Checklist for Subfamily Scarabaeinae Latreille, 1802 (Dung Beetles)
and region Europe (Eurasia)
Class
| Insecta
Linnaeus, 1758
Insects, True insects | 1408
35417
222091 | Families
Genera
Species |
Taxom
|
Subclass
| Pterygota
Lang, 1888
Winged insects | 1401
35364
221754 | Families
Genera
Species |
Taxom
|
Infraclass
| Neoptera
van der Wulp, 1890
Modern Wing-folding insects | 1211
34497
219210 | Families
Genera
Species |
Taxom
|
Superorder
| Holometabola
Heider, 1889
Holometabolous Insects | 647
26961
190296 | Families
Genera
Species |
Taxom
|
Order
| Coleoptera
Linnaeus, 1758
Beetles, Coleopterans | 169
9191
59961 | Families
Genera
Species |
Taxom
|
Suborder
| Polyphaga
Emery, 1886
Polyphags | 143
8084
50035 | Families
Genera
Species |
Taxom
|
Infraorder
| Scarabaeiformia
Crowson, 1960
| 14
778
3568 | Families
Genera
Species |
Taxom
|
Superfamily
| Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles | 14
778
3568 | Families
Genera
Species |
Taxom
| | Family | Scarabaeidae
Latreille, 1802
Dung beetles, Flower beetles, Fruit chafers, Rain beetles | 616
3000 | Genera
Species |
Taxom
| | Subfamily | Scarabaeinae
Latreille, 1802
Dung Beetles, True dung beetles | 99
765 | Genera
Species |
Taxom
|
Genera for Europe
99 Genera with 765 Species #ABCDEGHIKLMNOPSTUX
| Allogymnopleurus Janssens, 1855 | | 6 | Species | Taxom | | Amphistomus van Lansberge, 1974 | | 7 | Species | Taxom | Anachalcos Hope, 1837 | | 0 | Species | Taxom | | Aptenocanthon Matthews, 1936 | | 1 | Species | Taxom | | Ateuchites Meunier, 1898 | | 1 | Species | Taxom | | Ateuchus Weber, 1886 | | 4 | Species | Taxom | | Aulacopris White, 1974 | | 1 | Species | Taxom | | Bdelyropsis Pereira, Vul..., 1976 | | 1 | Species | Taxom | | Boletoscapter Matthews, 1974 | | 2 | Species | Taxom | | Bubas Mulsant, 1811 | | 3 | Species | Taxom | | Caccobius Thomson, 1767 | | 8 | Species | Taxom | | Canthidium Erichson, 1981 | | 8 | Species | Taxom | | Canthon Hoffmannsegg, 1953 | Tumblebug | 21 | Species | Taxom | Canthonella Chapin, 1930 | | 0 | Species | Taxom | | Canthonosoma MacLeay, 1868 | | 2 | Species | Taxom | | Catharsius Hope, 1857 | | 17 | Species | Taxom | | Cheironitis van Lansberge, 1789 | | 14 | Species | Taxom | | Chironitis Lansberge, 1790 | | 3 | Species | Taxom | | Circellium Latreille, 1781 | | 1 | Species | Taxom | | Cleptocaccobius Cambefort, 1857 | | 1 | Species | Taxom | | Copris Geoffroy, 1962 | Mexican dung beetle | 42 | Species | Taxom | | Coproecus Reiche, 1844 | | 1 | Species | Taxom | | Coprophanaeus Olsoufieff, 1847 | | 7 | Species | Taxom | | Coptodactyla Burmeister, 1877 | | 7 | Species | Taxom | | Coptorhina Hope, 1833 | | 2 | Species | Taxom | | Cryptocanthon Balthasar, 1973 | | 1 | Species | Taxom | | Cyptochirus Lesne, 1828 | | 1 | Species | Taxom | | Deltochilum Eschscholtz, 1848 | | 4 | Species | Taxom | | Demarziella Balthasar, 1976 | | 6 | Species | Taxom | Dendropaemon Perty, 1830 | | 0 | Species | Taxom | | Diastellopalpus van Lansberge, 1884 | | 3 | Species | Taxom | | Dichotomius Hope, 1867 | | 10 | Species | Taxom | | Digitonthophagus Balthasar, 2017 | | 9 | Species | Taxom | | Diorygopyx Matthews, 1871 | | 3 | Species | Taxom | | Disphysema Harold, 1873 | | 1 | Species | Taxom | | Drepanocerus Kirby, 1851 | | 3 | Species | Taxom | | Eodrepanus Barbero, Pal..., 1901 | | 2 | Species | Taxom | | Epactoides Olsoufieff, 1960 | | 1 | Species | Taxom | | Epirinus Reiche, 1880 | | 8 | Species | Taxom | | Euoniticellus Janssens, 1873 | | 9 | Species | Taxom | | Euonthophagus Balthasar, 1790 | | 10 | Species | Taxom | | Eurysternus Dalman, 2009 | | 8 | Species | Taxom | | Garreta Janssens, 1857 | | 4 | Species | Taxom | | Glauconia Paulian, 0 | | 2 | Species | Taxom | | Gromphas Brullé, 1830 | | 1 | Species | Taxom | | Gymnopleurus Illiger, 1843 | | 22 | Species | Taxom | | Helictopleurus Orbigny, 2000 | | 1 | Species | Taxom | | Heliocopris Hope, 1798 | | 11 | Species | Taxom | | Heteronitis Gillet, 1862 | | 1 | Species | Taxom | | Hyalonthophagus Palestrini &..., 1855 | | 1 | Species | Taxom | | Isocopris Pereira & Ma..., 1901 | | 1 | Species | Taxom | | Ixodina Roth, 1851 | | 1 | Species | Taxom | | Kheper Janssens, 1843 | | 5 | Species | Taxom | | Labroma Sharp, 1974 | | 1 | Species | Taxom | | Latodrepanus Krikken, 1857 | | 1 | Species | Taxom | | Lepanus Balthasar, 1874 | | 8 | Species | Taxom | | Liatongus Reitter, 1851 | | 2 | Species | Taxom | | Litocopris Waterhouse, 1869 | | 1 | Species | Taxom | | Lobateuchus Montreuil & al., 2010 | | 1 | Species | Taxom | | Macroderes Westwood, 1880 | | 2 | Species | Taxom | | Melanocanthon Halffter, 1920 | | 1 | Species | Taxom | | Metacatharsius Paulian, 1857 | | 10 | Species | Taxom | | Milichus Péringuey, 1871 | | 2 | Species | Taxom | | Mimonthophagus Balthasar, 1855 | | 2 | Species | Taxom | | Monoplistes Lansberge, 1888 | | 2 | Species | Taxom | | Neosisyphus Müller, 1877 | | 7 | Species | Taxom | | Odontoloma Boheman, 1868 | | 5 | Species | Taxom | | Oniticellus Serville, 1840 | | 10 | Species | Taxom | | Onitis Fabricius, 1886 | | 28 | Species | Taxom | | Ontherus Erichson, 1996 | | 5 | Species | Taxom | | Onthophagus Latreille, 1972 | | 296 | Species | Taxom | | Pachylomera Griffith & P..., 1828 | | 1 | Species | Taxom | | Pachysoma MacLeay, 1953 | | 4 | Species | Taxom | | Parachorius Harold, 2011 | | 2 | Species | Taxom | | Paragymnopleurus Shipp, 1875 | | 1 | Species | Taxom | | Peckolus Scholtz & Ho..., 1988 | | 1 | Species | Taxom | | Pedaria Laporte, 1901 | | 4 | Species | Taxom | | Phalops Erichson, 1867 | | 10 | Species | Taxom | | Phanaeus MacLeay, 1994 | Burrowing Dung Beetles | 10 | Species | Taxom | | Proagoderus Lansberge, 1901 | | 7 | Species | Taxom | | Pseudocanthon Bates, 1846 | | 3 | Species | Taxom | | Sarophorus Erichson, 1840 | | 2 | Species | Taxom | | Sauvagesinella Paulian, 2002 | | 2 | Species | Taxom | | Scarabaeus Linnaeus, 1901 | | 37 | Species | Taxom | | Sceliages Westwood, 2002 | | 1 | Species | Taxom | | Scybalocanthon Martínez, 1855 | | 2 | Species | Taxom | | Sinodrepanus Simonis, 1875 | | 1 | Species | Taxom | | Sisyphus Latreille, 1818 | | 15 | Species | Taxom | | Stiptopodius Harold, 1923 | | 1 | Species | Taxom | | Sulcophanaeus Olsoufieff, 1840 | | 1 | Species | Taxom | Sylvicanthon Halffter & M..., 1977 | | 0 | Species | Taxom | | Synapsis Forster, 1869 | | 1 | Species | Taxom | | Temnoplectron Westwood, 1841 | | 4 | Species | Taxom | | Tesserodon Hope, 1775 | | 1 | Species | Taxom | | Tibiodrepanus Krikken, 1840 | | 1 | Species | Taxom | | Tiniocellus Péringuey, 1851 | | 4 | Species | Taxom | Trichillum Harold, 1868 | | 0 | Species | Taxom | | Uroxys Westwood, 1874 | | 4 | Species | Taxom | | Xinidium Harold, 1869 | | 1 | Species | Taxom |
