sericania babai (Nomura, 1976) | ⇒ Sericania galloisi Niijima & Ki..., 1927 | | Sericania chikuzensis Sawada, 1938 | | | Sericania elongata Nomura, 1976 | | | Sericania fulgida (Niijima & Ki..., 1927) | ⇒ Sericania fuscolineata fulgida Niijima & Ki..., 1927 | | Sericania fulgida lineata (Sawada, 1939) | ⇒ Sericania fuscolineata fulgida Niijima & Ki..., 1927 | | Sericania fuscolineata Motschulsky, 1860 | | | Sericania fuscolineata ezoensis Nomura, 1976 | | | Sericania fuscolineata fulgida Niijima & Ki..., 1927 | | | Sericania fuscolineata fuscolineata | | | Sericania fuscolineata minuscula Nomura, 1976 | | | Sericania fuscolineata nipponensis (Nomura, 1976) | ⇒ Sericania fuscolineata Motschulsky, 1860 | | Sericania galloisi Niijima & Ki..., 1927 | | | Sericania hidana Niijima & Ki..., 1923 | | | Sericania imadatei (Sawada, 1955) | ⇒ Sericania ohtakei Sawada, 1955 | | Sericania kadowakii Nakane, 1983 | | | Sericania kirai Sawada, 1938 | | | Sericania kobayashii Nomura, 1976 | | | Sericania kompira Miyake & Sano, 1996 | | | Sericania lewisi Arrow, 1913 | | | Sericania lineata (Sawada, 1939) | ⇒ Sericania fuscolineata fulgida Niijima & Ki..., 1927 | | Sericania mara Ahrens, 2004 | | | Sericania marginata Nomura, 1973 | | | Sericania matusitai Sawada, 1955 | | | Sericania mimica Lewis, 1895 | | | Sericania mimica kompira Miyake & Sano, 1996 | | | Sericania mimica mimica | | | Sericania minuscula (Nomura, 1976) | ⇒ Sericania fuscolineata minuscula Nomura, 1976 | | Sericania miyakei Nomura, 1960 | | | Sericania ohirai Sawada, 1960 | | | Sericania ohtakei Sawada, 1955 | | | Sericania opaca Nomura, 1973 | | | Sericania pacis Ahrens, 2004 | | | Sericania quadrifoliata Lewis, 1895 | | | Sericania sachalinensis (Matsumura, 1911) | ⇒ Mesoserica sachalinensis Matsumura, 1914 | | Sericania serripes Nomura, 1973 | | | Sericania shikokuana Nakane, 1954 | | | Sericania sinuata Nomura, 1959 | | | Sericania suturalis (Nomura, 1976) | ⇒ Sericania marginata Nomura, 1973 | | Sericania tohokuensis Sawada, 1955 | | | Sericania yamauchii Sawada, 1938 | | | Sericania yamauchii mochizukii Kobayashi, 2014 | | | Sericania yamauchii yamauchii | | | Sericania yamayai Kobayashi & ..., 2008 | | | Sericania yanoi Kobayashi, 2014 | | | Setylaides foveatus Schmidt, 1909 | | | Setylaides nakanoi Ochi & Kawahara, 2019 | | | Setylaides tsujii Kakizoe, Kaw..., 2021 | | | Sigorus porcus Fabricius, 1792 | Mahogany Ridgeback scarab | | Silluvia elongata Landin, 1949 | | | Sinodiapterna troitzkyi Jacobson, 1897 | | | Smaragdesthes africana smaragdina Chevrolat, 1836 | | | Sophrops kawadai Nomura, 1959 | | | Sophrops kawadai kawadai | | | Sophrops kawadai okinawaensis Nomura, 1977 | | | Sophrops konishii Nomura, 1970 | | | Sophrops konishii konishii | | | Sophrops konishii yonaguniensis Nomura, 1970 | | | Sophrops takatoshii Itoh, 1989 | | | Stenotothorax hibernalis Nakane & Tsu..., 1956 | | | Stenotothorax