| Subgenus | Copris (Copris)
Geoffroy, 1762
Mexican dung beetle | 12 | Species |
| Subgenus | Copris (Microcopris)
Balthasar, 1958
| 1 | Species |
| Subgenus | Copris (Paracopris)
Balthasar, 1939
| 1 | Species |
| Subgenus | Copris (Sinocopris)
| 4 | Species |
| Species | Copris 4dentatus Fabricius, 1798
| | |
| Species | Copris achates Péringuey, 1901 ⇒ Copris cornifrons
| | |
| Species | Copris acornis Geoffroy, 1785 ⇒ Palaeonthophagus nuchicornis
| | |
| Species | Copris acutidens Motschulsky, 1860
| | |
| Species | Copris aenescens Wiedemann, 1823 ⇒ Colobonthophagus aenescens
| | |
| Species | Copris aesculapius Olivier, 1790
| | |
| Species | Copris affinis Sturm, 1800
| | |
| Species | Copris afgoi Cambefort, 1992
| | |
| Species | Copris agnus Sharp, 1875
| | |
| Species | Copris ajax Sturm, 1826
| | |
| Species | Copris albicornis Palisot de Beauvois, 1805 ⇒ Onthophagus albicornis
| | |
| Species | Copris alexis Blanchard, 1846 ⇒ Ontherus alexis
| | |
| Species | Copris algol Nguyen-phung, 1989
| | |
| Species | Copris alluaudi Gillet, 0 ⇒ Copris atropolitus
| | |
| Species | Copris alutacea Wiedemann, 1823
| | |
| Species | Copris amabile Ferreira, 1958 ⇒ Copris amabilis
| | |
| Species | Copris amabilis Kolbe, 1914
| | |
| Species | Copris amazonicus Darling & Génier, 2018
| | |
| Species | Copris ammon Fabricius, 1781
| | |
| Species | Copris amyntas Olivier, 1790
| | |
| Species | Copris amyntor Klug, 1855 ⇒ Copris harrisi
| | |
| Species | Copris anaglyptica Mannerheim, 1828
| | |
| Species | Copris anaglypticus Say, 1823 ⇒ Dichotomius bos
| | |
| Species | Copris analis Germar, 1813 ⇒ Gibbonthophagus limbatus
| | |
| Species | Copris anceus Olivier, 1789
| | |
| Species | Copris andrewesi Waterhouse, 1891
| | |
| Species | Copris angolensis Ferreira, 1962 ⇒ Copris cassius
| | |
| Species | Copris angusticornis Arrow, 1933
| | |
| Species | Copris angustus Nguyen-phung & Cambefort, 1986
| | |
| Species | Copris angusutus Nguyen-phung & Cambefort, 1986
| | |
| Species | Copris anomiopseoides Boucomont, 1924 ⇒ Paracopris anomiopseoides
| | |
| Species | Copris antares Bacescu & Carausu, 1947
| | |
| Species | Copris antenor Olivier, 1789
| | |
| Species | Copris antilope Fabricius, 1798 ⇒ Digitonthophagus gazella
| | |
| Species | Copris apicepunctatus Balthasar, 1942
| | |
| Species | Copris appendiculata Mannerheim, 1828
| | |
| Species | Copris arachnoides Fourcroy, 1785 ⇒ Sisyphus schaefferi schaefferi
| | |
| Species | Copris arcturus Gillet, 1907
| | |
| Species | Copris ares Zelenka, 1993
| | |
| Species | Copris aricius Blanchard, 1846 ⇒ Dichotomius triangulariceps
| | |
| Species | Copris arimotoi Ochi & Araya, 1994
| | |
| Species | Copris arizonensis Schaeffer, 1906
| | |
| Species | Copris armatus Harold, 1869
| | |
| Species | Copris armeniacus Falderman, 1835
| | |
| Species | Copris armiger Gillet, 1910
| | |
| Species | Copris arrowi Felsche, 1910
| | |
| Species | Copris aspericollis Gillet, 1910
| 1 | Subspecies |
| Species | Copris assifera Eschscholtz, 1822
| | |
| Species | Copris astyanax Olivier, 1790
| | |
| Species | Copris aterrimus Fabricius, 1798 ⇒ Caccobius aterrimus
| | |
| Species | Copris atropolitus Gillet, 1910
| | |
| Species | Copris aurata Fabricius, 1801 ⇒ Onthophagus auratus
| | |
| Species | Copris babirussa Eschscholtz, 1822
| | |
| Species | Copris bacchus Thunberg, 1787
| | |
| Species | Copris badia Castelnau, 1840
| | |
| Species | Copris bartolozzi Ochi, Kon & Pham, 2019
| | |
| Species | Copris basilewskyi Ferreira, 1962
| | |
| Species | Copris basipunctatus Balthasar, 1942
| | |
| Species | Copris belisama Schrank, 1798 ⇒ Copris lunaris
| | |
| Species | Copris bellator Chevrolat, 1844
| | |
| Species | Copris bellicosus Olivier, 1790
| | |
| Species | Copris bengalensis Shiino, 1978
