Checklist of Family Passalidae Leach, 1815 (bess beetle)
for Malaysia (Southeast Asia)
Class
| Insecta
Linnaeus, 1758
Insects, True insects | 562
8004
25384 | Families
Genera
Species |
Taxom
|
Subclass
| Pterygota
Lang, 1888
Winged insects | 558
7999
25379 | Families
Genera
Species |
Taxom
|
Infraclass
| Neoptera
van der Wulp, 1890
Modern Wing-folding insects | 535
7864
25001 | Families
Genera
Species |
Taxom
|
Superorder
| Holometabola
Heider, 1889
Holometabolous Insects | 321
5806
19788 | Families
Genera
Species |
Taxom
|
Order
| Coleoptera
Linnaeus, 1758
Beetles, Coleopterans | 98
2563
9929 | Families
Genera
Species |
Taxom
|
Suborder
| Polyphaga
Emery, 1886
Polyphags | 93
2366
9279 | Families
Genera
Species |
Taxom
|
Infraorder
| Scarabaeiformia
Crowson, 1960
| 10
308
1785 | Families
Genera
Species |
Taxom
|
Superfamily
| Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles | 10
308
1785 | Families
Genera
Species |
Taxom
| | Family | Passalidae
Leach, 1815
Bess beetle, Bess beetles, Betsy beetles, Passalid beetles | 11
115 | Genera
Species |
Taxom
|
Genera for Malaysia
11 Genera with 115 Species | Aceraius Kaup, 1898 | | 34 | Species | Taxom | | Basilianus Kaup, 1991 | | 13 | Species | Taxom | | Ceracupes Kaup, 1842 | | 1 | Species | Taxom | | Comacupes Kaup, 1914 | | 4 | Species | Taxom | | Didimus Kaup, 1844 | | 1 | Species | Taxom | | Gonatas Kaup, 1918 | | 1 | Species | Taxom | | Leptaulax Kaup, 2021 | | 37 | Species | Taxom | | Macrolinus Kaup, 1841 | | 5 | Species | Taxom | | Ophrygonius Zang, 1991 | | 16 | Species | Taxom | | Pelopides Kuwert, 1904 | | 6 | Species | Taxom | | Taeniocerus Kaup, 1868 | | 4 | Species | Taxom |
Species list for Malaysia
115 Species and 4 Subspecies | Aceraius alpinus Kon, Ueda & ..., 1995 | | | Aceraius alutaceosternus Kuwert, 1898 | | | Aceraius ashidai (Kon, Araya &..., 1992) | ⇒ Aceraius kazuhisai Kon, Johki &..., 2003 | | Aceraius borneanus (Kaup, 1871) | ⇒ Aceraius hikidai Kon, Ueda & ..., 1993 | | Aceraius boucheri Kon, Araya &..., 1993 | | | Aceraius fujiokae Iwase, 1994 | | | Aceraius grandis Burmeister, 1847 | | | Aceraius hassani Kon, Araya &..., 2006 | | | Aceraius helferi Kuwert, 1891 | | | Aceraius hidakai Kon, Araya &..., 1993 | | | Aceraius hikidai Kon, Ueda & ..., 1993 | | | Aceraius hirtimarginalis (Iwase, 1993) | ⇒ Basilianus hirtimarginalis Iwase, 1993 | | Aceraius illegalis Kuwert, 1891 | | | Aceraius jenisi Iwase, 1998 | | | Aceraius katsurae Iwase, 1995 | | | Aceraius kazuhisai Kon, Johki &..., 2003 | | | Aceraius kikutai Kon, Johki &..., 1995 | | | Aceraius kinabalensis Kon & Johki, 1989 | | | Aceraius kuwerti Zang, 1903 | | | Aceraius laevicollis Illiger, 1800 | | | Aceraius lamellatus Gravely, 1918 | | | Aceraius laniger Zang, 1905 | | | Aceraius masatakai Kon, Johki &..., 2003 | | | Aceraius moeschleri (Kon, Araya &..., 1992) | ⇒ Aceraius boucheri Kon, Araya &..., 1993 | | Aceraius oculidens Zang, 1905 | | | Aceraius parvulus (Hincks, 1938) | ⇒ Basilianus minor Gravely, 1914 | | Aceraius pilifer (Percheron, 1835) | ⇒ Aceraius helferi Kuwert, 1891 | | Aceraius riekoae Iwase, 1998 | | | Aceraius sabanus Kon, Ueda & ..., 1995 | | | Aceraius sawaii Kon, Araya &..., 2006 | | | Aceraius symmetricus (Iwase, 1995) | ⇒ Basilianus koni Boucher, 1995 | | Aceraius tricornis Zang, 1903 | | | Aceraius wallacei Kuwert, 1898 | | | Basilianus aequidens Gravely, 1914 | | | Basilianus bleuzeni Boucher, 1993 | | | Basilianus emas Iwase, 1998 | | | Basilianus ferreri Boucher, 1993 | | | Basilianus hirtimarginalis Iwase, 1993 | | | Basilianus koni Boucher, 1995 | | | Basilianus minor Gravely, 1914 | | | Basilianus pseudaequidens Boucher, 1995 | | | Basilianus rajanus Doesburg, 1941 | | | Basilianus singapurae Gravely, 1914 | | | Basilianus