Taxonomy of Genus Lethrus Scopoli, 1777
Class
| Insecta
Linnaeus, 1758
Insects, True insects | 4179
190570
2444470 | Families
Genera
Species |
Subclass
| Pterygota
Lang, 1888
Winged insects | 3816
189502
2439455 | Families
Genera
Species |
Infraclass
| Neoptera
van der Wulp, 1890
Modern Wing-folding insects | 3399
186457
2415487 | Families
Genera
Species |
Superorder
| Holometabola
Heider, 1889
Holometabolous Insects | 1891
142849
2048615 | Families
Genera
Species |
Order
| Coleoptera
Linnaeus, 1758
Beetles, Coleopterans | 592
62779
813516 | Families
Genera
Species |
Suborder
| Polyphaga
Emery, 1886
Polyphags | 274
54416
687671 | Families
Genera
Species |
Infraorder
| Scarabaeiformia
Crowson, 1960
| 32
5275
90050 | Families
Genera
Species |
Superfamily
| Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles | 32
5275
90050 | Families
Genera
Species | | Family | Geotrupidae
Latreille, 1802
Dor beetles, Dung beetles, Earth-Boring Dung Beetle | 165
2744 | Genera
Species | | Subfamily | Geotrupinae
Latreille, 1802
| 39
1325 | Genera
Species | | Tribe | Lethrini
Oken, 1843
| 13
947 | Genera
Species | | Genus | Lethrus
Scopoli, 1777
| 228 | Species |
Subnodes of Lethrus Scopoli, 1777 | Subgenus | Lethrus (Abrognathus)
Jakovlev, 1890
| 4 | Species | | Subgenus | Lethrus (Autolethrus)
| 3 | Species | | Subgenus | Lethrus (Ceratodirus)
Fischer von Waldheim, 1845
| 12 | Species | | Subgenus | Lethrus (Furcilethrus)
Nikolayev, 1968
| 20 | Species | | Subgenus | Lethrus (Heteroplistodus)
Jakovlev, 1890
| 19 | Species | | Subgenus | Lethrus (Lethrus)
Scopoli, 1777
| 35 | Species | | Subgenus | Lethrus (Mesolethrus)
Nikolajev, 2003
| 5 | Species | | Subgenus | Lethrus (Neolethrus)
Nikolaev, 1987
| 1 | Species | | Subgenus | Lethrus (Paralethrus)
Nikolajev, 2003
| 8 | Species | | Subgenus | Lethrus (Scelolethrus)
Semenov, 1892
| 25 | Species | | Species | Lethrus aenescens Fairmaire, 1892 ⇒ Paralethrus turkestanicus
| | | | Species | Lethrus aeneus Fabricius, 1792
| | | | Species | Lethrus afghanus Kral & Olexa, 1996 ⇒ Lethrus (Lethrus) appendiculatus
| | | | Species | Lethrus ahriman Pak & Gubin, 2020
| | | | Species | Lethrus asmodeus Pak & Gubin, 2020
| | | | Species | Lethrus baiocchii Hillert & Sechi, 2014
| | | | Species | Lethrus bituberculifrons Lebedev, 1926 ⇒ Paralethrus bituberculatus
| | | | Species | Lethrus bradytus Fairmaire, 1892 ⇒ Lethrus (Lethrus) scoparius
| | | | Species | Lethrus breviceps Ballion, 1870 ⇒ Abrognathus tuberculifrons
| | | | Species | Lethrus bucharicus Semenov, 1894 ⇒ Scelolethrus kozhantschikovi
| | | | Species | Lethrus conformis Frivaldsky, 1890 ⇒ Heteroplistodus potanini
| | | | Species | Lethrus coptotis Semenov, 1894 ⇒ Lethrus (Lethrus) appendiculatus
| | | | Species | Lethrus degener Medvedev, 1962 ⇒ Lethrus (Lethrus) appendiculatus
| | | | Species | Lethrus desertus Skopina, 1965
| | | | Species | Lethrus dinotherium Wilkins, 1885
| 1 | Subspecies | | Species | Lethrus dispar Jakovlev, 1890 ⇒ Ceratodirus cephalotes
| | | | Species | Lethrus dispars Fischer von Waldheim, 1845 ⇒ Ceratodirus longimanus