Species list for Europe
765 Species and 22 Subspecies | Allogymnopleurus aeneus Harold, 1875 | | | Allogymnopleurus alluaudi Garreta, 1914 | | | Allogymnopleurus consocius Péringuey, 1901 | | | Allogymnopleurus signaticollis Waterhouse, 1890 | | | Allogymnopleurus splendidus Bertoloni, 1849 | | | Allogymnopleurus thalassinus Klug, 1855 | | | Amphistomus calcaratus MacLeay, 1871 | | | Amphistomus complanatus Matthews, 1974 | | | Amphistomus primonactus Matthews, 1974 | | | Amphistomus speculifer Matthews, 1974 | | | Amphistomus squalidus MacLeay, 1887 | | | Amphistomus storeyi Matthews, 1974 | | | Amphistomus trispiculatus Matthews, 1974 | | | Aptenocanthon hopsoni Carter, 1936 | | | Ateuchites grandis ♰ Meunier, 1898 | | | Ateuchus aeneomicans Harold, 1868 | | | Ateuchus histeroides Weber, 1801 | | | Ateuchus pygidialis Harold, 1868 | | | Ateuchus rodriguezi Preudhomme d..., 1886 | | | Aulacopris maximus Matthews, 1974 | | | Bdelyropsis venezuelensis Howden, 1976 | | | Boletoscapter cornutus MacLeay, 1887 | | | Boletoscapter furcatus Matthews, 1974 | | | Bubas bison Linnaeus, 1767 | | | Bubas bubaloides Janssens, 1938 | | | Bubas bubalus Olivier, 1811 | | | Caccobius denticollis Harold, 1867 | | | Caccobius ferrugineus Fahraeus, 1857 | | | Caccobius histeroides Ménétriés, 1832 | | | Caccobius kelleri Olsoufieff, 1907 | | | Caccobius mundus Ménétriés, 1838 | | | Caccobius nigritulus Klug, 1855 | | | Caccobius obtusus Fahraeus, 1857 | | | Caccobius schreberi Linnaeus, 1767 | | | Canthidium annagabrielae Solis & Kohl..., 2004 | | | Canthidium ardens Bates, 1887 | | | Canthidium hespenheidei Howden & Young, 1981 | | | Canthidium pallidoalatum Howden & Young, 1981 | | | Canthidium perceptibile Howden & Young, 1981 | | | Canthidium smaragdinum Harold, 1867 | | | Canthidium subdopuncticolle Howden & Young, 1981 | | | Canthidium vespertinum Howden & Young, 1981 | | | Canthon (Boreocanthon) ebenus Say, 1823 | | | Canthon (Boreocanthon) mixtus Robinson, 1948 | | | Canthon (Boreocanthon) praticola Le Conte, 1859 | | | Canthon (Boreocanthon) simplex Le Conte, 1857 | | | Canthon (Canthon) chalcites Haldeman, 1843 | | | Canthon (Canthon) humectus Say, 1831 | Longhorn blenny | | Canthon (Canthon) imitator Brown, 1946 | | | Canthon (Canthon) indigaceus (LeConte, 1866) | ⇒ Canthon humectus Say, 1831 | | Canthon (Canthon) pilularius Linnaeus, 1758 | Dull tumblebug | | Canthon (Canthon) vigilans LeConte, 1858 | Kalia | | Canthon (Glaphyrocanthon) femoralis Chevrolat, 1834 | | | Canthon (Glaphyrocanthon) viridis (Palisot de B..., 1805) | ⇒ Epilissus viridis Klug, 1832 | | Canthon bicolor Castelnau, 1840 | | | Canthon deyrollei Harold, 1868 | | | Canthon dives Harold, 1868 | | | Canthon euryscelis Bates, 1887 | | | Canthon fulgidus Redtenbacher, 1867 | | | Canthon leechi Martinez, Ha..., 1964 | | | Canthon semiopacus Harold, 1868 | | | Canthon simulans Martínez, 1950 | | | Canthon staigi Pereira, 1953 | | | Canthonosoma castelnaui Harold, 1868 | | | Canthonosoma macleayi Harold, 1868 | | | Catharsius bradshawi Lansberge, 1887 | | | Catharsius brutus Harold, 1880 | | | Catharsius calaharicus Kolbe, 1893 | | | Catharsius furcillatus Guérin-Ménev..., 1847 | | | Catharsius guineensis Ferreira, 1963 | | | Catharsius harpagus Harold, 1877 | | | Catharsius heros Boheman, 1860 | | | Catharsius marcellus Kolbe, 1893 | | | Catharsius melancholicus (Boheman, 1860) | ⇒ Catharsius longiceps Gillet, 1907 | | Catharsius pandion Harold, 1877 | | | Catharsius phidias (Olivier, 1789) | ⇒ Catharsius platicornis Ferreira, 1960 | | Catharsius philus Kolbe, 1893 | | | Catharsius platicornis Ferreira, 1960 | | | Catharsius satyrus Kolbe, 1893 | | | Catharsius sesostris Waterhouse, 1888 | | | Catharsius tricornutus DeGeer, 1778 | | | Catharsius ulysses Boheman, 1857 | | | Cheironitis audens Péringuey, 1901 | | | Cheironitis furcifer Rossi, 1792 | | | Cheironitis haroldi Ballion, 1870 | | | Cheironitis hoplosternus Harold, 1868 | | | Cheironitis hungaricus Herbst, 1789 | | | Cheironitis indicus (Lansberge, 1875) | ⇒ Cheironitis arrowi Janssens, 1937 | | Cheironitis irroratus Rossi, 1790 | | | Cheironitis moeris Pallas, 1781 | | | Cheironitis pamphilus Ménétriés, 1849 | | | Cheironitis phoebus Reitter, 1892 | | | Cheironitis scabrosus Fabricius, 1776 | | | Cheironitis ungaricus Herbst, 1789 | | | Cheironitis ungaricus irroratus (Rossi, 1790) | ⇒ Cheironitis irroratus Rossi, 1790 | | Cheironitis ungaricus ungaricus Herbst, 1789 | | | Chironitis furcifer Rossi, 1792 | | | Chironitis hungaricus (Herbst, 1789) | ⇒ Cheironitis ungaricus Herbst, 1789 | | Chironitis irroratus Rossi, 1790 | | | Circellium bacchus Fabricius, 1781 | Flightless dung beetle | | Cleptocaccobius viridicollis Fahraeus, 1857 | | | Copris amyntor (Klug, 1855) | ⇒ Copris harrisi Waterhouse, 1891 | | Copris anceus Olivier, 1789 | | | Copris antares Bacescu & Ca..., 1947 | | | Copris armiger Gillet, 1910 | | | Copris caelatus Fabricius, 1794 | | | Copris capensis Waterhouse, 1891 | | | Copris cassius Péringuey, 