Checklist for Subfamily Scarabaeinae Latreille, 1802 (Dung Beetles)
and region Sudan (North Africa)
Class
| Insecta
Linnaeus, 1758
Insects, True insects | 327
2845
6248 | Families
Genera
Species |
Taxom
|
Subclass
| Pterygota
Lang, 1888
Winged insects | 326
2842
6245 | Families
Genera
Species |
Taxom
|
Infraclass
| Neoptera
van der Wulp, 1890
Modern Wing-folding insects | 312
2802
6154 | Families
Genera
Species |
Taxom
|
Superorder
| Holometabola
Heider, 1889
Holometabolous Insects | 191
1977
4576 | Families
Genera
Species |
Taxom
|
Order
| Coleoptera
Linnaeus, 1758
Beetles, Coleopterans | 50
685
1567 | Families
Genera
Species |
Taxom
|
Suborder
| Polyphaga
Emery, 1886
Polyphags | 46
573
1196 | Families
Genera
Species |
Taxom
|
Infraorder
| Scarabaeiformia
Crowson, 1960
| 7
153
399 | Families
Genera
Species |
Taxom
|
Superfamily
| Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles | 7
153
399 | Families
Genera
Species |
Taxom
| | Family | Scarabaeidae
Latreille, 1802
Dung beetles, Flower beetles, Fruit chafers, Rain beetles | 136
362 | Genera
Species |
Taxom
| | Subfamily | Scarabaeinae
Latreille, 1802
Dung Beetles, True dung beetles | 28
121 | Genera
Species |
Taxom
|
Genera for Sudan
28 Genera with 121 Species #ACDEGHKLMOPST
| Allogymnopleurus Janssens, 1871 | | 3 | Species | Taxom | | Caccobius Thomson, 1867 | | 3 | Species | Taxom | | Catharsius Hope, 1960 | | 1 | Species | Taxom | | Cheironitis van Lansberge, 1856 | | 2 | Species | Taxom | | Cleptocaccobius Cambefort, 1902 | | 1 | Species | Taxom | | Copris Geoffroy, 0 | Mexican dung beetle | 6 | Species | Taxom | | Coptorhina Hope, 1857 | | 1 | Species | Taxom | | Diastellopalpus van Lansberge, 1891 | | 2 | Species | Taxom | | Digitonthophagus Balthasar, 2017 | | 6 | Species | Taxom | | Euonthophagus Balthasar, 1897 | | 2 | Species | Taxom | | Garreta Janssens, 1840 | | 2 | Species | Taxom | | Gymnopleurus Illiger, 1843 | | 10 | Species | Taxom | | Heliocopris Hope, 0 | | 2 | Species | Taxom | | Kheper Janssens, 1886 | | 2 | Species | Taxom | | Liatongus Reitter, 1840 | | 1 | Species | Taxom | | Metacatharsius Paulian, 1960 | | 9 | Species | Taxom | | Milichus Péringuey, 1857 | | 1 | Species | Taxom | | Onitis Fabricius, 1789 | | 5 | Species | Taxom | | Onthophagus Latreille, 1902 | | 38 | Species | Taxom | | Pedaria Laporte, 1890 | | 3 | Species | Taxom | | Phalops Erichson, 1883 | | 7 | Species | Taxom | | Proagoderus Lansberge, 1851 | | 4 | Species | Taxom | | Pseudosaproecius Balthasar, 1884 | | 2 | Species | Taxom | | Pycnopanelus Arrow, 1913 | | 1 | Species | Taxom | | Scarabaeus Linnaeus, 1789 | | 6 | Species | Taxom | | Sisyphus Latreille, 0 | | 1 | Species | Taxom | | Stiptopodius Harold, 1992 | | 2 | Species | Taxom | | Tiniocellus Péringuey, 1895 | | 1 | Species | Taxom |
Species list for Sudan