| | | | Species | Lethrus dostojewskii Semenov, 1899
| | | | Species | Lethrus eous Semenov, 1894 ⇒ Lethrus (Paralethrus) bituberculatus impressifrons
| | | | Species | Lethrus eversmanni Krynicki, 1832 ⇒ Ceratodirus longimanus
| | | | Species | Lethrus eversmannii Krynicki, 1832
| | | | Species | Lethrus fedtschenkoi Semenov, 1894 ⇒ Lethrus (Lethrus) superbus
| | | | Species | Lethrus fedtshenkoi Semenov, 1894 ⇒ Lethrus (Lethrus) superbus
| | | | Species | Lethrus ferrugineus Eschscholtz, 1818 ⇒ Codocera ferruginea
| | | | Species | Lethrus forcipatus Jakovlev, 1892 ⇒ Lethrus (Lethrus) schaumi
| | | | Species | Lethrus glasunovi Koshantschikov, 1894 ⇒ Mesolethrus microbuccis
| | | | Species | Lethrus glasunowi D. Koshantschikov, 1894 ⇒ Lethrus (Mesolethrus) microbuccis
| | | | Species | Lethrus golestanicus Král & Hillert, 2021
| | | | Species | Lethrus halkidikiensis Kral & Hillert, 2013
| | | | Species | Lethrus hauseri Reitter, 1894 ⇒ Lethrus (Paralethrus) bituberculatus impressifrons
| 1 | Subspecies | | Species | Lethrus heydeni Fairmaire, 1866 ⇒ Lethrus (Lethrus) raymondi
| | | | Species | Lethrus heydenii Fairmaire, 1866 ⇒ Lethrus (Autolethrus) raymondi
| | | | Species | Lethrus impressifrons Ballion, 1870 ⇒ Paralethrus bituberculatus
| | | | Species | Lethrus inconspicuus S. I. Medvedev, 1959
| | | | Species | Lethrus inconspiquus Medvedev, 1959 ⇒ Furcilethrus banghaasi
| | | | Species | Lethrus jacobsoni Semenov & Medvedev, 1935 ⇒ Paralethrus bituberculatus
| | | | Species | Lethrus jakovlevi Semenov, 1891 ⇒ Heteroplistodus bulbocerus
| | | | Species | Lethrus jakowlewi Semenov, 1892 ⇒ Lethrus (Heteroplistodus) bulbocerus
| | | | Species | Lethrus kiritshenkoi Medvedev, 1965
| | | | Species | Lethrus klimenkoi Bagaturov, 2018
| | | | Species | Lethrus komarovi Semenov, 1894 ⇒ Scelolethrus sulcatus
| | | | Species | Lethrus komarowi Semenov, 1894
| | | | Species | Lethrus korzhinskii Semenov, 1899 ⇒ Lethrus (Heteroplistodus) korshinskii
| | | | Species | Lethrus koshantschikowi Semenov, 1892
| | | | Species | Lethrus kozhantchikovi
| | | | Species | Lethrus kozhantschikovi Semenov, 1891 ⇒ Lethrus (Scelolethrus) koshantschikovi
| | | | Species | Lethrus kuldshensis Lebedev, 1926
| | | | Species | Lethrus laevigatus Ballion, 1871 ⇒ Lethrus (Lethrus) scoparius
| | | | Species | Lethrus laevigatusus Ballion, 1870 ⇒ Lethrus (Lethrus) scoparius
| | | | Species | Lethrus litvinovi Semenov, 1900
| | | | Species | Lethrus litvinowi Semenov, 1900
| | | | Species | Lethrus liviae Pittino, 2011
| | | | Species | Lethrus lopatnini Medvedev, 1959
| | | | Species | Lethrus macrodon Nikolajev, 1969
| | | | Species | Lethrus mandibularis Jakovlev, 1892 ⇒ Lethrus (Lethrus) elephas
| | | | Species | Lethrus marquardti Reitter, 1902 ⇒ Lethrus (Lethrus) scoparius
| | | | Species | Lethrus miliaris Skopina, 1965
| | | | Species | Lethrus muskinabadensis Nikolajev & Ivanova, 2014
| | | | Species | Lethrus nazarovi Gusakov, 2017
| | | | Species | Lethrus nigroaeneus Skopin, 1955 ⇒ Lethrus (Paralethrus) bituberculatus impressifrons
