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Checklist for Family Passalidae Leach, 1815 (bess beetle)
and region Australasia (Oceania)


Class
 
Insecta
Linnaeus, 1758
Insects, True insects
942
23049
129157
Families
Genera
Species
 
Taxom
 

Subclass
 
Pterygota
Lang, 1888
Winged insects
938
23017
129086
Families
Genera
Species
 
Taxom
 

Infraclass
 
Neoptera
van der Wulp, 1890
Modern Wing-folding insects
883
22694
128014
Families
Genera
Species
 
Taxom
 

Superorder
 
Holometabola
Heider, 1889
Holometabolous Insects
534
17158
106564
Families
Genera
Species
 
Taxom
 

Order
 
Coleoptera
Linnaeus, 1758
Beetles, Coleopterans
144
6439
46908
Families
Genera
Species
 
Taxom
 

Suborder
 
Polyphaga
Emery, 1886
Polyphags
127
5615
39050
Families
Genera
Species
 
Taxom
 

Infraorder
 
Scarabaeiformia
Crowson, 1960
11
581
4660
Families
Genera
Species
 
Taxom
 

Superfamily
 
Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles
11
581
4660
Families
Genera
Species
 
Taxom
 
FamilyPassalidae
Leach, 1815
Bess beetle, Bess beetles, Betsy beetles, Passalid beetles
19
121
Genera
Species
 
Taxom
 


Genera for Australasia
19 Genera with 121 Species
Analaches Kuwert, 19103SpeciesTaxom
Arrox Zang, 1905
0SpeciesTaxom
Aulacocyclus Kaup, 193523SpeciesTaxom
Austropassalus Mjöberg, 19171SpeciesTaxom
Cetejus Kaup, 186813SpeciesTaxom
Gonatas Kaup, 18916SpeciesTaxom
Hincksius Boucher, 19933SpeciesTaxom
Labienus Kaup, 193820SpeciesTaxom
Leptaulax Kaup, 191817SpeciesTaxom
Mastachilus Kaup, 18915SpeciesTaxom
Mastochilus Kaup, 18926SpeciesTaxom
Passalus Fabricius, 1792
0SpeciesTaxom
Pharochilus Kaup, 18987SpeciesTaxom
Plesthenus Kaup, 18351SpeciesTaxom
Protomocoelus Zang, 19056SpeciesTaxom
Pseudepisphenus Gravely, 19909SpeciesTaxom
Publius Kaup, 1871
0SpeciesTaxom
Taeniocerus Kaup, 18681SpeciesTaxom
Tarquinius Kuwert, 18911SpeciesTaxom


