Checklist of Family Passalidae Leach, 1815 (bess beetle)
for Far East (Asia)
Class
| Insecta
Linnaeus, 1758
Insects, True insects | 1256
36222
255417 | Families
Genera
Species |
Taxom
|
Subclass
| Pterygota
Lang, 1888
Winged insects | 1250
36175
255323 | Families
Genera
Species |
Taxom
|
Infraclass
| Neoptera
van der Wulp, 1890
Modern Wing-folding insects | 1148
35562
252662 | Families
Genera
Species |
Taxom
|
Superorder
| Holometabola
Heider, 1889
Holometabolous Insects | 671
26426
204688 | Families
Genera
Species |
Taxom
|
Order
| Coleoptera
Linnaeus, 1758
Beetles, Coleopterans | 186
10806
97430 | Families
Genera
Species |
Taxom
|
Suborder
| Polyphaga
Emery, 1886
Polyphags | 152
9707
84101 | Families
Genera
Species |
Taxom
|
Infraorder
| Scarabaeiformia
Crowson, 1960
| 18
1070
13561 | Families
Genera
Species |
Taxom
|
Superfamily
| Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles | 18
1070
13561 | Families
Genera
Species |
Taxom
| | Family | Passalidae
Leach, 1815
Bess beetle, Bess beetles, Betsy beetles, Passalid beetles | 30
293 | Genera
Species |
Taxom
|
Genera for Far East
30 Genera with 293 Species
| Aceraius Kaup, 1898 | | 45 | Species | Taxom | | Aulacocyclus Kaup, 1935 | | 8 | Species | Taxom | | Basilianus Kaup, 1991 | | 28 | Species | Taxom | | Cacoius Boucher, 1993 | | 1 | Species | Taxom | | Ceracupes Kaup, 1988 | | 5 | Species | Taxom | | Ceracyclus Boucher & Bai, 2017 | | 2 | Species | Taxom | | Cetejus Kaup, 1868 | | 8 | Species | Taxom | | Comacupes Kaup, 1914 | | 12 | Species | Taxom | | Cylindrocaulus Fairmaire, 1883 | | 4 | Species | Taxom | | Didimus Kaup, 1843 | | 3 | Species | Taxom | | Episphenus Kaup, 1919 | | 1 | Species | Taxom | | Gonatas Kaup, 1918 | | 9 | Species | Taxom | | Labienus Kaup, 1868 | | 7 | Species | Taxom | | Leptaulax Kaup, 1995 | | 88 | Species | Taxom | | Macrolinus Kaup, 1868 | | 20 | Species | Taxom | | Mastachilus Kaup, 1841 | | 1 | Species | Taxom | | Mastochilus Kaup, 1841 | | 1 | Species | Taxom | | Ophrygonius Zang, 1991 | | 27 | Species | Taxom | | Paratiberioides Iwase, 1994 | | 5 | Species | Taxom | | Passalus Fabricius, 1823 | | 3 | Species | Taxom | | Pelopides Kuwert, 1823 | | 10 | Species | Taxom | | Pentalobus Kaup, 1844 | | 2 | Species | Taxom | | Plesthenus Kaup, 1903 | | 9 | Species | Taxom | | Pleurarius Kaup, 1868 | | 1 | Species | Taxom | | Proculus Kaup, 1868 | | 1 | Species | Taxom | | Protomocoelus Zang, 1904 | | 5 | Species | Taxom | | Serrulus Hong, 1983 | | 1 | Species | Taxom | | Taeniocerus Kaup, 1868 | | 5 | Species | Taxom | | Tiberioides Gravely, 1907 | | 4 | Species | Taxom | | Trichostigmus Kaup, 1992 | | 6 | Species | Taxom |
Species list for Far East
