Species list, Checklist of Supertribe Harpalitae for Yugoslavia (Southeast Europe)

Class	Insecta Linnaeus, 1758 Insects, True insects	505 6035 20701	Families Genera Species	Taxom
Subclass	Pterygota Lang, 1888 Winged insects	503 6027 20691	Families Genera Species	Taxom
Infraclass	Neoptera van der Wulp, 1890 Modern Wing-folding insects	477 5958 20563	Families Genera Species	Taxom
Superorder	Holometabola Heider, 1889 Holometabolous Insects	316 4667 17402	Families Genera Species	Taxom
Order	Coleoptera Linnaeus, 1758 Beetles, Coleopterans	91 1829 7556	Families Genera Species	Taxom
Suborder	Adephaga Schellenberg, 1806 Adephagans, Ground Beetles and Water Beetles	5 203 1475	Families Genera Species	Taxom
Superfamily	Caraboidea Latreille, 1802	1 179 1418	Family Genera Species	Taxom
Family	Carabidae Latreille, 1802 Carabid beetles, Carabids, Ground Beetles, Tiger Beetles	179 1418	Genera Species	Taxom
Subfamily	Harpalinae Bonelli, 1810	89 668	Genera Species	Taxom
Supertribe	Harpalitae Erwin, 1985	25 182	Genera Species	Taxom

Genera for Yugoslavia
25 Genera with 182 Species

Acinopus Dejean, 1829		5	Species	Taxom
Acupalpus Latreille, 1884		14	Species	Taxom
Amblystomus Erichson, 1883		8	Species	Taxom
Anisodactylus Dejean, 1933		10	Species	Taxom
Anthracus Motschulsky, 1862		5	Species	Taxom
Bradycellus Erichson, 1812		8	Species	Taxom
Carterus Dana, 1873		4	Species	Taxom
Cryptophonus Brandmayr & ..., 1829		3	Species	Taxom
Daptus Fischer von ..., 1823		1	Species	Taxom
Diachromus Erichson, 1758		1	Species	Taxom
Dicheirotrichus Jacquelin du..., 1827		4	Species	Taxom
Ditomus Dana, 1792		3	Species	Taxom
Dixus Billberg, 1825		3	Species	Taxom
Graniger Motschulsky, 1821		1	Species	Taxom
Gynandromorphus Dejean, 1905		3	Species	Taxom
Harpalus Latreille, 1847		90	Species	Taxom
Ophonus Dejean, 1828		27	Species	Taxom
Pachycarus Solier, 1997		4	Species	Taxom
Pangus Dejean, 1818		1	Species	Taxom
Parophonus Ganglbauer, 1829		6	Species	Taxom
Pseudoophonus Motschulsky, 1774		3	Species	Taxom
Scybalicus Schaum, 1829		1	Species	Taxom
Stenolophus Dejean, 1781	Seedcorn Beetle	10	Species	Taxom
Trichotichnus A. Morawitz, 1812		2	Species	Taxom
Tschitscherinellus Csiki, 1825		1	Species	Taxom

