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TAXFAMGENSPESUB
Checklist of Family Passalidae Leach, 1815 (bess beetle)
for Asia (Eurasia)


Class
 
Insecta
Linnaeus, 1758
Insects, True insects
1422
46785
383423
Families
Genera
Species
 
Taxom
 

Subclass
 
Pterygota
Lang, 1888
Winged insects
1416
46713
383230
Families
Genera
Species
 
Taxom
 

Infraclass
 
Neoptera
van der Wulp, 1890
Modern Wing-folding insects
1294
45948
379619
Families
Genera
Species
 
Taxom
 

Superorder
 
Holometabola
Heider, 1889
Holometabolous Insects
722
33606
307348
Families
Genera
Species
 
Taxom
 

Order
 
Coleoptera
Linnaeus, 1758
Beetles, Coleopterans
193
13338
138720
Families
Genera
Species
 
Taxom
 

Suborder
 
Polyphaga
Emery, 1886
Polyphags
157
11884
117601
Families
Genera
Species
 
Taxom
 

Infraorder
 
Scarabaeiformia
Crowson, 1960
18
1318
18244
Families
Genera
Species
 
Taxom
 

Superfamily
 
Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles
18
1318
18244
Families
Genera
Species
 
Taxom
 
FamilyPassalidae
Leach, 1815
Bess beetle, Bess beetles, Betsy beetles, Passalid beetles
31
317
Genera
Species
 
Taxom
 


Genera for Asia
31 Genera with 317 Species
#
A
B
C
D
E
G
L
M
O
P
S
T

Aceraius Kaup, 189848SpeciesTaxom
Aulacocyclus Kaup, 193510SpeciesTaxom
Basilianus Kaup, 199128SpeciesTaxom
Cacoius Boucher, 19931SpeciesTaxom
Ceracupes Kaup, 19885SpeciesTaxom
Ceracyclus Boucher & Bai, 20172SpeciesTaxom
Cetejus Kaup, 18688SpeciesTaxom
Comacupes Kaup, 191413SpeciesTaxom
Cylindrocaulus Fairmaire, 18835SpeciesTaxom
Didimus Kaup, 18433SpeciesTaxom
Episphenus Kaup, 18715SpeciesTaxom
Gonatas Kaup, 19189SpeciesTaxom
Labienus Kaup, 18687SpeciesTaxom
Leptaulax Kaup, 199590SpeciesTaxom
Macrolinus Kaup, 186826SpeciesTaxom
Mastachilus Kaup, 18411SpeciesTaxom
Mastochilus Kaup, 18411SpeciesTaxom
Ophrygonius Zang, 199130SpeciesTaxom
Paratiberioides Iwase, 19945SpeciesTaxom
Passalus Fabricius, 18233SpeciesTaxom
Pelopides Kuwert, 182310SpeciesTaxom
Pentalobus Kaup, 18442SpeciesTaxom
Plesthenus Kaup, 19039SpeciesTaxom
Pleurarius Kaup, 18682SpeciesTaxom
Proculus Kaup, 18681SpeciesTaxom
Protomocoelus Zang, 19045SpeciesTaxom
Serrulus Hong, 19831SpeciesTaxom
Stephanocephalus Kaup, 18731SpeciesTaxom
Taeniocerus Kaup, 18685SpeciesTaxom
Tiberioides Gravely, 19075SpeciesTaxom
Trichostigmus Kaup, 19926SpeciesTaxom