293 Species and 19 Subspecies | Aceraius aequalis Gravely, 1918 | | | Aceraius alpinus Kon, Ueda & ..., 1995 | | | Aceraius alutaceosternus Kuwert, 1898 | | | Aceraius ashidai (Kon, Araya &..., 1992) | ⇒ Aceraius kazuhisai Kon, Johki &..., 2003 | | Aceraius borneanus (Kaup, 1871) | ⇒ Aceraius hikidai Kon, Ueda & ..., 1993 | | Aceraius boucheri Kon, Araya &..., 1993 | | | Aceraius chinensis (Kuwert, 1898) | ⇒ Aceraius grandis Burmeister, 1847 | | Aceraius comptoni Kaup, 1868 | | | Aceraius fujiokae Iwase, 1994 | | | Aceraius grandis Burmeister, 1847 | | | Aceraius hassani Kon, Araya &..., 2006 | | | Aceraius helferi Kuwert, 1891 | | | Aceraius hidakai Kon, Araya &..., 1993 | | | Aceraius hikidai Kon, Ueda & ..., 1993 | | | Aceraius hirsutus Kuwert, 1891 | | | Aceraius hirtimarginalis (Iwase, 1993) | ⇒ Basilianus hirtimarginalis Iwase, 1993 | | Aceraius illegalis Kuwert, 1891 | | | Aceraius jenisi Iwase, 1998 | | | Aceraius katsurae Iwase, 1995 | | | Aceraius kazuhisai Kon, Johki &..., 2003 | | | Aceraius kikutai Kon, Johki &..., 1995 | | | Aceraius kinabalensis Kon & Johki, 1989 | | | Aceraius kuwerti Zang, 1903 | | | Aceraius laevicollis Illiger, 1800 | | | Aceraius laevimargo Zang, 1905 | | | Aceraius lamellatus Gravely, 1918 | | | Aceraius laniger Zang, 1905 | | | Aceraius luzonicus (Kuwert, 1898) | ⇒ Aceraius laevicollis Illiger, 1800 | | Aceraius masatakai Kon, Johki &..., 2003 | | | Aceraius moescheri Kuwert, 1891 | | | Aceraius moeschleri (Kon, Araya &..., 1992) | ⇒ Aceraius boucheri Kon, Araya &..., 1993 | | Aceraius neelgheriensis (Kaup, 1868) | ⇒ Basilianus indicus Stoliczka, 1873 | | Aceraius oculidens Zang, 1905 | | | Aceraius palawanus Zang, 1905 | | | Aceraius parvulus (Hincks, 1938) | ⇒ Basilianus minor Gravely, 1914 | | Aceraius pilifer (Percheron, 1835) | ⇒ Aceraius helferi Kuwert, 1891 | | Aceraius riekoae Iwase, 1998 | | | Aceraius sabanus Kon, Ueda & ..., 1995 | | | Aceraius sawaii Kon, Araya &..., 2006 | | | Aceraius sipolae Iwase, 1998 | | | Aceraius symmetricus (Iwase, 1995) | ⇒ Basilianus koni Boucher, 1995 | | Aceraius tobae Iwase, 1994 | | | Aceraius tricornis Zang, 1903 | | | Aceraius wallacei Kuwert, 1898 | | | Aulacocyclus celebensis Heller, 1899 | | | Aulacocyclus celebesiensis Heller, 1898 | | | Aulacocyclus felderi Stolickza, 1873 | | | Aulacocyclus glabriusculus Kuwert, 1891 | | | Aulacocyclus laevipennis Doesburg, 1945 | | | Aulacocyclus papuanus Heller, 1910 | | | Aulacocyclus perlatus Kaup, 1868 | | | Aulacocyclus zangi Dibb, 1935 | | | Basilianus aequalis Gravely, 1918 | | | Basilianus aequidens Gravely, 1914 | | | Basilianus birmanicus Gravely, 1914 | | | Basilianus bleuzeni Boucher, 1993 | | | Basilianus cantori Percheron, 1844 | | | Basilianus chinensis Endrodi, 1955 | | | Basilianus convexifrons Zang, 1904 | | | Basilianus emas Iwase, 1998 | | | Basilianus ferreri Boucher, 1993 | | | Basilianus griveaui Boucher, 1993 | | | Basilianus hirtimarginalis Iwase, 1993 | | | Basilianus indicus Stoliczka, 1873 | | | Basilianus inopinus Boucher, 1997 | | | Basilianus jacquesi Boucher, 