Species list for Yugoslavia
182 Species and 37 Subspecies

Acinopus (Acinopus) laevigatus Ménétriés, 1832
Acinopus laevigatus laevigatus Ménétriés..., 1832
Acinopus (Acinopus) picipes Olivier, 1795
Acinopus (Oedematicus) megacephalus (P. Rossi, 1794)	⇒ Acinopus picipes Olivier, 1795
Acinopus (Osimus) ammophilus Dejean, 1829
Acupalpus (Acupalpus) brunnipes Sturm, 1825
Acupalpus (Acupalpus) dubius Schilsky, 1888
Acupalpus (Acupalpus) elegans Dejean, 1829
Acupalpus (Acupalpus) exiguus Dejean, 1829
Acupalpus (Acupalpus) flavicollis Sturm, 1825
Acupalpus (Acupalpus) luteatus (Duftschmid, 1812)	⇒ Acupalpus dubius Schilsky, 1888
Acupalpus (Acupalpus) maculatus Schaum, 1860
Acupalpus (Acupalpus) meridianus Linnaeus, 1761	Midday harp ground beetle
Acupalpus (Acupalpus) parvulus Sturm, 1825
Acupalpus (Acupalpus) suturalis (Dejean, 1829)	⇒ Acupalpus meridianus Linnaeus, 1761
Acupalpus suturalis suturalis Dejean, 1829
Acupalpus (Ancylostria) interstitialis Reitter, 1884
Acupalpus (Tachistodes) notatus Mulsant & Rey, 1861
Acupalpus (Tachistodes) paludicola Reitter, 1884
Amblystomus levantinus Reitter, 1883
Amblystomus levantinus levantinus Reitter, 1883
Amblystomus metallescens Dejean, 1829
Amblystomus niger (Heer, 1841)	⇒ Amblystomus nigrinus Csiki, 1931
Amblystomus nigrinus Csiki, 1931
Amblystomus palustris Blackburn, 1888
Amblystomus picinus Baudi Di Selva, 1864
Amblystomus rectangulus Reitter, 1883
Anisodactylus (Anisodactylus) binotatus Fabricius, 1787	Two-marked harp ground beetle
Anisodactylus (Anisodactylus) nemorivagus Duftschmid, 1812	Heath Short-spur
Anisodactylus (Hexatrichus) poeciloides Stephens, 1828	Saltmarsh Short-spur
Anisodactylus poeciloides poeciloides Stephens, 1828
Anisodactylus poeciloides pseudoaeneus Dejean, 1829
Anisodactylus (Hexatrichus) virens Dejean, 1829
Anisodactylus (Pseudanisodactylus) signatus Panzer, 1796
Anisodactylus (Pseudodichirus) intermedius Dejean, 1829
Anisodactylus (Spongopus) pueli Schauberger, 1933
Anisodactylus pueli pueli Schauberger, 1933
Anthracus consputus Duftschmid, 1812
Anthracus insignis Reitter, 1884
Anthracus longicornis Schaum, 1857
Anthracus quarnerensis Reitter, 1884
Anthracus transversalis Schaum, 1862
Bradycellus (Stenocellus) nebulosus LeConte, 1853
Bradycellus (Triliarthrus) caucasicus Chaudoir, 1846
Bradycellus (Triliarthrus) csikii Laczó, 1912
Bradycellus (Triliarthrus) distinctus Dejean, 1829
Bradycellus (Triliarthrus) ganglbaueri Apfelbeck, 1904
Bradycellus (Triliarthrus) harpalinus Audinet-Serv..., 1821	European harp ground beetle
Bradycellus (Triliarthrus) ruficollis Stephens, 1828
Bradycellus (Triliarthrus) verbasci Duftschmid, 1812
Carterus (Pristocarterus) angustipennis Chaudoir, 1852
Carterus angustipennis lutshniki Zamotailov, 1988
Carterus (Pristocarterus) dama P.Rossi, 1792
Carterus (Pristocarterus) gilvipes Piochard de ..., 1873
Cryptophonus litigiosus Dejean, 1829
Cryptophonus melancholicus Dejean, 1829
Cryptophonus tenebrosus Dejean, 1829
Daptus vittatus Fischer von ..., 1823
Diachromus germanus Linnaeus, 1758
Dicheirotrichus (Trichocellus) discicollis Dejean, 1829
Dicheirotrichus (Trichocellus) lacustris L.Redtenbacher, 1858
Dicheirotrichus (Trichocellus) obsoletus Dejean, 1829
Dicheirotrichus (Trichocellus) rufithorax C.R.Sahlberg, 1827
Ditomus calydonius P. Rossi, 1790
Ditomus calydonius calydonius P. Rossi, 1790
Ditomus tricuspidatus Fabricius, 1792
Dixus clypeatus P. Rossi, 1790
Dixus eremita Dejean, 1825
Dixus obscurus Dejean, 1825
Graniger cordicollis Audinet-Serv..., 1821
Gynandromorphus etruscus Quensel, 1806
Gynandromorphus etruscus etruscus Quensel, 1806
Gynandromorphus etruscus peyroni Carret, 1905
Harpalus (Acardystus) atratus Latreille, 1804
Harpalus (Acardystus) attenuatus Stephens, 1828