1901 | | | Copris corniculatus (Mulsant, 1842) | ⇒ Copris lunaris Linnaeus, 1758 | | Copris cornifrons Boheman, 1860 | | | Copris corniger (Sahlberg, 1823) | ⇒ Onthophagus taurus Schreber, 1759 | | Copris costaricensis Gahan, 1894 | | | Copris denticulatus Nguyen-phung, 1988 | | | Copris druidum ♰ O. Heer, 1862 | | | Copris elphenor Klug, 1855 | | | Copris evanidus Klug, 1855 | | | Copris fallaciosus Gillet, 1907 | Kenyan dung beetle | | Copris fidius Olivier, 1789 | | | Copris gracilis Waterhouse, 1891 | | | Copris harrisi Waterhouse, 1891 | | | Copris hispanus Linnaeus, 1764 | | | Copris hispanus cavolinii Petagna, 1792 | | | Copris hispanus hispanus Linné, 1764 | | | Copris howdeni Matthews & H..., 1959 | | | Copris jacchoides Nguyen-phung..., 1987 | | | Copris lugubris Boheman, 1858 | | | Copris lunaris Linnaeus, 1758 | Horned Dung Beetle | | Copris macer Péringuey, 1901 | | | Copris mesacanthus Harold, 1878 | | | Copris misellus Péringuey, 1901 | | | Copris nepos Gillet, 1908 | | | Copris obesus Boheman, 1857 | | | Copris orion Klug, 1835 | | | Copris orphanus (Guérin-Ménev..., 1847) | ⇒ Copris evanidus Klug, 1855 | | Copris pueli Mollandin De..., 1905 | | | Copris puncticollis Boheman, 1857 | | | Copris subsidens Péringuey, 1901 | | | Copris subterranea ♰ O. Heer, 1862 | | | Copris subterraneus Heer, 1862 | | | Copris typhoeus Gerstaecker, 1884 | | | Copris umbilicatus Abeille de P..., 1901 | | | Copris urus (Boheman, 1857) | ⇒ Catharsius molossus Linnaeus, 1758 | | Copris vilhenai Ferreira, 1962 | | | Coproecus hemisphaericus Guérin-Ménev..., 1844 | | | Coprophanaeus (Coprophanaeus) pluto Harold, 1863 | | | Coprophanaeus (Megaphanaeus) bellicosus Olivier, 1789 | | | Coprophanaeus (Megaphanaeus) ensifer Germar, 1821 | | | Coprophanaeus (Megaphanaeus) lancifer Linnaeus, 1767 | | | Coprophanaeus boucardi Nevinson, 1891 | | | Coprophanaeus corythus Harold, 1863 | | | Coprophanaeus telamon Erichson, 1847 | | | Coptodactyla depressa Paulian, 1933 | | | Coptodactyla glabricollis Hope, 1842 | | | Coptodactyla meridionalis Matthews, 1976 | | | Coptodactyla monstrosa Felsche, 1909 | | | Coptodactyla nitida (Paulian, 1933) | ⇒ Coptodactyla subaenea Harold, 1877 | | Coptodactyla onitoides Gillet, 1925 | | | Coptodactyla subaenea Harold, 1877 | | | Coptorhina auspicata Péringuey, 1901 | | | Coptorhina klugi Hope, 1833 | | | Cryptocanthon nebulinus Howden, 1973 | | | Cyptochirus ambiguus (Kirby, 1828) | ⇒ Drepanocerus trogiformis Roth, 1851 | | Deltochilum gibbosum Fabricius, 1775 | Humpback Dung Beetle | | Deltochilum hypponum Buquet, 1844 | | | Deltochilum lobipes Bates, 1887 | | | Deltochilum mexicanum Burmeister, 1848 | | | Demarziella geminata MacLeay, 1871 | | | Demarziella imitatrix Balthasar, 1967 | | | Demarziella interrupta Carter, 1936 | | | Demarziella metallica Carter, 1936 | | | Demarziella mirifica Balthasar, 1961 | | | Demarziella pratensis Matthews, 1976 | | | Diastellopalpus infernalis Lansberge, 1887 | | | Diastellopalpus johnstoni Waterhouse, 1885 | | | Diastellopalpus lamellicollis Quedenfeldt, 1884 | | | Dichotomius (Luederwaldtinia) centralis Harold, 1869 | | | Dichotomius (Luederwaldtinia) schiffleri Vaz-de-Mello..., 2001 | | | Dichotomius (Luederwaldtinia) yucatanus Bates, 1887 | | | Dichotomius amicitiae Kohlmann & S..., 1997 | | | Dichotomius annae Linnaeus, 1767 | | | Dichotomius carolinus Linnaeus, 1767 | Carolina copris | | Dichotomius colonicus (Say, 1835) | ⇒ Dichotomius carolinus Linnaeus, 1767 | | Dichotomius melzeri Luederwaldt, 1922 | | | Dichotomius robustus Luederwaldt, 1935 | Apex mesh weaver | | Dichotomius satanas Harold, 1867 | | | Digitonthophagus aksumensis Génier, 2017 | | | Digitonthophagus biflagellatus Génier, 2017 | | | Digitonthophagus eucatta Génier, 2017 | | | Digitonthophagus falciger Génier, 2017 | | | Digitonthophagus fimator Génier, 2017 | | | Digitonthophagus gazella Fabricius, 1787 | Brown dung beetle | | Digitonthophagus sahelicus Moretto, 2017 | | | Digitonthophagus uks Génier, 2017 | | | Digitonthophagus ulcerosus Génier, 2017 | | | Diorygopyx asciculifer Matthews, 1974 | | | Diorygopyx simpliciclunis Matthews, 1974 | | | Diorygopyx tibialis MacLeay, 1871 | | | Disphysema candezei Harold, 1873 | | | Drepanocerus ambiguus Kirby, 1828 | | | Drepanocerus kirbyi Kirby, 1828 | | | Drepanocerus trogiformis Roth, 1851 | | | Eodrepanus coopei Barbero, Pal..., 2009 | | | Eodrepanus fastiditus Péringuey, 1901 | | | Epactoides basilewskyi Balthasar, 1960 | | | Epirinus aeneus Wiedemann, 1823 | | | Epirinus asper Péringuey, 1901 | | | Epirinus comosus Péringuey, 1901 | | | Epirinus flagellatus Fabricius, 1775 | | | Epirinus gratus Péringuey, 1901 | | | Epirinus obtusus Boheman, 1857 | | | Epirinus pseudorugosus Medina & Sch..., 2005 | | | Epirinus scrobiculatus Harold, 1880 | | | Euoniticellus africanus Harold, 1873 | Greater sandy dung beetle | | Euoniticellus fulvus Goeze, 1777 | | | Euoniticellus inaequalis Reiche, 1850 | | | Euoniticellus intermedius Reiche, 1848 | Intermediate sandy dung beetle | | Euoniticellus kawanus Janssens, 1939 | | | Euoniticellus nasicornis Reiche, 1849 | | | Euoniticellus pallens Olivier, 1789 | | | Euoniticellus pallipes Fabricius, 1781 | Southern sandy dung beetle | | Euoniticellus triangulatus Harold, 1873 | | | Euonthophagus amyntas Olivier, 1789 | | | Euonthophagus amyntas alces Fabricius, 1792 | | | Euonthophagus amyntas amyntas Olivier, 1789 | | | Euonthophagus atramentarius Ménétriés, 1832 | | | Euonthophagus carbonarius Klug, 1855 | | | Euonthophagus crocatus Mulsant & Go..., 1870 | | | Euonthophagus crocatus crocatus | | | Euonthophagus dorbignyi Olsoufieff, 1900 | | | Euonthophagus dorbignyi neutto Olsoufieff, 1900 | | | Euonthophagus gibbosus Scriba, 1790 | | | Eurysternus arnaudi Génier, 2009 | | | Eurysternus caribaeus Herbst, 1789 | | | Eurysternus cayennensis Castelnau, 1840 | | | Eurysternus