sternibarbis Boucher, 1995 | | | Basilianus tuberculosus Boucher & Kon, 1999 | | | Basilianus uedai Kon & Johki, 1991 | | | Ceracupes fronticornis Westwood, 1842 | | | Comacupes cavicornis Kaup, 1868 | | | Comacupes cylindraceus (Perty, 1831) | ⇒ Comacupes masoni Stoliczka, 1873 | | Comacupes masoni Stoliczka, 1873 | | | Comacupes stoliczkae Gravely, 1914 | | | Didimus africanus Percheron, 1844 | | | Gonatas minor Gravely, 1918 | | | Leptaulax acutangularis Iwase, 1998 | | | Leptaulax anibarbis Kuwert, 1898 | | | Leptaulax apicalis Iwase, 1996 | | | Leptaulax arayai Johki & Kon, 2003 | | | Leptaulax barbicauda Zang, 1905 | | | Leptaulax beccarii Kuwert, 1891 | | | Leptaulax bicolor (Fabricius, 1801) | ⇒ Leptaulax dentatus Fabricius, 1792 | | Leptaulax bicolor bicolor | | | Leptaulax castilloae Kon, Johki &..., 2005 | | | Leptaulax circulus Iwase, 1996 | | | Leptaulax crockerensis Iwase, 1998 | | | Leptaulax cyclotaenius Kuwert, 1891 | | | Leptaulax datar Iwase, 1996 | | | Leptaulax dentatus Fabricius, 1792 | | | Leptaulax dentifrons Iwase, 1996 | | | Leptaulax eschscholtzi Kaup, 1868 | | | Leptaulax hamidi Kon, Araya &..., 2021 | | | Leptaulax hidakai Johki, Araya..., 2003 | | | Leptaulax humerosus (Kuwert, 1891) | ⇒ Leptaulax dentatus Fabricius, 1792 | | Leptaulax jenisi Iwase, 1998 | | | Leptaulax malaccae Kuwert, 1891 | | | Leptaulax manis Iwase, 1996 | | | Leptaulax masumotoi Johki, Araya..., 2011 | | | Leptaulax parvus Iwase, 1995 | | | Leptaulax parvus parvus balita Iwase, 1998 | | | Leptaulax planus (Illiger, 1800) | ⇒ Leptaulax roepstorffi Kuwert, 1898 | | Leptaulax roepstorffi Kuwert, 1898 | | | Leptaulax sabahensis Iwase, 1996 | | | Leptaulax sarawakus Iwase, 1996 | | | Leptaulax sarawakus kashizakii Johki, Araya..., 2015 | | | Leptaulax sarawakus sarawakus | | | Leptaulax sawaii Johki, Araya..., 2015 | | | Leptaulax schillhammeri Iwase, 1998 | | | Leptaulax shunichii Kon, Johki &..., 2021 | | | Leptaulax similis Iwase, 1996 | | | Leptaulax strbai Iwase, 1998 | | | Leptaulax tamijii Kon, Araya &..., 2021 | | | Macrolinus batesi (Kuwert, 1898) | ⇒ Macrolinus cartereti Boucher, 1996 | | Macrolinus cartereti Boucher, 1996 | | | Macrolinus johkii Boucher & Kon, 1998 | | | Macrolinus latipennis Percheron, 1841 | | | Ophrygonius aequidens Gravely, 1914 | | | Ophrygonius bleuzeni (Boucher, 1993) | ⇒ Basilianus bleuzeni Boucher, 1993 | | Ophrygonius emas Iwase, 1998 | | | Ophrygonius ferreri (Boucher, 1993) | ⇒ Basilianus ferreri Boucher, 1993 | | Ophrygonius hirtimarginalis Iwase, 1993 | | | Ophrygonius inaequalis Burmeister, 1847 | | | Ophrygonius koni (Boucher, 1995) | ⇒ Basilianus koni Boucher, 1995 | | Ophrygonius minor Gravely, 1914 | | | Ophrygonius planus Iwase, 1995 | | | Ophrygonius pseudaequidens (Boucher, 1995) | ⇒ Basilianus pseudaequidens Boucher, 1995 | | Ophrygonius rajanus (Doesburg, 1941) | ⇒ Basilianus rajanus Doesburg, 1941 | | Ophrygonius singapurae (Gravely, 1914) | ⇒ Basilianus singapurae Gravely, 1914 | | Ophrygonius sternibarbis (Boucher, 1995) | ⇒ Basilianus sternibarbis Boucher, 1995 | | Ophrygonius tuberculosus (Boucher & Kon, 1999) | ⇒ Basilianus tuberculosus Boucher & Kon, 1999 | | Ophrygonius tuberculus Boucher, 1995 | | | Ophrygonius uedai (Kon & Johki, 1991) | ⇒ Basilianus uedai Kon & Johki, 1991 | | Pelopides dorsalis Kaup, 1871 | | | Pelopides monticulosus Smith, 1852 | | | Pelopides pedroi Kon, Johki &..., 2005 | | | Pelopides simplex Gravely, 1914 | | | Pelopides symmetricus Zang, 1904 | | | Taeniocerus bicanthatus Percheron, 1841 | | | Taeniocerus bicuspis Kaup, 1868 | | | Taeniocerus platypus Kaup, 1868 | | | Taeniocerus pygmaeus Kaup, 1868 | |
Subnodes of Passalidae with occurrences in Malaysia
Links, sources and further information
|