1993 | | | Basilianus javensis Gravely, 1918 | | | Basilianus koni Boucher, 1995 | | | Basilianus lacroixi Boucher, 1993 | | | Basilianus longiruga Boucher, 1993 | | | Basilianus megatarsi Boucher & Kon, 1999 | | | Basilianus minor Gravely, 1914 | | | Basilianus pseudaequidens Boucher, 1995 | | | Basilianus rajanus Doesburg, 1941 | | | Basilianus singapurae Gravely, 1914 | | | Basilianus sternibarbis Boucher, 1995 | | | Basilianus tanae Boucher, 1993 | | | Basilianus tuberculosus Boucher & Kon, 1999 | | | Basilianus tuberculus Boucher, 1995 | | | Basilianus uedai Kon & Johki, 1991 | | | Cacoius jacquesi (Boucher, 1993) | ⇒ Basilianus jacquesi Boucher, 1993 | | Ceracupes arrowi Heller, 1910 | | | Ceracupes chingkini Okano, 1988 | | | Ceracupes fronticornis Westwood, 1842 | | | Ceracupes yui Okano, 1988 | | | Ceracyclus jirouxi Boucher, Bai..., 2017 | | | Ceracyclus lotus Boucher, Bai..., 2017 | | | Cetejus gracilis Stoliczka, 1873 | | | Cetejus halmaheirae Kaup, 1871 | | | Cetejus laevigatus Zang, 1903 | | | Cetejus peltostictus Kaup, 1868 | | | Cetejus peltosticus Kaup, 1868 | | | Cetejus pseudamericanus Boucher, 1995 | | | Cetejus sodalis Kaup, 1868 | | | Comacupes basalis Smith, 1852 | | | Comacupes cavicornis Kaup, 1868 | | | Comacupes cylindraceus (Perty, 1831) | ⇒ Comacupes masoni Stoliczka, 1873 | | Comacupes foveicollis Kuwert, 1891 | | | Comacupes foveicollis foveicollis | | | Comacupes foveicollis minor Heller, 1896 | | | Comacupes intermedius Dibb, 1937 | | | Comacupes masoni Stoliczka, 1873 | | | Comacupes meridialis Boucher, 2015 | | | Comacupes minor (Heller, 1897) | ⇒ Comacupes foveicollis minor Heller, 1896 | | Comacupes palawanicus Boucher, 2015 | | | Comacupes stoliczkae Gravely, 1914 | | | Cylindrocaulus bucerus Fairmaire, 1880 | | | Cylindrocaulus davidi Boucher & Ca..., 1996 | | | Cylindrocaulus patalis Lewis, 1883 | | | Didimus africanus Percheron, 1844 | | | Didimus knutsoni Aurivillius, 1886 | | | Didimus parastictus (Imhoff, 1843) | ⇒ Didimus haroldi Kuwert, 1898 | | Episphenus annamensis Gravely, 1919 | | | Gonatas brevis (Kuwert, 1898) | ⇒ Gonatas germari Kaup, 1868 | | Gonatas germari Kaup, 1868 | | | Gonatas minimus Kuwert, 1891 | | | Gonatas minor Gravely, 1918 | | | Gonatas naviculator Percheron, 1844 | | | Gonatas odiosus Kuwert, 1891 | | | Gonatas pumilio Kaup, 1868 | | | Gonatas schellongi Kuwert, 1891 | | | Gonatas tenimbrensis Gravely, 1918 | | | Labienus compergus Boisduval, 1835 | | | Labienus crassus Kaup, 1868 | | | Labienus elisae Kuwert, 1898 | | | Labienus gigas Kaup, 1868 | | | Labienus inaequalis Gravely, 1918 | | | Labienus moluccanus Percheron, 1835 | | | Labienus ptox Kaup, 1868 | | | Leptaulax acutangularis Iwase, 1998 | | | Leptaulax anaulax Zang, 1905 | | | Leptaulax anibarbis Kuwert, 1898 | | | Leptaulax anna Zang, 1905 | | | Leptaulax apicalis Iwase, 1996 | | | Leptaulax