hirtellus Dalman, 1824 | | | Eurysternus magnus Castelnau, 1840 | | | Eurysternus marmoreus Castelnau, 1840 | | | Eurysternus mexicanus Harold, 1869 | | | Eurysternus superbus Génier, 2009 | | | Garreta matabelensis Janssens, 1938 | | | Garreta nitens Olivier, 1789 | | | Garreta unicolor Fahraeus, 1857 | | | Garreta wahlbergi Fahraeus, 1857 | | | Glauconia kefersteini | | | Glauconia undulata | | | Gromphas aeruginosa Perty, 1830 | | | Gymnopleurus aenescens Wiedemann, 1821 | | | Gymnopleurus coriarius (Herbst, 1789) | ⇒ Gymnopleurus flagellatus Fabricius, 1787 | | Gymnopleurus cyaneus (Fabricius, 1798) | ⇒ Gymnopleurus sericeifrons Fairmaire, 1887 | | Gymnopleurus deperditus ♰ O. Heer, 1862 | | | Gymnopleurus eocaenicus ♰ F. Meunier, 1921 | | | Gymnopleurus flagellatus Fabricius, 1787 | | | Gymnopleurus flagellatus flagellatus Fabricius, 1787 | | | Gymnopleurus geoffroyi Fuessly, 1775 | | | Gymnopleurus geoffroyi geoffroyi | | | Gymnopleurus geoffroyi virescens Müller, 1902 | | | Gymnopleurus humanus MacLeay, 1821 | | | Gymnopleurus humeralis Klug, 1855 | | | Gymnopleurus ignitus Klug, 1855 | | | Gymnopleurus leei Fabricius, 1792 | | | Gymnopleurus mopsus Pallas, 1781 | | | Gymnopleurus mopsus mopsus | | | Gymnopleurus rotundatus ♰ O. Heer, 1862 | | | Gymnopleurus sericeifrons Fairmaire, 1887 | | | Gymnopleurus sisyphus ♰ O. Heer, 1847 | | | Gymnopleurus sturmi (MacLeay, 1821) | ⇒ Gymnopleurus sturmii MacLeay, 1821 | | Gymnopleurus sturmii MacLeay, 1821 | | | Gymnopleurus virens Erichson, 1843 | | | Helictopleurus neoamplicollis Krell, 2000 | | | Heliocopris andersoni Bates, 1868 | | | Heliocopris atropos Boheman, 1860 | | | Heliocopris erycoides Felsche, 1907 | | | Heliocopris faunus Boheman, 1857 | | | Heliocopris hamadryas Fabricius, 1775 | | | Heliocopris hermes Gillet, 1911 | | | Heliocopris japetus Klug, 1855 | | | Heliocopris marshalli Péringuey, 1901 | | | Heliocopris neptunus Boheman, 1857 | | | Heliocopris pauliani Janssens, 1939 | | | Heliocopris pirmal Fabricius, 1798 | | | Heteronitis castelnaui Harold, 1862 | | | Hyalonthophagus alcyon Klug, 1855 | | | Isocopris imitator Felsche, 1901 | | | Ixodina abyssinica Roth, 1851 | | | Kheper aegyptiorum Latreille, 1823 | | | Kheper bonellii MacLeay, 1821 | | | Kheper lamarcki MacLeay, 1821 | | | Kheper nigroaeneus Boheman, 1857 | | | Kheper prodigiosus Erichson, 1843 | | | Labroma umbratilis Matthews, 1974 | | | Latodrepanus laticollis Fahraeus, 1857 | | | Lepanus australis Matthews, 1974 | | | Lepanus globulus MacLeay, 1887 | | | Lepanus latheticus Matthews, 1974 | | | Lepanus monteithi Matthews, 1974 | | | Lepanus occidentalis Matthews, 1974 | | | Lepanus pygmaeus MacLeay, 1888 | | | Lepanus tozerensis Gunter & Weir, 2019 | | | Lepanus ustulatus Lansberge, 1874 | | | Liatongus militaris Castelnau, 1840 | Yellow Shouldered Dung Beetle | | Liatongus spathulatus Roth, 1851 | | | Litocopris simplex Harold, 1869 | | | Lobateuchus parisii ♰ O. Montreuil, 2010 | | | Macroderes nitidus Harold, 1877 | | | Macroderes politulus (Preudhomme d..., 1880) | ⇒ Macroderes bias Olivier, 1789 | | Melanocanthon granulifer Schmidt, 1920 | Pearl blenny | | Metacatharsius dentinum Ferreira, 1964 | | | Metacatharsius exiguiformis Ferreira, 1964 | | | Metacatharsius exiguus Boheman, 1860 | | | Metacatharsius ferreirae Balthasar, 1965 | | | Metacatharsius marani Balthasar, 1940 | | | Metacatharsius opacus Waterhouse, 1891 | | | Metacatharsius pseudoopacus Ferreira, 1965 | | | Metacatharsius pumilionis Wallengren, 1881 | | | Metacatharsius pusio Kolbe, 1908 | | | Metacatharsius troglodytes (Boheman, 1857) | ⇒ Metacatharsius anderseni Waterhouse, 1891 | | Milichus apicalis (Fahraeus, 1857) | ⇒ Milichus rhodesianus Péringuey, 1904 | | Milichus picticollis Gerstaecker, 1871 | | | Mimonthophagus ambiguus Péringuey, 1901 | | | Mimonthophagus anomalus Klug, 1855 | | | Monoplistes leai Paulian, 1934 | | | Monoplistes occidentalis MacLeay, 1888 | | | Neosisyphus barbarossa Wiedemann, 1823 | | | Neosisyphus calcaratus Klug, 1855 | | | Neosisyphus fortuitus Péringuey, 1901 | | | Neosisyphus kuehni Haaf, 1955 | | | Neosisyphus rubrus Paschalidis, 1974 | | | Neosisyphus spinipes Thunberg, 1818 | Grey dungball roller | | Neosisyphus tibialis Raffray, 1877 | | | Odontoloma dentinum Harold, 1868 | | | Odontoloma endroedyi Howden & Sch..., 1987 | | | Odontoloma louwi Howden & Sch..., 1987 | | | Odontoloma pusillum Howden & Sch..., 1987 | | | Odontoloma sculpturatum Harold, 1868 | | | Oniticellus (Paroniticellus) festivus Steven, 1809 | | | Oniticellus amplicollis ♰ (O. Heer, 1862) | ⇒ Helictopleurus neoamplicollis Krell, 2000 | | Oniticellus egregius Klug, 1855 | | | Oniticellus falsus (Sharp, 1875) | ⇒ Sinodrepanus falsus Sharp, 1875 | | Oniticellus formosus (Chevrolat, 1830) | ⇒ Oniticellus pictus Hausmann, 1807 | | Oniticellus fulvus (Goeze, 1777) | ⇒ Euoniticellus fulvus Goeze, 1777 | | Oniticellus pallipes (Fairmaire, 1887) | ⇒ Euoniticellus inaequalis Reiche, 1850 | | Oniticellus pictus Hausmann, 1807 | | | Oniticellus planatus Castelnau, 1840 | | | Onitis aeruginosus Klug, 1855 | | | Onitis alexis (Klug, 1835) | ⇒ Cheironitis ungaricus ungaricus Herbst, 1789 | | Onitis alexis septentrionalis Balthasar, 1942 | | | Onitis anthracinus Felsche, 1907 | | | Onitis aygulus Fabricius, 1781 | | | Onitis belial Fabricius, 1798 | | | Onitis caffer Boheman, 1857 | | | Onitis confusus Boheman, 1860 | | | Onitis cryptodus Gillet, 1918 | | | Onitis curvipes Lansberge, 1875 | | | Onitis damoetas Steven, 1806 | | | Onitis deceptor Péringuey, 1901 | | | Onitis fulgidus (Klug, 1855) | ⇒ Pleuronitis fulgidus Castelnau, 1840 | | Onitis humerosus Pallas, 1771 | | | Onitis ion Olivier, 1789 | | | Onitis magus ♰ C. von Heyden, 1862 | | | Onitis mendax Gillet, 1918 | | | Onitis minutus Lansberge, 1875 | | | Onitis obscurus Lansberge, 1886 | | | Onitis pecuarius