Harpalus (Acardystus) flavescens Piller &Mitt..., 1783
Harpalus (Acardystus) pygmaeus Dejean, 1829
Harpalus (Actephilus) albanicus Reitter, 1900
Harpalus (Actephilus) picipennis Duftschmid, 1812
Harpalus (Actephilus) pumilus Sturm, 1818
Harpalus (Amblystus) latus Linnaeus, 1758
Harpalus (Amblystus) luteicornis Dufischmid, 1812
Harpalus (Amblystus) marginellus Gyllenhal, 1827
Harpalus (Amblystus) rubripes Dufischmid, 1812	Bareyed goby
Harpalus (Amblystus) solitaris Dejean, 1829	Checked goby
Harpalus (Artabas) dispar Dejean, 1829
Harpalus dispar splendens Gebler, 1829
Harpalus (Artabas) punctatostriatus (Dejean, 1829)	⇒ Harpalus rubefactus rubefactus Bates, 1873
Harpalus (Artabas) rumelicus Apfelbeck, 1904
Harpalus (Cephalophonus) cephalotes (Fairmaire & ..., 1854)	⇒ Harpalus capito A.Morawitz, 1862
Harpalus cephalotes cephalotes
Harpalus (Cryptophonus) litigiosus (Dejean, 1829)	⇒ Cryptophonus litigiosus Dejean, 1829
Harpalus litigiosus litigiosus Dejean, 1829
Harpalus (Cryptophonus) melancholicus (Dejean, 1829)	⇒ Cryptophonus melancholicus Dejean, 1829
Harpalus melancholicus melancholicus
Harpalus melancholicus reicheianus Kataev, 2012
Harpalus (Cryptophonus) tenebrosus (Dejean, 1829)	⇒
Harpalus tenebrosus tenebrosus
Harpalus (Euharpalops) fraternus LeConte, 1852
Harpalus (Harpalobius) autumnalis Duftschmid, 1812
Harpalus (Harpalobius) froelichii Sturm, 1818	Brush-thighed Seed-eater
Harpalus (Harpalobius) fuscipalpis Sturm, 1818
Harpalus (Harpalobius) hirtipes Panzer, 1796
Harpalus (Harpaloderus) honestus Dufischmid, 1812	St. Bees Seed-eater
Harpalus (Harpaloderus) neglectus Serville, 1821
Harpalus neglectus neglectus Audinet-Serv..., 1821
Harpalus (Harpaloderus) rufipalpis Sturm, 1818
Harpalus rufipalpis rufipalpis Sturm, 1818
Harpalus (Harpaloderus) sulphuripes Germar, 1824
Harpalus sulphuripes sulphuripes Germar, 1824
Harpalus (Harpalophonus) hospes Sturm, 1818
Harpalus hospes hospes Sturm, 1818
Harpalus (Harpalus) affinis (Schrank, 1781)	⇒ Harpalus serripes ernsti Kataev, 1995
Harpalus (Harpalus) brevicornis Germar, 1824
Harpalus (Harpalus) caspius Steven, 1806
Harpalus (Harpalus) kazanensis JedliÄka, 1958
Harpalus (Harpalus) oblitus (Dejean, 1829)	⇒ Harpalus fraternus LeConte, 1852
Harpalus oblitus oblitus Dejean, 1829
Harpalus (Harpalus) progrediens Schauberger, 1922
Harpalus (Harpalus) saxicola Dejean, 1829
Harpalus (Harpalus) somnulentus Dejean, 1829	Checked goby
Harpalus (Harpalus) subcylindricus Dejean, 1829
Harpalus (Harpalus) xanthopus Gemminger & ..., 1868
Harpalus xanthopus winkleri Schauberger, 1923
Harpalus (Harpalus) zabroides Dejean, 1829
Harpalus (Lasioharpalus) cupreus Dejean, 1829
Harpalus cupreus fastuosus Faldermann, 1835
Harpalus (Lasioharpalus) dimidiatus P. Rossi, 1790
Harpalus (Lasioharpalus) distinguendus Duftschmid, 1812
Harpalus distinguendus distinguendus Duftschmid, 1812
Harpalus (Lasioharpalus) smaragdinus Duftschmid, 1812
Harpalus rubefactus rubefactus Bates, 1873
Harpalus (Opadius) laevipes Zetterstedt, 1828
Harpalus (Pheuginus) anxius Duftschmid, 1812
Harpalus (Pheuginus) flavicornis Dejean, 1829
Harpalus flavicornis flavicornis Dejean, 1829
Harpalus (Pheuginus) modestus Dejean, 1829
Harpalus (Pheuginus) politus Dejean, 1829
Harpalus politus politus Dejean, 1829
Harpalus (Pheuginus) serripes Quensel, 1806
Harpalus serripes ernsti Kataev, 1995
Harpalus serripes serripes Quensel, 1806
Harpalus (Pheuginus) servus Duftschmid, 1812
Harpalus (Pheuginus) taciturnus Dejean, 1829
Harpalus (Pheuginus) tardus Panzer, 1796
Harpalus (Plectralidus) angulatus Putzeys, 1878
Harpalus angulatus scytha Tschitschér..., 1899
Harpalus (Plectralidus) danieli Reitter, 1900
Harpalus (Plectralidus) euchlorus Ménétriés, 1839
Harpalus (Plectralidus) fuscicornis Ménétriés, 1832