Species list for Asia
317 Species and 19 Subspecies
Aceraius aequalis Gravely, 1918
Aceraius alpinus Kon, Ueda & ..., 1995
Aceraius alutaceosternus Kuwert, 1898
Aceraius ashidai (Kon, Araya &..., 1992)⇒ Aceraius kazuhisai Kon, Johki &..., 2003
Aceraius borneanus (Kaup, 1871)⇒ Aceraius hikidai Kon, Ueda & ..., 1993
Aceraius boucheri Kon, Araya &..., 1993
Aceraius cantori (Kaup, 1868)⇒ Basilianus indicus Stoliczka, 1873
Aceraius chinensis (Kuwert, 1898)⇒ Aceraius grandis Burmeister, 1847
Aceraius comptoni Kaup, 1868
Aceraius fujiokae Iwase, 1994
Aceraius grandis Burmeister, 1847
Aceraius hassani Kon, Araya &..., 2006
Aceraius helferi Kuwert, 1891
Aceraius hidakai Kon, Araya &..., 1993
Aceraius hikidai Kon, Ueda & ..., 1993
Aceraius himalayensis (Gravely, 1914)⇒ Aceraius helferi Kuwert, 1891
Aceraius hirsutus Kuwert, 1891
Aceraius hirtimarginalis (Iwase, 1993)⇒ Basilianus hirtimarginalis Iwase, 1993
Aceraius illegalis Kuwert, 1891
Aceraius jenisi Iwase, 1998
Aceraius katsurae Iwase, 1995
Aceraius kazuhisai Kon, Johki &..., 2003
Aceraius kikutai Kon, Johki &..., 1995
Aceraius kinabalensis Kon & Johki, 1989
Aceraius kuwerti Zang, 1903
Aceraius laevicollis Illiger, 1800
Aceraius laevimargo Zang, 1905
Aceraius lamellatus Gravely, 1918
Aceraius laniger Zang, 1905
Aceraius luzonicus (Kuwert, 1898)⇒ Aceraius laevicollis Illiger, 1800
Aceraius masatakai Kon, Johki &..., 2003
Aceraius meyeri (Kuwert, 1891)⇒ Aceraius laevicollis Illiger, 1800
Aceraius moescheri Kuwert, 1891
Aceraius moeschleri (Kon, Araya &..., 1992)⇒ Aceraius boucheri Kon, Araya &..., 1993
Aceraius neelgheriensis (Kaup, 1868)⇒ Basilianus indicus Stoliczka, 1873
Aceraius oculidens Zang, 1905
Aceraius palawanus Zang, 1905
Aceraius parvulus (Hincks, 1938)⇒ Basilianus minor Gravely, 1914
Aceraius pilifer (Percheron, 1835)⇒ Aceraius helferi Kuwert, 1891
Aceraius riekoae Iwase, 1998
Aceraius sabanus Kon, Ueda & ..., 1995
Aceraius sawaii Kon, Araya &..., 2006
Aceraius sipolae Iwase, 1998
Aceraius symmetricus (Iwase, 1995)⇒ Basilianus koni Boucher, 1995
Aceraius tobae Iwase, 1994
Aceraius tricornis Zang, 1903
Aceraius wallacei Kuwert, 1898
Aulacocyclus andrewesi Gravely, 1914
Aulacocyclus andrewsi Gravely, 1914
Aulacocyclus celebensis Heller, 1899
Aulacocyclus celebesiensis Heller, 1898
Aulacocyclus felderi Stolickza, 1873
Aulacocyclus glabriusculus Kuwert, 1891
Aulacocyclus laevipennis Doesburg, 1945
Aulacocyclus papuanus Heller, 1910
Aulacocyclus perlatus Kaup, 1868
Aulacocyclus zangi Dibb, 1935
Basilianus aequalis Gravely, 1918
Basilianus aequidens Gravely, 1914
Basilianus birmanicus Gravely, 1914
Basilianus bleuzeni Boucher, 1993
Basilianus cantori Percheron, 1844
Basilianus chinensis Endrodi, 1955
Basilianus convexifrons Zang, 1904
Basilianus emas Iwase, 1998