| | |
| Species | Copris bicuspis Germar, 1824 ⇒ Dichotomius (Selenocopris) bicuspis
| | |
| Species | Copris bidens Pfeffer, 1888 ⇒ Liatongus femoratus
| | |
| Species | Copris bifidus Poda von Neuhaus, 1761
| | |
| Species | Copris bihamatus Balthasar, 1965
| | |
| Species | Copris binodis Thunberg, 1818
| | |
| Species | Copris bituberculatus Scriba, 1790 ⇒ Palaeonthophagus verticicornis
| | |
| Species | Copris bootes Klug, 1855
| | |
| Species | Copris boreus Olivier, 1790
| | |
| Species | Copris bornemisszai Ferreira, 1968 ⇒ Copris bootes
| | |
| Species | Copris bos Blanchard, 1846 ⇒ Dichotomius bos
| | |
| Species | Copris boucardi Harold, 1869
| | |
| Species | Copris bovinus Gillet, 1908
| | |
| Species | Copris brachypterus Nomura, 1964
| 2 | Subspecies |
| Species | Copris brahminus Hope, 1831 ⇒ Synapsis brahminus
| | |
| Species | Copris brasiliana Germar, 1824 ⇒ Isocopris inhiatus
| | |
| Species | Copris breve Germar, 1824 ⇒ Eucanthidium breve
| | |
| Species | Copris brevicornis Darling & Génier, 2018
| | |
| Species | Copris brevis Germar, 1823
| | |
| Species | Copris brittoni Ferreira, 1962
| | |
| Species | Copris bucephalus Fabricius, 1775 ⇒ Heliocopris bucephalus
| | |
| Species | Copris buqueti Lucas, 1857 ⇒ Dichotomius buqueti
| | |
| Species | Copris caelatus Fabricius, 1794
| | |
| Species | Copris caerulescens Olivier, 1790
| | |
| Species | Copris caffer Gillet, 1908
| | |
| Species | Copris caliginosus Kohlmann, Cano & Delgado, 2003
| | |
| Species | Copris calvus Sharp, 1875
| | |
| Species | Copris cambeforti Nguyen-phung, 1988
| | |
| Species | Copris camerunus Felsche, 1904
| | |
| Species | Copris canadensis Fabricius, 1801 ⇒ Onthophagus (Onthophagus) orpheus
| | |
| Species | Copris caobangensis Bui, Dumack & Bonkowski, 2018
| | |
| Species | Copris capensis Waterhouse, 1891
| | |
| Species | Copris caraniceps Felsche, 1910
| | |
| Species | Copris carbonaria Mannerheim, 1828
| | |
| Species | Copris carbonarius Mannerheim, 1829 ⇒ Luederwaldtinia carbonarius
| | |
| Species | Copris carinicus Gillet, 1910
| | |
| Species | Copris carmelita Fabricius, 1801
| | |
| Species | Copris cartwrighti Robinson, 1941 ⇒ Copris fricator cartwrighti
| | |
| Species | Copris cassius Péringuey, 1901
| 1 | Subspecies |
| Species | Copris castaneus Mulsant, 1842 ⇒ Copris lunaris
| | |
| Species | Copris cavolinii V. Petagna, 1792
| | |
| Species | Copris celebensis Lansberge, 1886
| | |
| Species | Copris centricornis Fabricius, 1798 ⇒ Onthophagus centricornis
| | |
| Species | Copris cephalotes Péringuey, 1892
| | |
| Species | Copris cervus Fabricius, 1798
| | |
| Species | Copris cheni Ochi, Kon & Bai, 2007
| | |
| Species | Copris chimalapensis Mora-Aguilar & Delgado, 2015
| | |
| Species | Copris chinmontanus Ochi, Kon & Kawahara, 2019
| | |
| Species | Copris ciconia Fabricius, 1801 ⇒ Phalops iphis
| | |
| Species | Copris ciliatus
| | |
| Species | Copris ciprianii Balthasar, 1939
| | |
| Species | Copris claudius Harold, 1877 ⇒ Copris repertus
| | |
| Species | Copris clavicornis Matthews & Halffter, 1959 ⇒ Copris moecha
| | |
| Species | Copris coenosus Erichson, 1849
| | |
| Species | Copris colmanti Gillet, 1908
| | |
| Species | Copris colonica Say, 1835
| | |
| Species | Copris colonicus Say, 1835 ⇒ Dichotomius carolinus
| | |
| Species | Copris complexus Nguyen-phung, 1989
Robber Baron Cave meshweaver | | |
| Species | Copris compressipennis Gillet, 1910 ⇒ Paracopris compressipennis
| | |
| Species | Copris confucius Harold, 1877
| | |
| Species | Copris confusa Boheman, 1857
| | |
| Species | Copris confusus Boheman, 1857
| 1 | Subspecies |
| Species | Copris conicollis Blanchard, 1846 ⇒ Dichotomius conicollis
| | |
| Species | Copris consobrinus Lansberge, 1886 ⇒ Copris doriae
| | |
