Checklist for Subfamily Scarabaeinae Latreille, 1802 (Dung Beetles) and region Caucasus (Eurasia)
Class | Insecta Linnaeus, 1758 Insects, True insects | 523 6140 18845 | Families Genera Species | Taxom | Subclass | Pterygota Lang, 1888 Winged insects | 521 6137 18842 | Families Genera Species | Taxom | Infraclass | Neoptera van der Wulp, 1890 Modern Wing-folding insects | 497 6044 18637 | Families Genera Species | Taxom | Superorder | Holometabola Heider, 1889 Holometabolous Insects | 331 4886 16042 | Families Genera Species | Taxom | Order | Coleoptera Linnaeus, 1758 Beetles, Coleopterans | 98 1952 8628 | Families Genera Species | Taxom | Suborder | Polyphaga Emery, 1886 Polyphags | 93 1701 7123 | Families Genera Species | Taxom | Infraorder | Scarabaeiformia Crowson, 1960
| 11 207 655 | Families Genera Species | Taxom | Superfamily | Scarabaeoidea Latreille, 1802 Bess Beetles, Chafer, Dung Beetles, D´or Beetles | 11 207 655 | Families Genera Species | Taxom | | Family | Scarabaeidae Latreille, 1802 Dung beetles, Flower beetles, Fruit chafers, Rain beetles | 177 565 | Genera Species | Taxom | | Subfamily | Scarabaeinae Latreille, 1802 Dung Beetles, True dung beetles | 19 101 | Genera Species | Taxom |
Genera for Caucasus 19 Genera with 101 Species | Caccobius Thomson, 1767 | | 3 | Species | Taxom | | Canthon Hoffmannsegg, 1858 | Tumblebug | 6 | Species | Taxom | | Cheironitis van Lansberge, 1849 | | 3 | Species | Taxom | | Copris Geoffroy, 1773 | Mexican dung beetle | 7 | Species | Taxom | | Deltochilum Eschscholtz, 1887 | | 2 | Species | Taxom | | Dichotomius Hope, 1767 | | 1 | Species | Taxom | | Digitonthophagus Balthasar, 1787 | | 1 | Species | Taxom | | Epirinus Reiche, 1775 | | 1 | Species | Taxom | | Euoniticellus Janssens, 1781 | | 3 | Species | Taxom | | Euonthophagus Balthasar, 1790 | | 4 | Species | Taxom | | Gymnopleurus Illiger, 2011 | | 10 | Species | Taxom | | Melanocanthon Halffter, 1920 | | 1 | Species | Taxom | | Onitis Fabricius, 1771 | | 2 | Species | Taxom | | Onthophagus Latreille, 1939 | | 56 | Species | Taxom | | Paroniticellus Balthasar, 1809 | | 1 | Species | Taxom | | Phanaeus MacLeay, 1819 | Burrowing Dung Beetles | 2 | Species | Taxom | | Pseudocanthon Bates, 1847 | | 1 | Species | Taxom | | Scarabaeus Linnaeus, 1849 | | 6 | Species | Taxom | | Sisyphus Latreille, 1758 | | 2 | Species | Taxom |
Species list for Caucasus 101 Species and 15 Subspecies | Caccobius histeroides Ménétriés, 1832 | | | Caccobius mundus Ménétriés, 1838 | | | Caccobius schreberi Linnaeus, 1767 | | | Canthon (Boreocanthon) depressipennis LeConte, 1859 | | | Canthon (Boreocanthon) lecontei Harold, 1868 | | | Canthon (Boreocanthon) probus Germar, 1824 | | | Canthon (Canthon) imitator Brown, 1946 | | | Canthon (Canthon) pilularius Linnaeus, 1758 | Dull tumblebug | | Canthon (Canthon) vigilans LeConte, 1858 | Kalia | | Cheironitis haroldi Ballion, 1870 | | | Cheironitis moeris Pallas, 1781 | | | Cheironitis pamphilus Ménétriés, 1849 | | | Copris (Copris) kartlinus ♰ O. N. Kabakov, 1988 | | | Copris armeniacus Falderman, 1835 | | | Copris felschei (Reitter, 1893) | ⇒ Copris armeniacus Falderman, 1835 | | Copris hispanus cavolinii Petagna, 1792 | | | Copris howdeni Matthews & H..., 1959 | | | Copris