121 Species and 6 Subspecies | Allogymnopleurus anthracinus Klug, 1845 | | | Allogymnopleurus signaticollis Waterhouse, 1890 | | | Allogymnopleurus umbrinus Gerstaecker, 1871 | | | Caccobius ferrugineus Fahraeus, 1857 | | | Caccobius polygonus d´Orbigny, 1911 | | | Caccobius punctatissimus Harold, 1867 | | | Catharsius africanus Ferreira, 1960 | | | Cheironitis imitator Balthasar, 1963 | | | Cheironitis osiridis Reiche, 1856 | | | Cleptocaccobius signaticollis d´Orbigny, 1902 | | | Copris diversus Waterhouse, 1891 | | | Copris gazellarum Gillet, 1918 | | | Copris harrisi Waterhouse, 1891 | | | Copris laius Harold, 1868 | | | Copris laius angusticornis Nguyen-phung..., 1987 | | | Copris laius laius | | | Coptorhina nitidipennis Boheman, 1857 | | | Diastellopalpus thomsoni Bates, 1888 | | | Diastellopalpus tridens (Fairmaire, 1891) | ⇒ Diastellopalpus thomsoni Bates, 1888 | | Digitonthophagus aksumensis Génier, 2017 | | | Digitonthophagus eucatta Génier, 2017 | | | Digitonthophagus gazella Fabricius, 1787 | Brown dung beetle | | Digitonthophagus lusinganus Orbigny, 1905 | | | Digitonthophagus petilus Génier, 2017 | | | Digitonthophagus sahelicus Moretto, 2017 | | | Euonthophagus carbonarius Klug, 1855 | | | Euonthophagus maindroni d´Orbigny, 1897 | | | Garreta crenulatus Kolbe, 1895 | | | Garreta rutilans Castelnau, 1840 | | | Gymnopleurus aeruginosus Harold, 1867 | | | Gymnopleurus bicolor (Latreille, 1823) | ⇒ Gymnopleurus leei Fabricius, 1792 | | Gymnopleurus coerulescens Olivier, 1789 | | | Gymnopleurus fulgidus Olivier, 1789 | | | Gymnopleurus laevicollis (Castelnau, 1840) | ⇒ Gymnopleurus purpureus Garreta, 1914 | | Gymnopleurus leei Fabricius, 1792 | | | Gymnopleurus puncticollis (Gillet, 1909) | ⇒ Gymnopleurus virens Erichson, 1843 | | Gymnopleurus purpureus Garreta, 1914 | | | Gymnopleurus sericeifrons Fairmaire, 1887 | | | Gymnopleurus virens Erichson, 1843 | | | Heliocopris gigas Linnaeus, 1758 | | | Heliocopris gigas gigas | | | Kheper aegyptiorum aegyptiorum | | | Kheper vanderkelleni Lansberge, 1886 | | | Liatongus militaris Castelnau, 1840 | Yellow Shouldered Dung Beetle | | Metacatharsius abortivus Fairmaire, 1891 | | | Metacatharsius auberti Fairmaire, 1891 | | | Metacatharsius darfurensis Balthasar, 1965 | | | Metacatharsius holyi Balthasar, 1965 | | | Metacatharsius inermis Castelnau, 1840 | | | Metacatharsius minutus Lansberge, 1882 | | | Metacatharsius nubiensis Harold, 1878 | | | Metacatharsius peleus Olivier, 1789 | | | Metacatharsius renaudi Frey, 1960 | | | Milichus apicalis (Fahraeus, 1857) | ⇒ Milichus rhodesianus Péringuey, 1904 | | Onitis alexis alexis | | | Onitis guineensis Gillet, 1918 | | | Onitis laticollis Lansberge, 1875 | | | Onitis mraci Moretto, 2021 | | | Onitis unguiculatus Olivier, 1789 | | | Onthophagus (Onthophagus) margaritifer d´Orbigny, 1898 | | | Onthophagus (Strandius) obliquus Olivier, 1789 | | | Onthophagus acutus d´Orbigny, 1908 | | | Onthophagus aeneopiceus d´Orbigny, 1902 | | | Onthophagus aerestriatus d´Orbigny, 1913 | | | Onthophagus