Species list for Australasia
121 Species and 1 Subspecies
Analaches australiensis Stoliczka, 1873
Analaches bicavis Zang, 1905
Analaches dubius (Heller, 1910)⇒ Analaches bicavis Zang, 1905
Aulacocyclus aliicornis Kuwert, 1897
Aulacocyclus collaris Blackburn, 1896
Aulacocyclus deyrollei Kaup, 1868
Aulacocyclus deyrolli Kaup, 1868
Aulacocyclus edentulus W.S. Macleay, 1826Bess Beetle
Aulacocyclus errans (Blackburn, 1896)⇒ Aulacocyclus fracticornis Kuwert, 1891
Aulacocyclus fracticornis Kuwert, 1891
Aulacocyclus gravelyi Dibb, 1933
Aulacocyclus hangayi Doesburg, 1992
Aulacocyclus kaupi MacLeay, 1869
Aulacocyclus kaupii MacLeay, 1871
Aulacocyclus macleayi Kaup, 1871
Aulacocyclus mastersi MacLeay, 1871
Aulacocyclus mastersii MacLeay, 1871
Aulacocyclus papuanus Heller, 1910
Aulacocyclus perlatus Kaup, 1868
Aulacocyclus rosenbergi Kaup, 1868
Aulacocyclus rosenbergii Kaup, 1868
Aulacocyclus rotundatoclypeatus (Kuwert, 1891)⇒ Aulacocyclus fracticornis Kuwert, 1891
Aulacocyclus tambourinensis Mjoeberg, 1917
Aulacocyclus teres Percheron, 1841
Aulacocyclus yorkensis Reyes-Castil..., 2016
Aulacocyclus zangi Dibb, 1935
Austropassalus hultgreni Mjoeberg, 1917
Cetejus acutangulus Heller, 1910
Cetejus bretoni Boucher, 1993
Cetejus convexus Doesburg, 1956
Cetejus glabriventris Kuwert, 1898
Cetejus grabowskyi Kuwert, 1891
Cetejus imbecillis Kuwert, 1898
Cetejus innovator Boucher, 1993
Cetejus lateralis Hincks, 1938
Cetejus longicornis Doesburg, 1956
Cetejus obliquus Kirsche, 1877
Cetejus paradoxus Hincks, 1937
Cetejus schenklingi Heller, 1910
Cetejus sodalis Kaup, 1868
Gonatas altidens Heller, 1910
Gonatas germari Kaup, 1868
Gonatas minimus Kuwert, 1891
Gonatas naviculator Percheron, 1844
Gonatas pumilio Kaup, 1868
Gonatas schellongi Kuwert, 1891
Hincksius macrolinoides Boucher, 1993
Hincksius papuanus Hincks, 1956
Hincksius praestabilis Boucher, 1993
Labienus aberrans Hincks, 1938
Labienus cavifrons Hincks, 1935
Labienus cheesmanae Hincks, 1935
Labienus compergus Boisduval, 1835
Labienus dohrni Kuwert, 1891
Labienus glaber Gravely, 1913
Labienus glabriceps Doesburg, 1956
Labienus gracilis Heller, 1910
Labienus impar Kuwert, 1898
Labienus inaequalis Gravely, 1918
Labienus kjellanderi Endrodi, 1967
Labienus moluccanus Percheron, 1835
Labienus opalus Boucher, 1993
Labienus protomocoeloides Hincks, 1933
Labienus proximus Hincks, 1933
Labienus ptox Kaup, 1868
Labienus ptoxoides Gravely, 1918
Labienus toxopeusi Doesburg, 1956
Labienus trigonophorus Zang, 1905
Labienus truncatus Hincks, 1938
Leptaulax analis Zang, 1906
Leptaulax anibarbis Kuwert, 1898
Leptaulax barbicauda Zang, 1905
Leptaulax bicolor (Fabricius, 1801)⇒ Leptaulax dentatus Fabricius, 1792
Leptaulax bicolor bicolor
Leptaulax biroi Endrodi, 1953
Leptaulax dentatus Fabricius, 1792
Leptaulax drescheri Doesburg, 1956
Leptaulax hansemani Kuwert, 1898
Leptaulax hirsutus Hincks, 1956
Leptaulax joliveti Endrodi, 1971
Leptaulax novaeguineae Kuwert, 1891
Leptaulax obtusidens Kuwert, 1891
Leptaulax papua (Kuwert, 1898)⇒ Leptaulax anibarbis Kuwert, 1898
Leptaulax papuanus Zang, 1906
Leptaulax pulchellus Arrow, 1907
Leptaulax sambawae Gravely, 1918
Mastachilus australasicus Percheron, 1841
Mastachilus lottinii Boisduval, 1835
Mastachilus parvifrons Kuwert, 1891
Mastachilus polyphyllus W. S. MacLeay, 1826
Mastachilus quaestionis Kuwert, 1891Bess beetle
Mastochilus australasicus Percheron, 1841
Mastochilus parvifrons Kuwert, 1891
Mastochilus pectiniger Heller, 1910
Mastochilus polyphyllus MacLeay, 1827
Mastochilus quaestionis Kuwert, 1891
Mastochilus subobliquus Tyron, 1892
Pharochilus dilatatus Dalman, 1817
Pharochilus nitidulus MacLeay, 1871
Pharochilus politus Burmeister, 1847Common Passalid Beetle
Pharochilus puncticollis MacLeay, 1871
Pharochilus punctiger Gravely, 1918
Pharochilus rugiceps Hope & Westwood, 1845Common passalid beetle
Pharochilus scutellonotus Kuwert, 1898
Plesthenus lottinii Boisduval, 1835
Protomocoelus australis Boisduval, 1835
Protomocoelus fallax Kuwert, 1898
Protomocoelus heynei Kuwert, 1891
Protomocoelus labrinotus Kuwert, 1891
Protomocoelus otiosus Kuwert, 1898
Protomocoelus sternbergi Zang, 1905
Pseudepisphenus forceps Boucher, 1993
Pseudepisphenus hodeberti Boucher, 1992
Pseudepisphenus humeribarbis Boucher, 1991
Pseudepisphenus longulus Boucher, 1990
Pseudepisphenus merkli Boucher, 1992
Pseudepisphenus morettoi Boucher, 1992
Pseudepisphenus perplexus Gravely, 1914
Pseudepisphenus proditor Boucher, 1990
Pseudepisphenus scitulus Boucher, 1990
Taeniocerus deyrollei Kaup, 1868
Tarquinius paradoxus Kuwert, 1891

Subnodes of Passalidae with occurrences in Australasia
SubfamilyAulacocyclinae
Kaup, 1868
2
24
Genera
Species

Taxom

SubfamilyPassalinae
Leach, 1815
16
92
Genera
Species

Taxom



Links, sources and further information
Passalidae at gbif.org
Passalidae at irmng.org
Passalidae at itis.gov