arayai Johki & Kon, 2003 | | | Leptaulax arrowi Hincks, 1933 | | | Leptaulax barbicauda Zang, 1905 | | | Leptaulax batschianae Kuwert, 1891 | | | Leptaulax beccarii Kuwert, 1891 | | | Leptaulax bicolor (Fabricius, 1801) | ⇒ Leptaulax dentatus Fabricius, 1792 | | Leptaulax bicolor bicolor | | | Leptaulax castilloae Kon, Johki &..., 2005 | | | Leptaulax circulus Iwase, 1996 | | | Leptaulax consequens Kuwert, 1898 | | | Leptaulax crockerensis Iwase, 1998 | | | Leptaulax cyclotaenius Kuwert, 1891 | | | Leptaulax datar Iwase, 1996 | | | Leptaulax dentatus Fabricius, 1792 | | | Leptaulax dentifrons Iwase, 1996 | | | Leptaulax differentispina Kuwert, 1891 | | | Leptaulax elegans Iwase, 1996 | | | Leptaulax eschscholtzi Kaup, 1868 | | | Leptaulax formosanus Doesburg, 1992 | | | Leptaulax fujiokae Iwase, 1995 | | | Leptaulax hamidi Kon, Araya &..., 2021 | | | Leptaulax hidakai Johki, Araya..., 2003 | | | Leptaulax himalayae (Kuwert, 1898) | ⇒ Leptaulax cyclotaenius Kuwert, 1891 | | Leptaulax humerosus (Kuwert, 1891) | ⇒ Leptaulax dentatus Fabricius, 1792 | | Leptaulax immarginalis Iwase, 1996 | | | Leptaulax immarginalis immarginalis immarginalis..., 1996 | | | Leptaulax immarginalis minangkabauensis Iwase, 1997 | | | Leptaulax intermedius Iwase, 1996 | | | Leptaulax iwasei Johki, Araya..., 2007 | | | Leptaulax jaechi Iwase, 1998 | | | Leptaulax jenisi Iwase, 1998 | | | Leptaulax karoensis Iwase, 1996 | | | Leptaulax kecil Iwase, 1996 | | | Leptaulax koreanus Nomura, Kon,..., 1993 | | | Leptaulax labyrinthus Iwase, 1998 | | | Leptaulax laevis Iwase, 1998 | | | Leptaulax loebli Kon, Johki &..., 2003 | | | Leptaulax luzonicus Iwase, 1996 | | | Leptaulax macassariensis (Kuwert, 1898) | ⇒ Leptaulax differentispina Kuwert, 1891 | | Leptaulax malaccae Kuwert, 1891 | | | Leptaulax manillae Kuwert, 1891 | | | Leptaulax manis Iwase, 1996 | | | Leptaulax masatakai Johki, Araya..., 2003 | | | Leptaulax masayukii Iwase, 1997 | | | Leptaulax masumotoi Johki, Araya..., 2011 | | | Leptaulax matsumotoi Kon, Johki &..., 2003 | | | Leptaulax matsumotoi kangfangensis Kon & Abe, 2009 | | | Leptaulax matsumotoi matsumotoi | | | Leptaulax mindanaoensis Iwase, 1996 | | | Leptaulax negrosensis Iwase, 1996 | | | Leptaulax niae Kuwert, 1898 | | | Leptaulax niisatoi Kon, Johki &..., 2017 | | | Leptaulax pacholatkoi Iwase, 1998 | | | Leptaulax pacholatkoi mutoniatus Neumann & Kon, 2013 | | | Leptaulax pacholatkoi pacholatkoi | | | Leptaulax palawanicus Zang, 1905 | | | Leptaulax parvus Iwase, 1995 | | | Leptaulax parvus balita Iwase, 1997 | | | Leptaulax parvus parvus balita Iwase, 1998 | | | Leptaulax planus (Illiger, 1800) | ⇒ Leptaulax roepstorffi Kuwert, 1898 | | Leptaulax punctatus Iwase, 1995 | | | Leptaulax riekoae Iwase, 1998 | | | Leptaulax riekoae kachinensis Kon, Araya &..., 2005 | | | Leptaulax riekoae riekoae kachinensis ..., 2005 | | | Leptaulax