Basilianus ferreri Boucher, 1993
Basilianus griveaui Boucher, 1993
Basilianus hirtimarginalis Iwase, 1993
Basilianus indicus Stoliczka, 1873
Basilianus inopinus Boucher, 1997
Basilianus jacquesi Boucher, 1993
Basilianus javensis Gravely, 1918
Basilianus koni Boucher, 1995
Basilianus lacroixi Boucher, 1993
Basilianus longiruga Boucher, 1993
Basilianus megatarsi Boucher & Kon, 1999
Basilianus minor Gravely, 1914
Basilianus pseudaequidens Boucher, 1995
Basilianus rajanus Doesburg, 1941
Basilianus singapurae Gravely, 1914
Basilianus sternibarbis Boucher, 1995
Basilianus tanae Boucher, 1993
Basilianus tuberculosus Boucher & Kon, 1999
Basilianus tuberculus Boucher, 1995
Basilianus uedai Kon & Johki, 1991
Cacoius jacquesi (Boucher, 1993)⇒ Basilianus jacquesi Boucher, 1993
Ceracupes arrowi Heller, 1910
Ceracupes chingkini Okano, 1988
Ceracupes fronticornis Westwood, 1842
Ceracupes yui Okano, 1988
Ceracyclus jirouxi Boucher, Bai..., 2017
Ceracyclus lotus Boucher, Bai..., 2017
Cetejus gracilis Stoliczka, 1873
Cetejus halmaheirae Kaup, 1871
Cetejus laevigatus Zang, 1903
Cetejus peltostictus Kaup, 1868
Cetejus peltosticus Kaup, 1868
Cetejus pseudamericanus Boucher, 1995
Cetejus sodalis Kaup, 1868
Comacupes basalis Smith, 1852
Comacupes cavicornis Kaup, 1868
Comacupes cylindraceus (Perty, 1831)⇒ Comacupes masoni Stoliczka, 1873
Comacupes foveicollis Kuwert, 1891
Comacupes foveicollis foveicollis
Comacupes foveicollis minor Heller, 1896
Comacupes intermedius Dibb, 1937
Comacupes kaupi Boucher, 2004
Comacupes masoni Stoliczka, 1873
Comacupes meridialis Boucher, 2015
Comacupes minor (Heller, 1897)⇒ Comacupes foveicollis minor Heller, 1896
Comacupes palawanicus Boucher, 2015
Comacupes stoliczkae Gravely, 1914
Cylindrocaulus bucerus Fairmaire, 1880
Cylindrocaulus davidi Boucher & Ca..., 1996
Cylindrocaulus mishmi Kon, Araya &..., 2015
Cylindrocaulus patalis Lewis, 1883
Didimus africanus Percheron, 1844
Didimus knutsoni Aurivillius, 1886
Didimus parastictus (Imhoff, 1843)⇒ Didimus haroldi Kuwert, 1898
Episphenus annamensis Gravely, 1919
Episphenus comptoni Kaup, 1868
Episphenus flachi Kuwert, 1891
Episphenus moorei Kaup, 1871
Gonatas brevis (Kuwert, 1898)⇒ Gonatas germari Kaup, 1868
Gonatas germari Kaup, 1868
Gonatas minimus Kuwert, 1891
Gonatas minor Gravely, 1918
Gonatas naviculator Percheron, 1844
Gonatas odiosus Kuwert, 1891
Gonatas pumilio Kaup, 1868
Gonatas schellongi Kuwert, 1891
Gonatas tenimbrensis Gravely, 1918
Labienus compergus Boisduval, 1835
Labienus crassus Kaup, 1868
Labienus elisae Kuwert, 1898
Labienus gigas Kaup, 1868
Labienus inaequalis Gravely, 1918
Labienus moluccanus Percheron, 1835
Labienus ptox Kaup, 1868
Leptaulax acutangularis Iwase, 1998
Leptaulax anaulax Zang, 1905
Leptaulax anibarbis Kuwert, 1898