hibernalis hibernalis | | | Subrinus sturmi Harold, 1870 | | | Taeniodera marmorata Wallace, 1867 | | | Teuchestes analis Fabricius, 1787 | | | Teuchestes brachysomus Solsky, 1874 | | | Torynorrhina opalina Hope, 1831 | | | Trematodes tenebrioides Pallas, 1871 | | | Trichiorhyssemus asperulus Waterhouse, 1875 | | | Trichiorhyssemus boninensis (Nakane, 1960) | ⇒ Neotrichiorhyssemus esakii Nomura, 1943 | | Trichiorhyssemus hatakei Ochi, Kawaha..., 2019 | | | Trichiorhyssemus kitayamai Ochi & Kawah..., 2001 | | | Trichiorhyssemus lasionotus Clouët, 1901 | | | Trichiorhyssemus microtrichius Ochi, Kawaha..., 2011 | | | Trichiorhyssemus nakoshii Ochi, Kawaha..., 2021 | | | Trichiorhyssemus nishinoi Ochi, Kawaha..., 2020 | | | Trichiorhyssemus yumikoae Pittino & Kawai, 2007 | | | Trichius fasciatus Linnaeus, 1758 | Bee Beetle | | Trichius japonicus Janson, 1885 | | | Trichius sexualis Bedel, 1906 | | | Trichius succinctus (Sawada, 1943) | ⇒ Trichius sexualis Bedel, 1906 | | Tricholontha papagena Nomura, 1952 | | | Tricholontha papagena inouei Kobayashi & ..., 2013 | | | Tricholontha papagena papagena | | | Trichoserica opacifrons (Nomura, 1971) | ⇒ Serica planifrons Nomura, 1972 | | Trigonophorinus riaulti Fairmaire, 1897 | | | Trigonophorus riaulti (Fairmaire, 1897) | ⇒ Trigonophorinus riaulti Fairmaire, 1897 | | Trypoxylus (Allomyrina) dichotomus (Linnaeus, 1771) | ⇒ Allomyrina dichotoma Linnaeus, 1771 | | Trypoxylus dichotomus septentrionalis Kono, 1931 | | | Trypoxylus dichotomus shizuae Adachi, 2017 | | | Trypoxylus dichotomus takarai Kusui, 1976 | | | Trypoxylus dichotomus tsuchiyai Nagai, 2006 | | | Trypoxylus dichotoma Linnaeus, 1771 | | | Valgus smithii MacLeay, 1838 | | | Xylonychus eucalypti pentaphyllus Arrow, 1924 | | | Xylotrupes gideon Linnaeus, 1767 | Brown rhinoceros beetle | | Yanovalgus planiusculus Nomura, 1952 | |
Subnodes of Scarabaeidae with occurrences in Japan | Subfamily | Aphodiinae
Leach, 1815
Aphodiine Dung Beetles, Small dung beetles | 71
199 | Genera
Species |
Taxom
| | Subfamily | Cetoniinae
Leach, 1815
Flower beetles, Flower chafers, Flower scarabs, Fruit and flower chafers | 59
107 | Genera
Species |
Taxom
| | Subfamily | Dynastinae
MacLeay, 1819
Horn beetles, Rhinoceros beetles | 9
16 | Genera
Species |
Taxom
| | Subfamily | Euchirinae
Hope, 1840
| 1
2 | Genus
Species |
Taxom
| | Subfamily | Melolonthinae
Samouelle, 1819
May Beetles and Junebugs, White grubs | 42
199 | Genera
Species |
Taxom
| | Subfamily | Rutelinae
MacLeay, 1819
Shining Flower Chafers, Shining Leaf Chafers | 15
98 | Genera
Species |
Taxom
| | Subfamily | Scarabaeinae
Latreille, 1802
Dung Beetles, True dung beetles | 11
102 | Genera
Species |
Taxom
|
 | | Scarabaeidae (Latreille 1802) |
Links, sources and further information Scarabaeidae at artfakta.se
Scarabaeidae at dyntaxa.se
Scarabaeidae at gbif.org
Scarabaeidae at irmng.org
Scarabaeidae at itis.gov
|