lunaris Linnaeus, 1758 | Horned Dung Beetle | | Copris minutus Drury, 1773 | Small Black Dung Beetle | | Deltochilum gibbosum Fabricius, 1775 | Humpback Dung Beetle | | Deltochilum valgum acropyge Bates, 1887 | | | Dichotomius carolinus Linnaeus, 1767 | Carolina copris | | Digitonthophagus gazella Fabricius, 1787 | Brown dung beetle | | Epirinus flagellatus Fabricius, 1775 | | | Euoniticellus fulvus Goeze, 1777 | | | Euoniticellus pallens Olivier, 1789 | | | Euoniticellus pallipes Fabricius, 1781 | Southern sandy dung beetle | | Euonthophagus amyntas Olivier, 1789 | | | Euonthophagus amyntas subviolaceus Ménétries, 1832 | | | Euonthophagus atramentarius Ménétriés, 1832 | | | Euonthophagus gibbosus Scriba, 1790 | | | Gymnopleurus aciculatus Gebler, 1841 | | | Gymnopleurus flagellatus Fabricius, 1787 | | | Gymnopleurus flagellatus asperatus Motschulsky, 1849 | | | Gymnopleurus flagellatus flagellatus Fabricius, 1787 | | | Gymnopleurus geoffroyi Fuessly, 1775 | | | Gymnopleurus geoffroyi cyanescens Motschulsky, 1849 | | | Gymnopleurus geoffroyi geoffroyi | | | Gymnopleurus mopsus Pallas, 1781 | | | Gymnopleurus mopsus mopsus | | | Gymnopleurus qurosh Montreuil, 2011 | | | Melanocanthon granulifer Schmidt, 1920 | Pearl blenny | | Onitis damoetas Steven, 1806 | | | Onitis humerosus Pallas, 1771 | | | Onthophagus (Furconthophagus) furcatus (Fabricius, 1781) | ⇒ Onthophagus furcaticeps Masters, 1886 | | Onthophagus (Onthophagus) illyricus Scopoli, 1763 | | | Onthophagus (Onthophagus) oklahomensis Brown, 1927 | Oklahoma dung beetle | | Onthophagus (Onthophagus) taurus Schreber, 1759 | Bull Headed Dung Beetle | | Onthophagus (Onthophagus) tuberculifrons Harold, 1871 | Dung beetle | | Onthophagus (Palaeonthophagus) angorensis Petrovitz, 1963 | | | Onthophagus (Palaeonthophagus) coenobita Herbst, 1783 | | | Onthophagus (Palaeonthophagus) conspersus Reitter, 1893 | | | Onthophagus (Palaeonthophagus) cruciatus (Ménétriés, 1832) | ⇒ Onthophagus lemur Fabricius, 1781 | | Onthophagus (Palaeonthophagus) fissicornis Steven, 1809 | | | Onthophagus (Palaeonthophagus) fortigibber Reitter, 1909 | | | Onthophagus (Palaeonthophagus) fracticornis (Preyssler, 1790) | ⇒ Onthophagus similis Scriba, 1790 | | Onthophagus (Palaeonthophagus) gibbulus Pallas, 1781 | | | Onthophagus gibbulus rostrifer Reitter, 1892 | | | Onthophagus (Palaeonthophagus) grossepunctatus Reitter, 1905 | | | Onthophagus (Palaeonthophagus) kolenatii Reitter, 1893 | | | Onthophagus (Palaeonthophagus) lemur Fabricius, 1781 | | | Onthophagus (Palaeonthophagus) lucidus Illiger, 1800 | | | Onthophagus (Palaeonthophagus) marginalis Gebler, 1817 | | | Onthophagus marginalis marginalis | | | Onthophagus (Palaeonthophagus) medius (Kugelann, 1792) | ⇒ Onthophagus vacca Linnaeus, 1767 | | Onthophagus (Palaeonthophagus) nuchicornis Linnaeus, 1758 | Mottled dung beetle | | Onthophagus (Palaeonthophagus) opacicollis Reitter, 1893 | | | Onthophagus (Palaeonthophagus) ovatus Linnaeus, 1767 | | | Onthophagus (Palaeonthophagus) parmatus Reitter, 1892 | | | Onthophagus (Palaeonthophagus) ponticus Harold, 1883 | | | Onthophagus (Palaeonthophagus) pygargus Motschulsky, 1845 | | | Onthophagus (Palaeonthophagus) rakovici Pittino, 2004 | | | Onthophagus ruficapillus ruficapillus