Leptaulax anna Zang, 1905
Leptaulax apicalis Iwase, 1996
Leptaulax arayai Johki & Kon, 2003
Leptaulax arrowi Hincks, 1933
Leptaulax aurivillii (Kuwert, 1891)⇒ Leptaulax differentispina Kuwert, 1891
Leptaulax barbicauda Zang, 1905
Leptaulax batschianae Kuwert, 1891
Leptaulax beccarii Kuwert, 1891
Leptaulax bicolor (Fabricius, 1801)⇒ Leptaulax dentatus Fabricius, 1792
Leptaulax bicolor bicolor
Leptaulax castilloae Kon, Johki &..., 2005
Leptaulax circulus Iwase, 1996
Leptaulax consequens Kuwert, 1898
Leptaulax crockerensis Iwase, 1998
Leptaulax cyclotaenius Kuwert, 1891
Leptaulax datar Iwase, 1996
Leptaulax dentatus Fabricius, 1792
Leptaulax dentifrons Iwase, 1996
Leptaulax differentispina Kuwert, 1891
Leptaulax elegans Iwase, 1996
Leptaulax eschscholtzi Kaup, 1868
Leptaulax formosanus Doesburg, 1992
Leptaulax fujiokae Iwase, 1995
Leptaulax hamidi Kon, Araya &..., 2021
Leptaulax hidakai Johki, Araya..., 2003
Leptaulax himalayae (Kuwert, 1898)⇒ Leptaulax cyclotaenius Kuwert, 1891
Leptaulax humerosus (Kuwert, 1891)⇒ Leptaulax dentatus Fabricius, 1792
Leptaulax immarginalis Iwase, 1996
Leptaulax immarginalis immarginalis immarginalis..., 1996
Leptaulax immarginalis minangkabauensis Iwase, 1997
Leptaulax intermedius Iwase, 1996
Leptaulax iwasei Johki, Araya..., 2007
Leptaulax jaechi Iwase, 1998
Leptaulax jenisi Iwase, 1998
Leptaulax karoensis Iwase, 1996
Leptaulax kecil Iwase, 1996
Leptaulax koreanus Nomura, Kon,..., 1993
Leptaulax labyrinthus Iwase, 1998
Leptaulax laevis Iwase, 1998
Leptaulax loebli Kon, Johki &..., 2003
Leptaulax luzonicus Iwase, 1996
Leptaulax macassariensis (Kuwert, 1898)⇒ Leptaulax differentispina Kuwert, 1891
Leptaulax malaccae Kuwert, 1891
Leptaulax manillae Kuwert, 1891
Leptaulax manis Iwase, 1996
Leptaulax masatakai Johki, Araya..., 2003
Leptaulax masayukii Iwase, 1997
Leptaulax masumotoi Johki, Araya..., 2011
Leptaulax matsumotoi Kon, Johki &..., 2003
Leptaulax matsumotoi kangfangensis Kon & Abe, 2009
Leptaulax matsumotoi matsumotoi
Leptaulax mindanaoensis Iwase, 1996
Leptaulax negrosensis Iwase, 1996
Leptaulax niae Kuwert, 1898
Leptaulax niisatoi Kon, Johki &..., 2017
Leptaulax pacholatkoi Iwase, 1998
Leptaulax pacholatkoi mutoniatus Neumann & Kon, 2013
Leptaulax pacholatkoi pacholatkoi
Leptaulax palawanicus Zang, 1905
Leptaulax parvus Iwase, 1995
Leptaulax parvus balita Iwase, 1997
Leptaulax parvus parvus balita Iwase, 1998
Leptaulax planus (Illiger, 1800)⇒ Leptaulax roepstorffi Kuwert, 1898
Leptaulax punctatus Iwase, 1995
Leptaulax riekoae Iwase, 1998
Leptaulax riekoae kachinensis Kon, Araya &..., 2005
Leptaulax riekoae riekoae kachinensis ..., 2005
Leptaulax roepstorffi Kuwert, 1898
Leptaulax roepstorfi Kuwert, 1898
Leptaulax sabahensis Iwase, 1996
Leptaulax sakaii Iwase, 1996