roepstorffi Kuwert, 1898 | | | Leptaulax sabahensis Iwase, 1996 | | | Leptaulax sakaii Iwase, 1996 | | | Leptaulax sambawae Gravely, 1918 | | | Leptaulax sarawakus Iwase, 1996 | | | Leptaulax sarawakus kashizakii Johki, Araya..., 2015 | | | Leptaulax sarawakus sarawakus | | | Leptaulax sawaii Johki, Araya..., 2015 | | | Leptaulax schillhammeri Iwase, 1998 | | | Leptaulax schoedli Iwase, 1998 | | | Leptaulax separandus Kuwert, 1891 | | | Leptaulax shunichii Kon, Johki &..., 2021 | | | Leptaulax similis Iwase, 1996 | | | Leptaulax strbai Iwase, 1998 | | | Leptaulax tamijii Kon, Araya &..., 2021 | | | Leptaulax timoriensis Percheron, 1841 | | | Leptaulax trajae Iwase, 1998 | | | Leptaulax uenoi (Iwase, 1995) | ⇒ Leptaulax arrowi Hincks, 1933 | | Macrolinus batesi (Kuwert, 1898) | ⇒ Macrolinus cartereti Boucher, 1996 | | Macrolinus cartereti Boucher, 1996 | | | Macrolinus depressus Gravely, 1918 | | | Macrolinus duivenbodei Kaup, 1868 | | | Macrolinus foveolatus Ma, 1988 | | | Macrolinus johkii Boucher & Kon, 1998 | | | Macrolinus latipennis Percheron, 1841 | | | Macrolinus medogensis Zang, 1981 | | | Macrolinus mineti Boucher, 1996 | | | Macrolinus puncticollis Hincks, 1938 | | | Macrolinus punctipectus Hincks, 1838 | | | Macrolinus salibabuensis Boucher, 1996 | | | Macrolinus sikkimensis Stoliczka, 1873 | | | Macrolinus sikkimensis sikkimensis sikkimensis ..., 1873 | | | Macrolinus sikkimensis tavoyanus Gravely, 1914 | | | Macrolinus sinicus Hong, 1984 | | | Macrolinus sulciperfectus Kuwert, 1891 | | | Macrolinus urus Heller, 1898 | | | Macrolinus weberi (Kaup, 1868) | ⇒ Macrolinus latipennis Percheron, 1841 | | Mastachilus australasicus Percheron, 1841 | | | Mastochilus australasicus Percheron, 1841 | | | Ophrygonius aequidens Gravely, 1914 | | | Ophrygonius birmanicus (Gravely, 1914) | ⇒ Basilianus birmanicus Gravely, 1914 | | Ophrygonius bleuzeni (Boucher, 1993) | ⇒ Basilianus bleuzeni Boucher, 1993 | | Ophrygonius cantori Percheron, 1844 | | | Ophrygonius chinensis (Endrodi, 1955) | ⇒ Basilianus chinensis Endrodi, 1955 | | Ophrygonius convexifrons Zang, 1904 | | | Ophrygonius emas Iwase, 1998 | | | Ophrygonius ferreri (Boucher, 1993) | ⇒ Basilianus ferreri Boucher, 1993 | | Ophrygonius griveaui (Boucher, 1993) | ⇒ Basilianus griveaui Boucher, 1993 | | Ophrygonius hirtimarginalis Iwase, 1993 | | | Ophrygonius inaequalis Burmeister, 1847 | | | Ophrygonius inopinus (Boucher, 1997) | ⇒ Basilianus inopinus Boucher, 1997 | | Ophrygonius javensis (Gravely, 1918) | ⇒ Basilianus javensis Gravely, 1918 | | Ophrygonius koni (Boucher, 1995) | ⇒ Basilianus koni Boucher, 1995 | | Ophrygonius lacroixi (Boucher, 1993) | ⇒ Basilianus lacroixi Boucher, 1993 | | Ophrygonius longiruga (Boucher, 1993) | ⇒ Basilianus longiruga Boucher, 1993 | | Ophrygonius megatarsi (Boucher & Kon, 1999) | ⇒ Basilianus megatarsi Boucher & Kon, 1999 | | Ophrygonius