arcifer d´Orbigny, 1904 | | | Onthophagus atrovittatus d´Orbigny, 1908 | | | Onthophagus bicolor (Raffray, 1877) | ⇒ Onthophagus flavolimbatus Klug, 1855 | | Onthophagus bidens Olivier, 1789 | | | Onthophagus bituberculatus (Olivier, 1789) | ⇒ Onthophagus acutus d´Orbigny, 1908 | | Onthophagus blanchardi Harold, 1869 | | | Onthophagus cornifrons Thomson, 1858 | | | Onthophagus cupreus Harold, 1880 | | | Onthophagus filicornis Harold, 1873 | | | Onthophagus fimetarius Roth, 1851 | | | Onthophagus flavolimbatus Klug, 1855 | | | Onthophagus guttiger d´Orbigny, 1905 | | | Onthophagus heteroclitus d´Orbigny, 1911 | | | Onthophagus melanocephalus Klug, 1845 | | | Onthophagus mocquerysi d´Orbigny, 1902 | | | Onthophagus nilicola d´Orbigny, 1902 | | | Onthophagus niloticus Harold, 1879 | | | Onthophagus nitidulus Klug, 1845 | | | Onthophagus paravinctus Tagliaferri ..., 2012 | | | Onthophagus pullatus d´Orbigny, 1905 | | | Onthophagus reticulatus d´Orbigny, 1902 | | | Onthophagus scaber Roth, 1851 | | | Onthophagus sellatus Klug, 1845 | | | Onthophagus semivirescens d´Orbigny, 1902 | | | Onthophagus sericeicollis Boucomont, 1928 | | | Onthophagus subdivisus d´Orbigny, 1908 | | | Onthophagus subulifer d´Orbigny, 1908 | | | Onthophagus thomsoni Bates, 1888 | | | Onthophagus transcaspicus Koenig, 1888 | Gazella scarab | | Onthophagus trinominatus Goidanich, 1926 | | | Onthophagus variegatus (Fabricius, 1798) | ⇒ Onthophagus filicornis Harold, 1873 | | Onthophagus vinctus Erichson, 1843 | | | Onthophagus viridicatus d´Orbigny, 1902 | | | Pedaria criberrima Waterhouse, 1890 | | | Pedaria nigra Laporte, 1832 | | | Pedaria tuberculigera Waterhouse, 1890 | | | Phalops barbicornis Lansberge, 1883 | | | Phalops beccarii Harold, 1875 | | | Phalops djuricus Kuntzen, 1913 | | | Phalops fimbriatus Klug, 1835 | | | Phalops iphis Olivier, 1789 | | | Phalops princeps d´Orbigny, 1908 | | | Phalops sulcatus Lansberge, 1883 | | | Proagoderus multicornis d´Orbigny, 1908 | | | Proagoderus patinatus Marcus, 1917 | | | Proagoderus quadrispinosus d´Orbigny, 1908 | | | Proagoderus worsissa Roth, 1851 | | | Pseudosaproecius portentosus Felsche, 1907 | | | Pseudosaproecius validicornis Quedenfeldt, 1884 | | | Pycnopanelus parvicollis d´Orbigny, 1913 | | | Scarabaeus (Scarabaeus) pius Illiger, 1803 | | | Scarabaeus (Scarabaeus) sacer (Linnaeus, 1758) | ⇒ Scarabaeus pius Illiger, 1803 | | Scarabaeus goryi Castelnau, 1840 | | | Scarabaeus isidis Castelnau, 1840 | | | Scarabaeus jalof Castelnau, 1840 | | | Scarabaeus palemo Olivier, 1789 | | | Sisyphus crispatus crispatus | | | Stiptopodius doriae Harold, 1871 | | | Stiptopodius glaber Branco, 1992 | | | Tiniocellus setifer Kraatz, 1895 | |
Subnodes of Scarabaeinae with occurrences in Sudan
Links, sources and further information Scarabaeinae at artfakta.se
Scarabaeinae at dyntaxa.se
Scarabaeinae at itis.gov
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