Harpalus (Plectralidus) karamani Apfelbeck, 1902
Harpalus karamani hoffmanni Schauberger, 1928
Harpalus karamani karamani Apfelbeck, 1902
Harpalus (Plectralidus) triseriatus A. Fleischer, 1897
Harpalus triseriatus babunensis Mlynar, 1979
Harpalus triseriatus triseriatus Fleischer, 1897
Harpalus (Pseudoophonus) calceatus (Duftschmid, 1812)	⇒ Pseudoophonus calceatus Duftschmid, 1812
Harpalus (Pseudoophonus) griseus (Panzer, 1796)	⇒ Pseudoophonus griseus Panzer, 1797
Harpalus (Pseudoophonus) rufipes (DeGeer, 1774)	⇒
Harpalus (Semiophonus) signaticornis Duftschmid, 1812
Harpalus peritus (Casey, 1914)	⇒ Harpalus somnulentus Dejean, 1829
Harpalus reitteri Wrase & Kataev, 2011
Harpalus tabidus ♰ O. Heer, 1847
Ophonus (Hesperophonus) azureus Fabricius, 1775
Ophonus (Hesperophonus) convexicollis Ménétriés, 1832
Ophonus (Hesperophonus) cribricollis Dejean, 1829
Ophonus (Hesperophonus) rotundatus Dejean, 1829
Ophonus (Hesperophonus) similis Dejean, 1829
Ophonus (Hesperophonus) subquadratus (Dejean, 1829)	⇒
Ophonus (Incisophonus) incisus Dejean, 1829
Ophonus (Macrophonus) oblongus Schaum, 1858
Ophonus (Metophonus) brevicollis (Audinet-Serv..., 1821)	⇒ Ophonus melletii Heer, 1837
Ophonus (Metophonus) cordatus Duftschmid, 1812
Ophonus (Metophonus) gammeli Schauberger, 1932
Ophonus (Metophonus) laticollis (Mannerheim, 1825)	⇒ Ophonus puncticollis Paykull, 1798
Ophonus (Metophonus) melletii Heer, 1837	Mellet's Downy-back
Ophonus melletii melletii Heer, 1837
Ophonus (Metophonus) parallelus (Dejean, 1829)	⇒ Ophonus xaxarsi Schauberger, 1928
Ophonus (Metophonus) puncticeps (Stephens, 1828)	⇒ Ophonus puncticollis Paykull, 1798
Ophonus (Metophonus) puncticollis Paykull, 1798
Ophonus (Metophonus) rufibarbis Fabricius, 1792
Ophonus (Metophonus) rupicola Sturm, 1818
Ophonus (Metophonus) schaubergerianus (Puel, 1937)	⇒ Ophonus melletii Heer, 1837
Ophonus (Metophonus) veluchianus G.Muller, 1931
Ophonus (Metophonus) xaxarsi Schauberger, 1928
Ophonus (Ophonus) ardosiacus Lutshnik, 1922
Ophonus (Ophonus) diffinis (Dejean, 1829)	⇒ Ophonus ardosiacus Lutshnik, 1922
Ophonus (Ophonus) franziniorum Sciaky, 1987
Ophonus (Ophonus) sabulicola Panzer, 1796
Ophonus (Ophonus) stictus Stephens, 1828	Oolite Downy-back
Pachycarus (Mystropterus) atrocoeruleus Dejean, 1830
Pachycarus atrocoeruleus atrocoeruleus Waltl, 1838
Pachycarus (Mystropterus) cyaneus Dejean, 1830
Pachycarus (Mystropterus) macedonicus V. & B.Gueor..., 1997
Pangus scaritides Sturm, 1818
Parophonus (Ophonomimus) hirsutulus Dejean, 1829
Parophonus (Parophonus) dejeani Csiki, 1932
Parophonus (Parophonus) laeviceps Ménétriés, 1832
Parophonus (Parophonus) maculicornis Duftschmid, 1812
Parophonus (Parophonus) mendax P.Rossi, 1790
Parophonus (Parophonus) planicollis (Dejean, 1829)	⇒ Parophonus laeviceps Ménétriés, 1832
Pseudoophonus (Platus) calceatus Duftschmid, 1812
Pseudoophonus (Pseudoophonus) griseus Panzer, 1797
Pseudoophonus (Pseudoophonus) rufipes De Geer, 1774
Scybalicus oblongiusculus Dejean, 1829
Stenolophus (Egadroma) marginatus Dejean, 1829
Stenolophus (Stenolophus) abdominalis Géné, 1836
Stenolophus abdominalis persicus Mannerheim, 1844
Stenolophus (Stenolophus) discophorus Fischer von ..., 1823
Stenolophus (Stenolophus) mixtus Herbst, 1784
Stenolophus (Stenolophus) narentinus Drovenik & Peks, 1999
Stenolophus (Stenolophus) proximus Dejean, 1829
Stenolophus (Stenolophus) skrimshiranus Stephens, 1828
Stenolophus (Stenolophus) steveni Krynicki, 1832
Stenolophus (Stenolophus) teutonus Schrank, 1781
Trichotichnus (Trichotichnus) knauthi Ganglbauer, 1901
Trichotichnus (Trichotichnus) laevicollis Duftschmid, 1812
Tschitscherinellus cordatus Dejean, 1825

Subnodes of Harpalitae with occurrences in Yugoslavia

Tribe

Harpalini
Bonelli, 1810

25
182

Genera
Species

Taxom