| | | | Species | Lethrus obliteratus Semenov, 1894 ⇒ Lethrus (Lethrus) mucronatus
| | | | Species | Lethrus obsoletus Semenov, 1894 ⇒ Lethrus (Furcilethrus) tenuidens obsoletus
| | | | Species | Lethrus paki Ivanova, 2012
| | | | Species | Lethrus percutens Gusakov & Klimenko, 2011
| | | | Species | Lethrus persicus Reitter, 1890 ⇒ Lethrus (Lethrus) acutangulus
| | | | Species | Lethrus perun Kral & Hillert, 2013
| | | | Species | Lethrus petri Semenov & S. I. Medvedev, 1935
| | | | Species | Lethrus petrii Semenov & Medvedev, 1935 ⇒ Heteroplistodus lebedevi
| | | | Species | Lethrus pigmaeus
| | | | Species | Lethrus podolicus Fischer von Waldheim, 1822 ⇒ Lethrus (Lethrus) apterus
| | | | Species | Lethrus puncticollis Kraatz, 1882 ⇒ Mesolethrus microbuccis
| | | | Species | Lethrus rugulosus S. I. Medvedev, 1959 ⇒ Lethrus (Furcilethrus) antovae
| | | | Species | Lethrus rustam Nikolajev & Ivanova, 2014
| | | | Species | Lethrus scarabaeoides Hochenwarth, 1785
| | | | Species | Lethrus scaraboeoides Hochenwarth, 1785
| | | | Species | Lethrus schaumii Reitter, 1890
| | | | Species | Lethrus schneideri Kral & Hillert, 2013
| | | | Species | Lethrus semenovi Kozhantschikov, 1893 ⇒ Furcilethrus furcatus
| | | | Species | Lethrus silus Reitter, 1894 ⇒ Paralethrus turkestanicus
| | | | Species | Lethrus spathulatus Lebedev, 1932
| | | | Species | Lethrus strymonensis Kral & Hillert, 2013
| | | | Species | Lethrus subaeneus Fairmaire, 1866 ⇒ Lethrus (Lethrus) apterus
| | | | Species | Lethrus subaurosus Reitter, 1909 ⇒ Ceratodirus karelini
| | | | Species | Lethrus talassicola Medvedev, 1965 ⇒ Heteroplistodus bispinus
| | | | Species | Lethrus tarbagataicus Nikolajev, 2001
| | | | Species | Lethrus taschkumyricus Protzenko, 1972 ⇒ Lethrus (Lethrus) mucronatus
| | | | Species | Lethrus thracicus Král & Hillert, 2018
| | | | Species | Lethrus toropovi Shapovalov, 2022
| | | | Species | Lethrus tschitscherini Semenov, 1894
| | | | Species | Lethrus tshatkalensis Protsenko, 1965
| | | | Species | Lethrus tshitsherini Seminov, 1894
| | | | Species | Lethrus turcomanicus Semenov, 1894 ⇒ Lethrus (Scelolethrus) substriatus
| | | | Species | Lethrus zarudnianus Semenov & Medvedev, 1935 ⇒ Scelolethrus mithras
| | | | Species | Lethrus zarudnyi Semonow & Medvedev, 1935 ⇒ Paralethrus turkestanicus
| | |
Synonyms | Autolethrus (Semenov, 1892) | | Bulbocerus (Acharius, 1781) | | Clunipes (Hochenwarth, 1785) | | Goniolethrus (Semenov, 1894) | | Lathrus (Gmelin in Linnaeus, 1789) | | Lethrulus (Semenov, 1892) | | Luthres (Latreille, 1802) | | Microlethrus (Semenov, 1894) | | Sclerolethrus (Reitter, 1892) | | Teratolethrus (Semenov, 1894) |
Distribution of Lethrus Scopoli, 1777 Afghanistan, Albania, Austria, Bosnia and Herzegovina, Bulgaria, China, Croatia, Czech Republic, Finland, Greece, Hungary, Iran, Italy, Kazakhstan, Kyrgyzstan, Latvia, Moldova, Mongolia, Norway, Poland, Romania, Russia, Serbia, Slovakia, Sweden, Tajikistan, Transnistria, Turkey, Turkmenistan, Ukraine, Uzbekistan
Links, sources and further information
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