Brullé, 1832 | | | Onthophagus (Palaeonthophagus) semicornis Panzer, 1798 | | | Onthophagus (Palaeonthophagus) sericatus Reitter, 1893 | | | Onthophagus (Palaeonthophagus) similis (Scriba, 1790) | ⇒ Onthophagus fracticornis Preyssler, 1790 | | Onthophagus (Palaeonthophagus) speculifer Solsky, 1876 | | | Onthophagus (Palaeonthophagus) suturellus Brullé, 1832 | | | Onthophagus (Palaeonthophagus) truchmenus Kolenati, 1846 | | | Onthophagus truchmenus truchmenus | | | Onthophagus (Palaeonthophagus) vacca Linnaeus, 1767 | | | Onthophagus (Palaeonthophagus) verticicornis (Laicharting, 1781) | ⇒ Onthophagus nuchicornis Linnaeus, 1758 | | Onthophagus (Palaeonthophagus) viridis Ménétriés, 1832 | | | Onthophagus (Palaeonthophagus) vitulus (Fabricius, 1776) | ⇒ Onthophagus immundus Boheman, 1858 | | Onthophagus (Paleonthophagus) andalusicus Waltl, 1835 | | | Onthophagus andalusicus persianus Olsoufieff, 1900 | | | Onthophagus (Paleonthophagus) nuchicornis Linnaeus, 1758 | | | Onthophagus (Paleonthophagus) similis Scriba, 1790 | | | Onthophagus (Paleonthophagus) vacca Linnaeus, 1767 | | | Onthophagus aerarius Reitter, 1892 | | | Onthophagus basipustulatus Heyden, 1879 | | | Onthophagus dorsosignatus d´Orbigny, 1898 | | | Onthophagus formaneki Reitter, 1897 | | | Onthophagus furcaticeps Masters, 1886 | | | Onthophagus immundus Boheman, 1858 | | | Onthophagus liothorax Koshantschikov, 1894 | | | Onthophagus sacharovskii Olsoufieff, 1918 | | | Onthophagus suermelii Petrovitz, 1963 | | | Onthophagus zinovskyi Qaryagdy, 1939 | | | Onthophagus zuvandi Qaryagdy, 1939 | | | Paroniticellus festivus Steven, 1809 | | | Phanaeus (Phanaeus) igneus MacLeay, 1819 | Laguna Madre fiddler | | Phanaeus (Phanaeus) vindex MacLeay, 1819 | Laguna Madre fiddler | | Pseudocanthon perplexus LeConte, 1847 | | | Scarabaeus (Ateuchetus) armeniacus Ménétriés, 1832 | | | Scarabaeus (Scarabaeus) babori Balthasar, 1934 | | | Scarabaeus (Scarabaeus) pius Illiger, 1803 | | | Scarabaeus (Scarabaeus) sacer (Linnaeus, 1758) | ⇒ Scarabaeus pius Illiger, 1803 | | Scarabaeus (Scarabaeus) typhon (Fischer von ..., 1823) | ⇒ Megasoma gyas Herbst, 1785 | | Scarabaeus acuticollis Motschulsky, 1849 | | | Sisyphus schaefferi boschniaki Fischer von ..., 1823 | | | Sisyphus schaefferi schaefferi Linnaeus, 1758 | |
Subnodes of Scarabaeinae with occurrences in Caucasus | Tribe | Ateuchini Montreuil, 1998
| 0 0 | Genera Species | Taxom
| | Tribe | Canthonini Péringuey, 1900
| 5 11 | Genera Species | Taxom
| | Tribe | Coprini Kolbe, 1805
| 2 8 | Genera Species | Taxom
| | Tribe | Gymnopleurini Lacordaire, 1856
| 1 5 | Genus Species | Taxom
| | Tribe | Oniticellini d´Orbigny, 1916
| 2 4 | Genera Species | Taxom
| | Tribe | Onitini Hope, 1837
| 2 5 | Genera Species | Taxom
| | Tribe | Onthophagini Lacordaire, 1856
| 4 59 | Genera Species | Taxom
| | Tribe | Phanaeini MacLeay, 1819
| 1 2 | Genus Species | Taxom
| | Tribe | Scarabaeini Latreille, 1802
| 1 6 | Genus Species | Taxom
| | Tribe | Sisyphini Mulsant, 1842
| 1 1 | Genus Species | Taxom
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Links, sources and further information Scarabaeinae at artfakta.se Scarabaeinae at dyntaxa.se Scarabaeinae at itis.gov
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