Leptaulax sambawae Gravely, 1918
Leptaulax sarawakus Iwase, 1996
Leptaulax sarawakus kashizakii Johki, Araya..., 2015
Leptaulax sarawakus sarawakus
Leptaulax sawaii Johki, Araya..., 2015
Leptaulax schillhammeri Iwase, 1998
Leptaulax schoedli Iwase, 1998
Leptaulax separandus Kuwert, 1891
Leptaulax shunichii Kon, Johki &..., 2021
Leptaulax similis Iwase, 1996
Leptaulax strbai Iwase, 1998
Leptaulax tamijii Kon, Araya &..., 2021
Leptaulax timoriensis Percheron, 1841
Leptaulax trajae Iwase, 1998
Leptaulax uenoi (Iwase, 1995)⇒ Leptaulax arrowi Hincks, 1933
Macrolinus andamanensis Stoliczka, 1873
Macrolinus batesi (Kuwert, 1898)⇒ Macrolinus cartereti Boucher, 1996
Macrolinus cartereti Boucher, 1996
Macrolinus crenatipennis Kuwert, 1898
Macrolinus depressus Gravely, 1918
Macrolinus duivenbodei Kaup, 1868
Macrolinus foveolatus Ma, 1988
Macrolinus johkii Boucher & Kon, 1998
Macrolinus latipennis Percheron, 1841
Macrolinus medogensis Zang, 1981
Macrolinus mineti Boucher, 1996
Macrolinus nicobaricus Gravely, 1914
Macrolinus obesus Gravely, 1918
Macrolinus puncticollis Hincks, 1938
Macrolinus punctipectus Hincks, 1838
Macrolinus rotundifrons Kaup, 1871
Macrolinus salibabuensis Boucher, 1996
Macrolinus sikkimensis Stoliczka, 1873
Macrolinus sikkimensis sikkimensis sikkimensis ..., 1873
Macrolinus sikkimensis tavoyanus Gravely, 1914
Macrolinus sinicus Hong, 1984
Macrolinus sulciperfectus Kuwert, 1891
Macrolinus urus Heller, 1898
Macrolinus waterhousei Kaup, 1871
Macrolinus weberi (Kaup, 1868)⇒ Macrolinus latipennis Percheron, 1841
Mastachilus australasicus Percheron, 1841
Mastochilus australasicus Percheron, 1841
Ophrygonius aequidens Gravely, 1914
Ophrygonius birmanicus (Gravely, 1914)⇒ Basilianus birmanicus Gravely, 1914
Ophrygonius bleuzeni (Boucher, 1993)⇒ Basilianus bleuzeni Boucher, 1993
Ophrygonius boxer Boucher, 1993
Ophrygonius cantori Percheron, 1844
Ophrygonius chinensis (Endrodi, 1955)⇒ Basilianus chinensis Endrodi, 1955
Ophrygonius convexifrons Zang, 1904
Ophrygonius emas Iwase, 1998
Ophrygonius ferreri (Boucher, 1993)⇒ Basilianus ferreri Boucher, 1993
Ophrygonius griveaui (Boucher, 1993)⇒ Basilianus griveaui Boucher, 1993
Ophrygonius hirtimarginalis Iwase, 1993
Ophrygonius inaequalis Burmeister, 1847
Ophrygonius indicus (Stoliczka, 1873)⇒ Basilianus indicus Stoliczka, 1873
Ophrygonius inopinus (Boucher, 1997)⇒ Basilianus inopinus Boucher, 1997
Ophrygonius javensis (Gravely, 1918)⇒ Basilianus javensis Gravely, 1918
Ophrygonius koni (Boucher, 1995)⇒ Basilianus koni Boucher, 1995
Ophrygonius lacroixi (Boucher, 1993)⇒ Basilianus lacroixi Boucher, 1993
Ophrygonius longiruga (Boucher, 1993)⇒ Basilianus longiruga Boucher, 1993
Ophrygonius megatarsi (Boucher & Kon, 1999)⇒ Basilianus megatarsi Boucher & Kon, 1999