Lansberge, 1875 | | | Onitis picticollis Boheman, 1857 | | | Onitis pseudosetosus Ferreira, 1976 | | | Onitis reichei Lansberge, 1875 | | | Onitis tortuosus Houston, 1983 | | | Onitis uncinatus Klug, 1855 | | | Onitis vanderkelleni Lansberge, 1886 | Dung beetle | | Onitis viridulus (Boheman, 1857) | ⇒ Onitis pecuarius Lansberge, 1875 | | Onitis westermanni Lansberge, 1886 | | | Ontherus brevicollis Kirsch, 1870 | | | Ontherus carinifrons Luederwaldt, 1930 | | | Ontherus pseudodidymus Génier, 1996 | | | Ontherus pubens Génier, 1996 | | | Ontherus sanctaemartae Génier, 1996 | | | Onthophagus (Afrostrandius) plebejus Klug, 1855 | | | Onthophagus (Amphionthophagus) melitaeus Fabricius, 1798 | | | Onthophagus (Euonthophagus) amyntas (Olivier, 1789) | ⇒ Euonthophagus amyntas Olivier, 1789 | | Onthophagus (Furconthophagus) furcatus (Fabricius, 1781) | ⇒ Onthophagus furcaticeps Masters, 1886 | | Onthophagus (Micronthophagus) quaestus Sharp, 1875 | | | Onthophagus (Onthophagus) batesi Howden and C..., 1963 | | | Onthophagus (Onthophagus) brevifrons (Horn, 1881) | ⇒ Onthophagus abyssinicus Gillet, 1925 | | Onthophagus (Onthophagus) depressus Harold, 1871 | Flat African dung beetle | | Onthophagus (Onthophagus) gibbulus Pallas, 1781 | | | Onthophagus (Onthophagus) hecate Panzer, 1794 | Scooped Scarab | | Onthophagus (Onthophagus) illyricus Scopoli, 1763 | | | Onthophagus (Onthophagus) incensus Say, 1835 | Dung beetle | | Onthophagus (Onthophagus) knausi Brown, 1927 | | | Onthophagus (Onthophagus) landolti (Harold, 1880) | ⇒ Onthophagus schaefferi Howden and C..., 1963 | | Onthophagus (Onthophagus) medorensis Brown, 1929 | | | Onthophagus (Onthophagus) nuchicornis Linnaeus, 1758 | | | Onthophagus (Onthophagus) oklahomensis Brown, 1927 | Oklahoma dung beetle | | Onthophagus (Onthophagus) orpheus Panzer, 1794 | | | Onthophagus (Onthophagus) pennsylvanicus Harold, 1871 | | | Onthophagus (Onthophagus) polyphemi Hubbard, 1894 | Gopher Tortoise Onthophagus Beetle | | Onthophagus (Onthophagus) striatulus Palisot de B..., 1809 | Surf blenny | | Onthophagus (Onthophagus) subaeneus (Palisot de B..., 1811) | ⇒ Onthophagus striatulus Palisot de B..., 1809 | | Onthophagus (Onthophagus) subtropicus Howden and C..., 1963 | | | Onthophagus (Onthophagus) taurus Schreber, 1759 | Bull Headed Dung Beetle | | Onthophagus (Onthophagus) tuberculifrons Harold, 1871 | Dung beetle | | Onthophagus (Onthophagus) velutinus Horn, 1875 | | | Onthophagus (Onthophagus) vitulus Péringuey, 1901 | | | Onthophagus (Palaeonthophagus) albarracinus Baraud, 1979 | | | Onthophagus (Palaeonthophagus) angorensis Petrovitz, 1963 | | | Onthophagus (Palaeonthophagus) baraudi Nicolas, 1964 | | | Onthophagus (Palaeonthophagus) coenobita Herbst, 1783 | | | Onthophagus (Palaeonthophagus) cruciatus (Ménétriés, 1832) | ⇒ Onthophagus lemur Fabricius, 1781 | | Onthophagus (Palaeonthophagus) dellacasai Pittino & Ma..., 1981 | | | Onthophagus (Palaeonthophagus) eulaminicornis Pittino, 2006 | | | Onthophagus (Palaeonthophagus) excisus Reiche & Saulcy, 1856 | | | Onthophagus (Palaeonthophagus) fissicornis Steven, 1809 | | | Onthophagus (Palaeonthophagus) fortigibber Reitter, 1909 | | | Onthophagus (Palaeonthophagus) fracticornis (Preyssler, 1790) | ⇒ Onthophagus similis Scriba, 1790 | | Onthophagus (Palaeonthophagus) gibbulus Pallas, 1781 | | | Onthophagus gibbulus gibbulus | | | Onthophagus (Palaeonthophagus) grossepunctatus Reitter, 1905 | | | Onthophagus (Palaeonthophagus) joannae Goljan, 1953 | | | Onthophagus (Palaeonthophagus) kindermanni Harold, 1877 | | | Onthophagus (Palaeonthophagus) kolenatii Reitter, 1893 | | | Onthophagus (Palaeonthophagus) latigena d´Orbigny, 1897 | | | Onthophagus (Palaeonthophagus) lemur Fabricius, 1781 | | | Onthophagus (Palaeonthophagus) leucostigma Steven, 1811 | | | Onthophagus (Palaeonthophagus) lucidus Illiger, 1800 | | | Onthophagus (Palaeonthophagus) macedonicus Miksic, 1959 | | | Onthophagus (Palaeonthophagus) marginalis Gebler, 1817 | | | Onthophagus marginalis marginalis | | | Onthophagus (Palaeonthophagus) massai Baraud, 1975 | | | Onthophagus (Palaeonthophagus) medius (Kugelann, 1792) | ⇒ Onthophagus vacca Linnaeus, 1767 | | Onthophagus (Palaeonthophagus) merdarius Chevrolat, 1865 | | | Onthophagus (Palaeonthophagus) nuchicornis Linnaeus, 1758 | Mottled dung beetle | | Onthophagus (Palaeonthophagus) opacicollis Reitter, 1893 | | | Onthophagus (Palaeonthophagus) ovatus Linnaeus, 1767 | | | Onthophagus (Palaeonthophagus) panici Petrovitz, 1964 | | | Onthophagus (Palaeonthophagus) parmatus Reitter, 1892 | | | Onthophagus (Palaeonthophagus) ponticus Harold, 1883 | | | Onthophagus (Palaeonthophagus) rodentium Pittino, 2004 | | | Onthophagus (Palaeonthophagus) ruficapillus Brullé, 1832 | | | Onthophagus ruficapillus ruficapillus Brullé, 1832 | | | Onthophagus (Palaeonthophagus) semicornis Panzer, 1798 | | | Onthophagus (Palaeonthophagus) sericatus Reitter, 1893 | | | Onthophagus (Palaeonthophagus) similis (Scriba, 1790) | ⇒ Onthophagus fracticornis Preyssler, 1790 | | Onthophagus (Palaeonthophagus) stylocerus Graells, 1851 | | | Onthophagus (Palaeonthophagus) suturellus Brullé, 1832 | | | Onthophagus (Palaeonthophagus) tesquorum Semenov & Me..., 1927 | | | Onthophagus (Palaeonthophagus) trigibber Reitter, 1893 | | | Onthophagus (Palaeonthophagus) truchmenus Kolenati, 1846 | | | Onthophagus truchmenus truchmenus | | | Onthophagus (Palaeonthophagus) vacca Linnaeus, 1767 | | | Onthophagus (Palaeonthophagus) verticicornis (Laicharting, 1781) | ⇒ Onthophagus nuchicornis Linnaeus, 1758 | | Onthophagus (Palaeonthophagus) vitulus (Fabricius, 1776) | ⇒ Onthophagus immundus Boheman, 1858 | | Onthophagus (Paleonthophagus) andalusicus Waltl, 1835 | | | Onthophagus andalusicus andalusicus | | | Onthophagus (Paleonthophagus) baraudi Nicolas, 1964 | | | Onthophagus (Paleonthophagus) coenobita Herbst, 1783 | | | Onthophagus (Paleonthophagus) fracticornis Preyssler, 1790 | | | Onthophagus (Paleonthophagus) gibbulus Pallas, 1781 | | | Onthophagus (Paleonthophagus) grossepunctatus Reitter, 1905 | | | Onthophagus (Paleonthophagus) joannae Goljan, 1953 | | | Onthophagus (Paleonthophagus) lemur Fabricius, 1781 | | | Onthophagus (Paleonthophagus) massai Baraud, 1975 | | | Onthophagus (Paleonthophagus) nuchicornis Linnaeus, 1758 | | | Onthophagus (Paleonthophagus) opacicollis Pierotti, 1959 | | | Onthophagus (Paleonthophagus) ovatus Linnaeus, 1767 | | | Onthophagus (Paleonthophagus) ruficapillus Brullé, 1832 | | | Onthophagus (Paleonthophagus) semicornis Panzer, 1798 | | | Onthophagus (Paleonthophagus) similis Scriba, 1790 | | | Onthophagus (Paleonthophagus) vacca Linnaeus, 1767 | | | Onthophagus (Paleonthophagus) verticicornis Laicharting, 1781 | | | Onthophagus (Paleonthophagus) vitulus Fabricius, 1776 | | | Onthophagus (Parentius) emarginatus Mulsant, 1842 | | | Onthophagus (Parentius) hispanicus Baraud, 1963 | | | Onthophagus (Parentius) punctatus Illiger, 1803 | | | Onthophagus brucei cyanopterus (d´Orbigny, 1905) | ⇒ Proagoderus cyanopterus d´Orbigny, 1905 | | Onthophagus (Proagoderus) pegesimallus Marcus, 1917 | | | Onthophagus (Relictonthophagus) emarginatus (Mulsant, 1842) | ⇒ Onthophagus rubrimaculatus MacLeay, 1864 | | Onthophagus (Relictonthophagus) nigellus (Illiger, 1803) | ⇒ Caccobius histeroides Ménétriés, 1832 | | Onthophagus (Relictonthophagus) punctatus (Illiger, 1803) | ⇒ Onthophagus emarginatus Mulsant, 1842 | | Onthophagus (Trichonthophagus) hirtus Illiger, 1803 | | | Onthophagus (Trichonthophagus) maki Illiger, 1803 | | | Onthophagus absyrtus Balthasar, 1946 | | | Onthophagus aeruginosus Roth, 1851 | | | Onthophagus albicornis Palisot de B..., 1805 | | | Onthophagus albipennis Péringuey, 1908 | | | Onthophagus amphicoma Boucomont, 1914 | | | Onthophagus anchommatus Lea, 1923 | | | Onthophagus anisocerus Erichson, 1842 | | | Onthophagus arvernus ♰ L. Piton, 1939 | | | Onthophagus asper MacLeay, 1864 | | | Onthophagus atrox Harold, 1867 | | | Onthophagus auritus Erichson, 1842 | | | Onthophagus australis Guérin-Ménev..., 1830 | Southern dung beetle | | Onthophagus axillaris (Boheman, 1860) | ⇒ Onthophagus criniger d´Orbigny, 1904 | | Onthophagus beiranus Péringuey, 1908 | | | Onthophagus bicarinaticeps Lea, 1923 | | | Onthophagus binodis Thunberg, 1818 | Humpbacked Dung Beetle | | Onthophagus bisontinus ♰ Heer, 1862 | | | Onthophagus blackwoodensis Blackburn, 1891 | | | Onthophagus bornemisszai Matthews, 1972 | | | Onthophagus brooksi Matthews, 1972 | | | Onthophagus bunamin Matthews, 1972 | | | Onthophagus camelus Fabricius, 1787 | | | Onthophagus capella (Kirby, 1818) | ⇒ Onthophagus ramosellus Bates, 1891 | | Onthophagus capelliformis Gillet, 1925 | | | Onthophagus capellinus Frey, 1963 | | | Onthophagus capitosus Harold, 1867 | | | Onthophagus carpophilus Pereira & Ha..., 1961 | | | Onthophagus championi Bates, 1887 | | | Onthophagus chepara Matthews, 1972 | | | Onthophagus chryses Bates, 1887 | | | Onthophagus confluens (d´Orbigny, 1902) | ⇒ Onthophagus vacca Linnaeus, 1767 | | Onthophagus consentaneus Harold, 1867 | Common northern dung beetle | | Onthophagus conspicuus MacLeay, 1864 | | | Onthophagus crassus ♰ (Heer, 1862) | ⇒ Parascatonomus tricornis Wiedemann, 1823 | | Onthophagus cuniculus MacLeay, 1864 | | | Onthophagus cyanochlorus d´Orbigny, 1902 | | | Onthophagus dandalu Matthews, 1972 | | | Onthophagus declivis Harold, 1869 | | | Onthophagus desectus MacLeay, 1871 | | | Onthophagus dinoderus d´Orbigny, 1913 | | | Onthophagus discolor (Hope, 1841) | ⇒ Onthophagus impurus Harold, 1868 | | Onthophagus dunningi Harold, 1869 | | | Onthophagus ebenus Péringuey, 1888 | | | Onthophagus emeritus Péringuey, 1901 | | | Onthophagus evanidus Harold, 1869 | | | Onthophagus fabricii Waterhouse, 1894 | | | Onthophagus ferox Harold, 1867 | Western dung beetle | | Onthophagus ferrari Matthews, 1972 | | | Onthophagus filicornis Harold, 1873 | | | Onthophagus fimetarius Roth, 1851 | | | Onthophagus fissiceps MacLeay, 1888 | | | Onthophagus fissinasus Fairmaire, 1895 | | | Onthophagus flavoapicalis Lea, 1923 | | | Onthophagus flavolimbatus Klug, 1855 | | | Onthophagus fletcheri Blackburn, 1903 | | | Onthophagus frenchi Blackburn, 1903 | | | Onthophagus froggattellus Monteith & R..., 2017 | | | Onthophagus fugitivus Péringuey, 1901 | | | Onthophagus fuliginosus Erichson, 1842 | | | Onthophagus furcaticeps Masters, 1886 | | | Onthophagus furcatoides (Lansberge, 1886) | ⇒ Onthophagus neofurcatus Goidanich, 1926 | | Onthophagus gangulu Matthews, 1972 | | | Onthophagus geelongensis Blackburn, 1891 | | | Onthophagus giraffa Hausmann, 1807 | | | Onthophagus giuseppecarpanetoi Tagliaferri ..., 2012 | | | Onthophagus glabratus Hope, 1842 | | | Onthophagus granulatus (Boheman, 1858) | ⇒ Onthophagus consentaneus Harold, 1867 | | Onthophagus granulifer (Harold, 1886) | ⇒ Onthophagus ponticus Harold, 1883 | | Onthophagus guatemalensis (Bates, 1887) | ⇒ Onthophagus medorensis Brown, 1929 | | Onthophagus gulmarri Matthews, 1972 | | | Onthophagus haagi Harold, 1867 | | | Onthophagus herus Péringuey, 1901 | | | Onthophagus hirtellus Frey, 1957 | | | Onthophagus hoplocerus Lea, 1923 | | | Onthophagus hyaena Fabricius, 1801 | | | Onthophagus immundus Boheman, 1858 | | | Onthophagus incanus MacLeay, 1888 | | | Onthophagus incornutus MacLeay, 1871 | | | Onthophagus indutus d´Orbigny, 1902 | | | Onthophagus infuscatus Klug, 1845 | | | Onthophagus interstitialis (Fahraeus, 1857) | ⇒ Euonthophagus carbonarius Klug, 1855 | | Onthophagus jangga Matthews, 1972 | | | Onthophagus jubatus Harold, 1869 | | | Onthophagus juvencus Klug, 1835 | | | Onthophagus kiambram Storey, 1977 | | | Onthophagus koebelei Blackburn, 1903 | | | Onthophagus kokereka Matthews, 1972 | | | Onthophagus lacustris Harold, 1877 | | | Onthophagus