minor Gravely, 1914 | | | Ophrygonius planus Iwase, 1995 | | | Ophrygonius pseudaequidens (Boucher, 1995) | ⇒ Basilianus pseudaequidens Boucher, 1995 | | Ophrygonius rajanus (Doesburg, 1941) | ⇒ Basilianus rajanus Doesburg, 1941 | | Ophrygonius singapurae (Gravely, 1914) | ⇒ Basilianus singapurae Gravely, 1914 | | Ophrygonius sternibarbis (Boucher, 1995) | ⇒ Basilianus sternibarbis Boucher, 1995 | | Ophrygonius tanae (Boucher, 1993) | ⇒ Basilianus tanae Boucher, 1993 | | Ophrygonius tuberculosus (Boucher & Kon, 1999) | ⇒ Basilianus tuberculosus Boucher & Kon, 1999 | | Ophrygonius tuberculus Boucher, 1995 | | | Ophrygonius uedai (Kon & Johki, 1991) | ⇒ Basilianus uedai Kon & Johki, 1991 | | Paratiberioides laminatus Hincks, 1938 | | | Paratiberioides mandibularis Heller, 1900 | | | Paratiberioides mirabilis Iwase, 1994 | | | Paratiberioides similis Iwase, 1995 | | | Paratiberioides trajae Iwase, 1994 | | | Passalus basalis (Smith, 1852) | ⇒ Comacupes basalis Smith, 1852 | | Passalus sinuatus Eschscholtz, 1829 | | | Passalus tridens (Wiedemann, 1823) | ⇒ Pelopides tridens Wiedemann, 1823 | | Pelopides burmeisteri Kaup, 1868 | | | Pelopides dorsalis Kaup, 1871 | | | Pelopides gravidus Kuwert, 1891 | | | Pelopides mniszechi Kaup, 1868 | | | Pelopides monticulosus Smith, 1852 | | | Pelopides pedroi Kon, Johki &..., 2005 | | | Pelopides simplex Gravely, 1914 | | | Pelopides symmetricus Zang, 1904 | | | Pelopides tridens Wiedemann, 1823 | | | Pentalobus barbatus Fabricius, 1801 | | | Pentalobus palinii Percheron, 1844 | | | Plesthenus anneae Boucher, 1996 | | | Plesthenus gelon Schaufuss, 1885 | | | Plesthenus invitus Kuwert, 1891 | | | Plesthenus lottini Boisduval, 1835 | | | Plesthenus lottini gelon Schaufuss, 1885 | | | Plesthenus lottini lottini | | | Plesthenus sanghiensis Boucher, 1996 | | | Plesthenus scutellopunctatus Zang, 1903 | | | Pleurarius pilipes Kaup, 1868 | | | Proculus mniszechi Kaup, 1868 | | | Protomocoelus australis Boisduval, 1835 | | | Protomocoelus heynei Kuwert, 1891 | | | Protomocoelus otiosus Kuwert, 1898 | | | Protomocoelus triumphator Zang, 1904 | | | Serrulus sinicus ♰ Hong, 1983 | | | Taeniocerus bicanthatus Percheron, 1841 | | | Taeniocerus bicuspis Kaup, 1868 | | | Taeniocerus mourzinei Boucher, 1998 | | | Taeniocerus platypus Kaup, 1868 | | | Taeniocerus pygmaeus Kaup, 1868 | | | Tiberioides borealis Arrow, 1907 | | | Tiberioides kerleyi Kon & Araya, 2007 | | | Tiberioides kuwerti Arrow, 1907 | | | Trichostigmus mindorensis Boucher, 1992 | | | Trichostigmus mindoroensis Boucher, 1992 | | | Trichostigmus semperi Boucher, 1992 | | | Trichostigmus thoreyi Kaup, 1868 | | | Trichostigmus ursulus Schaufuss, 1885 | | | Trichostigmus whiteheadi Boucher, 1992 | |
Subnodes of Passalidae with occurrences in Far East
Links, sources and further information
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