Ophrygonius minor Gravely, 1914
Ophrygonius neelgherriensis Percheron, 1841
Ophrygonius planus Iwase, 1995
Ophrygonius pseudaequidens (Boucher, 1995)⇒ Basilianus pseudaequidens Boucher, 1995
Ophrygonius rajanus (Doesburg, 1941)⇒ Basilianus rajanus Doesburg, 1941
Ophrygonius singapurae (Gravely, 1914)⇒ Basilianus singapurae Gravely, 1914
Ophrygonius sternibarbis (Boucher, 1995)⇒ Basilianus sternibarbis Boucher, 1995
Ophrygonius tanae (Boucher, 1993)⇒ Basilianus tanae Boucher, 1993
Ophrygonius tuberculosus (Boucher & Kon, 1999)⇒ Basilianus tuberculosus Boucher & Kon, 1999
Ophrygonius tuberculus Boucher, 1995
Ophrygonius uedai (Kon & Johki, 1991)⇒ Basilianus uedai Kon & Johki, 1991
Paratiberioides laminatus Hincks, 1938
Paratiberioides mandibularis Heller, 1900
Paratiberioides mirabilis Iwase, 1994
Paratiberioides similis Iwase, 1995
Paratiberioides trajae Iwase, 1994
Passalus basalis (Smith, 1852)⇒ Comacupes basalis Smith, 1852
Passalus sinuatus Eschscholtz, 1829
Passalus tridens (Wiedemann, 1823)⇒ Pelopides tridens Wiedemann, 1823
Pelopides burmeisteri Kaup, 1868
Pelopides dorsalis Kaup, 1871
Pelopides gravidus Kuwert, 1891
Pelopides mniszechi Kaup, 1868
Pelopides monticulosus Smith, 1852
Pelopides pedroi Kon, Johki &..., 2005
Pelopides simplex Gravely, 1914
Pelopides symmetricus Zang, 1904
Pelopides tridens Wiedemann, 1823
Pentalobus barbatus Fabricius, 1801
Pentalobus palinii Percheron, 1844
Plesthenus anneae Boucher, 1996
Plesthenus gelon Schaufuss, 1885
Plesthenus invitus Kuwert, 1891
Plesthenus lottini Boisduval, 1835
Plesthenus lottini gelon Schaufuss, 1885
Plesthenus lottini lottini
Plesthenus sanghiensis Boucher, 1996
Plesthenus scutellopunctatus Zang, 1903
Pleurarius brachyphyllus Stoliczka, 1873
Pleurarius pilipes Kaup, 1868
Proculus mniszechi Kaup, 1868
Protomocoelus australis Boisduval, 1835
Protomocoelus heynei Kuwert, 1891
Protomocoelus otiosus Kuwert, 1898
Protomocoelus triumphator Zang, 1904
Serrulus sinicus Hong, 1983
Stephanocephalus redtenbacheri Stoliczka, 1873
Taeniocerus bicanthatus Percheron, 1841
Taeniocerus bicuspis Kaup, 1868
Taeniocerus mourzinei Boucher, 1998
Taeniocerus platypus Kaup, 1868
Taeniocerus pygmaeus Kaup, 1868
Tiberioides austeni Gravely, 1914
Tiberioides borealis Arrow, 1907
Tiberioides kerleyi Kon & Araya, 2007
Tiberioides kuwerti Arrow, 1907
Trichostigmus mindorensis Boucher, 1992
Trichostigmus mindoroensis Boucher, 1992
Trichostigmus semperi Boucher, 1992
Trichostigmus thoreyi Kaup, 1868
Trichostigmus ursulus Schaufuss, 1885
Trichostigmus whiteheadi Boucher, 1992

Subnodes of Passalidae with occurrences in Asia
SubfamilyAulacocyclinae
Kaup, 1868
5
34
Genera
Species
 
Taxom
 
SubfamilyPassalinae
Leach, 1815
24
280
Genera
Species
 
Taxom
 


Links, sources and further information