lamelliger (Gerstaecker, 1871) | ⇒ Onthophagus ochropygus d´Orbigny, 1902 | | Onthophagus laminatus (MacLeay, 1864) | ⇒ Onthophagus lunatus Harold, 1868 | | Onthophagus latro Harold, 1877 | | | Onthophagus leai Blackburn, 1895 | | | Onthophagus lebasi Boucomont, 1932 | | | Onthophagus longipes Paulian, 1937 | | | Onthophagus lugubris (Fahraeus, 1857) | ⇒ Onthophagus lituratus Roth, 1851 | | Onthophagus lunatus Harold, 1868 | | | Onthophagus luteus ♰ Oustalet, 1874 | | | Onthophagus macleayi Blackburn, 1903 | | | Onthophagus macrocephalus Kirby, 1818 | | | Onthophagus malthinus Gillet, 1930 | | | Onthophagus mamillatus Lea, 1923 | | | Onthophagus manya Matthews, 1972 | | | Onthophagus marginicollis Harold, 1880 | | | Onthophagus mexicanus Bates, 1887 | | | Onthophagus minutus (Hausmann, 1807) | ⇒ Onthophagus centricornis Fabricius, 1798 | | Onthophagus mjobergi Gillet, 1925 | | | Onthophagus mniszechi Harold, 1869 | Kershaw´s burying beetle | | Onthophagus mulgravei Paulian, 1937 | | | Onthophagus mutatus Harold, 1859 | | | Onthophagus muticus MacLeay, 1864 | | | Onthophagus neostenocerus Goidanich, 1926 | | | Onthophagus nodulifer Harold, 1867 | | | Onthophagus nutans (Roth, 1851) | ⇒ Onthophagus schaufussi Harold, 1867 | | Onthophagus obtusicornis (Fahraeus, 1857) | ⇒ Onthophagus marginalis przewalski Kabakov, 2006 | | Onthophagus obtutus Péringuey, 1901 | | | Onthophagus ocelliger Harold, 1877 | | | Onthophagus orphnoides (Bates, 1887) | ⇒ Onthophagus mirabilis Bates, 1887 | | Onthophagus ovatulus ♰ Heer, 1847 | | | Onthophagus pallidipennis (Fahraeus, 1857) | ⇒ Onthophagus rasipennis d´Orbigny, 1908 | | Onthophagus paluma Matthews, 1972 | | | Onthophagus parumnotatus Fahraeus, 1857 | | | Onthophagus parvus Blanchard, 1853 | | | Onthophagus pentacanthus Harold, 1867 | Five horned dung beetle | | Onthophagus peramelinus Lea, 1923 | | | Onthophagus perpilosus MacLeay, 1871 | | | Onthophagus pexatus Harold, 1869 | | | Onthophagus pilosus (Fahraeus, 1857) | ⇒ Onthophagus echinus Boucomont, 1914 | | Onthophagus posticus Erichson, 1842 | | | Onthophagus praecellens Bates, 1887 | | | Onthophagus prehensilis Arrow, 1920 | | | Onthophagus probus Péringuey, 1901 | | | Onthophagus prodromus ♰ Heer, 1862 | | | Onthophagus pronus Erichson, 1842 | | | Onthophagus propinquus (MacLeay, 1888) | ⇒ Onthophagus vacca Linnaeus, 1767 | | Onthophagus pugionatus Fahraeus, 1857 | | | Onthophagus pugnacior Blackburn, 1903 | | | Onthophagus pugnax Harold, 1868 | | | Onthophagus purpureicollis MacLeay, 1864 | | | Onthophagus quadrinodosus Fahraeus, 1857 | | | Onthophagus quadripustulatus Fabricius, 1775 | | | Onthophagus queenslandicus Blackburn, 1903 | | | Onthophagus rhodesianus Frey, 1975 | | | Onthophagus rhyncophorus | | | Onthophagus rubicundulus MacLeay, 1871 | | | Onthophagus rubrimaculatus MacLeay, 1864 | | | Onthophagus rupicapra Waterhouse, 1894 | | | Onthophagus schatzmayri (Pierotti, 1959) | ⇒ Onthophagus opacicollis Reitter, 1893 | | Onthophagus schaufussi Harold, 1867 | | | Onthophagus sellatus Klug, 1845 | | | Onthophagus semiflavus (Boheman, 1860) | ⇒ Onthophagus flavolimbatus Klug, 1855 | | Onthophagus semimetallicus Lea, 1923 | | | Onthophagus sloanei Blackburn, 1903 | | | Onthophagus spitsbergeniensis Krell, 2010 | | | Onthophagus spitsbergensis ♰ O. Heer, 1870 | | | Onthophagus statzi ♰ Krell, 1990 | | | Onthophagus stellio Erichson, 1843 | | | Onthophagus sticticus Harold, 1867 | | | Onthophagus subocelliger (Blackburn, 1903) | ⇒ Onthophagus waterhousei Gillet, 1927 | | Onthophagus suffusus Klug, 1855 | | | Onthophagus sugillatus Klug, 1855 | | | Onthophagus suturalis (Péringuey, 1888) | ⇒ Onthophagus trituber Wiedemann, 1823 | | Onthophagus sydneyensis Blackburn, 1903 | | | Onthophagus tabellicornis MacLeay, 1864 | | | Onthophagus tamworthi Blackburn, 1903 | | | Onthophagus tenebrosus Harold, 1871 | | | Onthophagus thoreyi Harold, 1868 | | | Onthophagus transcaspicus Koenig, 1888 | Gazella scarab | | Onthophagus tricavicollis Lea, 1923 | | | Onthophagus trinodosus Fahraeus, 1857 | | | Onthophagus tuckonie Matthews, 1972 | | | Onthophagus tweedensis Blackburn, 1903 | | | Onthophagus urus (Harold, 1867) | ⇒ Onthophagus urusheeri Krell, 2000 | | Onthophagus urusheeri ♰ Krell, 2000 | | | Onthophagus vermiculatus Frey, 1970 | | | Onthophagus verticalis (Fahraeus, 1857) | ⇒ Onthophagus praedatus Harold, 1862 | | Onthophagus vigens Péringuey, 1901 | | | Onthophagus vilis Harold, 1877 | | | Onthophagus vinctus Erichson, 1843 | | | Onthophagus wakelbura Matthews, 1972 | | | Onthophagus walteri Josso & Prév..., 2000 | | | Onthophagus waminda Matthews, 1972 | | | Onthophagus weisei (Reitter, 1891) | ⇒ Euonthophagus gibbosus Scriba, 1790 | | Onthophagus xanthochlorus Walter & Cam..., 1977 | | | Onthophagus yaran Storey & Weir, 1990 | | | Onthophagus yeyeko Matthews, 1972 | | | Onthophagus yungaburra Matthews, 1972 | | | Pachylomera femoralis Kirby, 1828 | | | Pachysoma aesculapius Olivier, 1789 | | | Pachysoma bennigseni Felsche, 1907 | | | Pachysoma denticolle Péringuey, 1888 | | | Pachysoma gariepinus Ferreira, 1953 | | | Parachorius javanus Boucomont, 1914 | | | Parachorius semsanganus Tarasov & Keith, 2011 | | | Paragymnopleurus sinuatus abax Sharp, 1875 | | | Peckolus alpinus Howden & Sch..., 1988 | | | Pedaria barrei Josso & Prév..., 2006 | | | Pedaria cylindrica Fahraeus, 1857 | | | Pedaria picea Fahraeus, 1857 | | | Pedaria segregis Péringuey, 1901 | | | Phalops ardea Klug, 1855 | | | Phalops boschas Klug, 1855 | | | Phalops bubalus Harold, 1867 | | | Phalops dregei Harold, 1867 | | | Phalops flavocinctus Klug, 1855 | | | Phalops plancus Erichson, 1843 | | | Phalops prasinus (Erichson, 1843) | ⇒ Phalops beccarii Harold, 1875 | | Phalops rufosignatus (Lansberge, 1885) | ⇒ Phalops dregei Harold, 1867 | | Phalops vanellus Lansberge, 1883 | | | Phalops wittei Harold, 1867 | | | Phanaeus (Notiophanaeus) meleagris Blanchard, 1846 | | | Phanaeus (Notiophanaeus) pyrois Bates, 1887 | | | Phanaeus (Phanaeus) amithaon Harold, 1875 | | | Phanaeus (Phanaeus) palliatus Sturm, 1843 | | | Phanaeus (Phanaeus) vindex MacLeay, 1819 | Laguna Madre fiddler | | Phanaeus daphnis Harold, 1863 | Beetle | | Phanaeus demon Castelnau, 1840 | | | Phanaeus menelas (Castelnau, 1840) | ⇒ Sulcophanaeus menelas Castelnau, 1840 | | Phanaeus mexicanus Harold, 1863 | | | Phanaeus texensis Edmonds, 1994 | | | Proagoderus aciculatus Fahraeus, 1857 | | | Proagoderus bicallosus Klug, 1855 | | | Proagoderus lanista Castelnau, 1840 | | | Proagoderus loricatus Klug, 1855 | | | Proagoderus rangifer Klug, 1855 | | | Proagoderus rectefurcatus Fairmaire, 1891 | | | Proagoderus sapphirinus (Péringuey, 1901) | ⇒ Onthophagus aureiceps d´Orbigny, 1902 | | Pseudocanthon perplexus LeConte, 1847 | | | Pseudocanthon vitraci Fleutiaux & ..., 1889 | | | Pseudocanthon xanthurus Blanchard, 1846 | | | Sarophorus costatus Fahraeus, 1857 | | | Sarophorus tuberculatus Castelnau, 1840 | | | Sauvagesinella becki Paulian, 1934 | | | Sauvagesinella loftyensis Matthews & Weir, 2002 | | | Scarabaeus (Ateuchetus) armeniacus Ménétriés, 1832 | | | Scarabaeus (Ateuchetus) cicatricosus (Lucas, 1846) | ⇒ Scarabaeus palemo Olivier, 1789 | | Scarabaeus (Ateuchetus) laticollis Linnaeus, 1767 | | | Scarabaeus (Ateuchetus) puncticollis (Latreille, 1819) | ⇒ Geotrupes spiniger Marsham, 1802 | | Scarabaeus (Ateuchetus) semipunctatus Fabricius, 1792 | | | Scarabaeus (Ateuchetus) variolosus (Fabricius, 1787) | ⇒ Ateuchetus semipunctatus Fabricius, 1792 | | Scarabaeus (Scarabaeus) babori Balthasar, 1934 | | | Scarabaeus (Scarabaeus) bannuensis Janssens, 1940 | | | Scarabaeus (Scarabaeus) carinatus (Gebler, 1841) | ⇒ Calcodes carinatus Linnaeus, 1758 | | Scarabaeus (Scarabaeus) pius Illiger, 1803 | | | Scarabaeus (Scarabaeus) sacer (Linnaeus, 1758) | ⇒ Scarabaeus pius Illiger, 1803 | | Scarabaeus (Scarabaeus) typhon (Fischer von ..., 1823) | ⇒ Megasoma gyas Herbst, 1785 | | Scarabaeus (Scarabaeus) winkleri Stolfa, 1938 | | | Scarabaeus aegaeon Drury, 1773 | | | Scarabaeus aegyptiacus Stolfa, 1938 | | | Scarabaeus affinis (Sturm, 1800) | ⇒ Onthophagus vacca Linnaeus, 1767 | | Scarabaeus ambiguus (Boheman, 1857) | ⇒ Gnorimus variabilis Linnaeus, 1758 | | Scarabaeus bellicosus (Olivier, 1789) | ⇒ Coprophanaeus bellicosus Olivier, 1789 | | Scarabaeus caffer Boheman, 1857 | | | Scarabaeus cineta Drury, 1782 | | | Scarabaeus convexus Hausmann, 1807 | | | Scarabaeus deludens zur Strassen, 1961 | | | Scarabaeus galenus Westwood, 1847 | | | Scarabaeus goryi Castelnau, 1840 | | | Scarabaeus indus (Linnaeus, 1758) | ⇒ Euphoria inda Linnaeus, 1758 | | Scarabaeus isidis Castelnau, 1840 | | | Scarabaeus palemo Olivier, 1789 | | | Scarabaeus proboscideus Guérin-Ménev..., 1844 | | | Scarabaeus proserpinae O. Heer, 1862 | | | Scarabaeus rufus (DeGeer, 1778) | ⇒ Rhysothorax rufus Fabricius, 1792 | | Scarabaeus rugosus (Hausmann, 1807) | ⇒ Onthophagus taurus Schreber, 1759 | | Scarabaeus spretus zur Strassen, 1962 | | | Scarabaeus suri Hausmann, 1807 | | | Scarabaeus viator Péringuey, 1901 | | | Scarabaeus zambesianus Péringuey, 1901 | | | Sceliages granulatus Forgie & Gre..., 2002 | | | Scybalocanthon nigriceps Harold, 1868 | | | Scybalocanthon sexspilotus Guérin-Ménev..., 1855 | | | Sinodrepanus falsus Sharp, 1875 | | | Sisyphus caffer Boheman, 1857 | | | Sisyphus costatus Thunberg, 1818 | | | Sisyphus crispatus Gory, 1833 | | | Sisyphus fasciculatus Boheman, 1857 | | | Sisyphus fortuitus Péringuey, 1900 | | | Sisyphus gazanus Arrow, 1909 | | | Sisyphus goryi (Harold, 1859) | ⇒ Sisyphus caffer goryi Harold, 1859 | | Sisyphus impressipennis Lansberge, 1886 | | | Sisyphus manni Montreuil, 2015 | | | Sisyphus schaefferi Linnaeus, 1758 | | | Sisyphus schaefferi boschniaki Fischer von ..., 1823 | | | Sisyphus schaefferi schaefferi Linnaeus, 1758 | | | Sisyphus seminulum (Gerstaecker, 1871) | ⇒ Sisyphus costatus Thunberg, 1818 | | Sisyphus sordidus (Boheman, 1857) | ⇒ Sisyphus caffer Boheman, 1857 | | Sisyphus spinipes (Thunberg, 1818) | ⇒ Neosisyphus spinipes Thunberg, 1818 | | Stiptopodius patrizii Boucomont, 1923 | | | Sulcophanaeus menelas Castelnau, 1840 | | | Synapsis aquisgranensis Forster, 1869 | | | Temnoplectron aeneopiceum Matthews, 1974 | | | Temnoplectron cooki Reid & Storey, 2000 | | | Temnoplectron politulum MacLeay, 1887 | | | Temnoplectron rotundum Westwood, 1841 | | | Tesserodon novaehollandiae Fabricius, 1775 | | | Tibiodrepanus sulcicollis Castelnau, 1840 | | | Tiniocellus dolosus Branco, 2010 | | | Tiniocellus eurypygus Branco, 2010 | | | Tiniocellus setifer Kraatz, 1895 | | | Tiniocellus spinipes Roth, 1851 | | | Uroxys boneti Pereira & Ha..., 1961 | | | Uroxys micros Bates, 1887 | | | Uroxys pauliani Balthasar, 1940 | | | Uroxys tuberculatus Lansberge, 1874 | | | Xinidium dentilabris Harold, 1869 | |
Subnodes of Scarabaeinae with occurrences in Europe | Tribe | Ateuchini
Montreuil, 1998
| 8
20 | Genera
Species |
Taxom
| | Tribe | Canthonini
Péringuey, 1900
| 25
76 | Genera
Species |
Taxom
| | Tribe | Coprini
Kolbe, 1805
| 14
118 | Genera
Species |
Taxom
| | Tribe | Deltochilini
Lacordaire, 1856
| 2
9 | Genera
Species |
Taxom
| | Tribe | Gymnopleurini
Lacordaire, 1856
| 4
29 | Genera
Species |
Taxom
| | Tribe | Oniticellini
d´Orbigny, 1916
| 13
43 | Genera
Species |
Taxom
| | Tribe | Onitini
Hope, 1837
| 4
43 | Genera
Species |
Taxom
| | Tribe | Onthophagini
Lacordaire, 1856
| 13
336 | Genera
Species |
Taxom
| | Tribe | Phanaeini
MacLeay, 1819
| 5
19 | Genera
Species |
Taxom
| | Tribe | Scarabaeini
Latreille, 1802
| 5
46 | Genera
Species |
Taxom
| | Tribe | Sisyphini
Mulsant, 1842
| 2
20 | Genera
Species |
Taxom
|
Links, sources and further information Scarabaeinae at artfakta.se
Scarabaeinae at dyntaxa.se
Scarabaeinae at itis.gov
|