| Species | Copris conspicillata Weber, 1801
| | |
| Species | Copris conspurcatus Geoffroy, 1785 ⇒ Palaeonthophagus vacca
| | |
| Species | Copris continentalis Ochi & Kon, 2010
| | |
| Species | Copris contracta Boheman, 1857
| | |
| Species | Copris contractus Boheman, 1857 ⇒ Copris obesus
| | |
| Species | Copris convexiusculus Shipp, 1897 ⇒ Catharsius convexiusculus
| | |
| Species | Copris coriarius Gillet, 1907
| | |
| Species | Copris corniculatus Mulsant, 1842 ⇒ Copris lunaris
| | |
| Species | Copris cornifrons Boheman, 1860
| | |
| Species | Copris corniger Sahlberg, 1823 ⇒ Onthophagus (Onthophagus) taurus
| | |
| Species | Copris corpulentus Gillet, 1910
| | |
| Species | Copris corvus Fabricius, 1798
| | |
| Species | Copris costaricensis Gahan, 1894
| 2 | Subspecies |
| Species | Copris costariensis
| | |
| Species | Copris cotopaxi Guérin-Méneville, 1855
| | |
| Species | Copris crassus Deschodt & Davis, 2015
| | |
| Species | Copris crenatipennis Blanchard, 1846 ⇒ Dichotomius semiaeneus
| | |
| Species | Copris cribratus Gillet, 1927
| | |
| Species | Copris cribricollis Gillet, 1910 ⇒ Catharsius pithecius
| | |
| Species | Copris crinicollis Germar, 1824 ⇒ Dichotomius (Luederwaldtinia) crinicollis
| | |
| Species | Copris cristatus DeGeer, 1778 ⇒ Heliocopris bucephalus
| | |
| Species | Copris curlettii Moretto, 2011
| | |
| Species | Copris curvicornis Boheman, 1860 ⇒ Copris cornifrons
| | |
| Species | Copris curvifrons Péringuey, 1885 ⇒ Xinidium dewitzi
| | |
| Species | Copris cyaneus Fabricius, 1798
| | |
| Species | Copris dama Fabricius, 1798 ⇒ Colobonthophagus dama
| | |
| Species | Copris davidi Darling & Génier, 2018
| | |
| Species | Copris davidis Deyrolle, 1878
| | |
| Species | Copris davisi Nguyen-phung & Cambefort, 1986
| | |
| Species | Copris davisoni Waterhouse, 1891
| | |
| Species | Copris decellei Cambefort, 1992
| | |
| Species | Copris decorata Perty, 1830
| | |
| Species | Copris deletus Mulsant, 1842 ⇒ Copris lunaris
| | |
| Species | Copris delicatus Arrow, 1931
| | |
| Species | Copris denticulatus Nguyen-phung, 1988
| | |
| Species | Copris dicyrtus Matthews & Halffter, 1959
| | |
| Species | Copris diminutivus Walker, 1858 ⇒ Caccobius diminutivus
| | |
| Species | Copris discoideus Olivier, 1790
| | |
| Species | Copris diversus Waterhouse, 1891
| | |
| Species | Copris divisa Wiedemann, 1823 ⇒ Phalops divisus
| | |
| Species | Copris dorcas Olivier, 1790
| | |
| Species | Copris doriae Harold, 1877
| | |
| Species | Copris dorsopurcus Voet, 1769 ⇒ Canthon triangularis
| | |
| Species | Copris dorsoviridis Voet, 1769 ⇒ Canthon triangularis
| | |
| Species | Copris draco Arrow, 1906
| | |
| Species | Copris dracoides Frey, 0 ⇒ Copris megaceratoides
| | |
| Species | Copris dracunculus Fereira, 1959
| | |
| Species | Copris druidum ♰ O. Heer, 1862
| | |
| Species | Copris dudleyi Cambefort, 1992
| | |
| Species | Copris eburneus Cambefort, 1992
| | |
| Species | Copris egena Klug, 1835
| | |
| Species | Copris elpheneroides Felsche, 1910
| | |
| Species | Copris elphenor Klug, 1855
| | |
| Species | Copris emarginatus A. G. Olivier, 1789 ⇒ Copris lunaris
| | |
| Species | Copris ensifer Germar, 1821 ⇒ Coprophanaeus (Megaphanaeus) ensifer
| | |
| Species | Copris ephialtes Mannerheim, 1828
| | |
| Species | Copris erecta Wiedemann, 1819
| | |
| Species | Copris erectus Wiedeman, 1819 ⇒ Onthophagus (Serrophorus) sagittarius
| | |
| Species | Copris eridanus Olivier, 1790
| | |
| Species | Copris erratus Lansberge, 1886
| 2 | Subspecies |
| Species | Copris eryx Fabricius, 1801
| | |
| Species | Copris eteocles Castelnau, 1840
| | |
| Species | Copris evanidus Klug, 1855
| | |
| Species | Copris excavata Klug, 1855
| | |
| Species | Copris excavatus Klug, 1855 ⇒ Copris bootes
| | |
| Species | Copris excisus Waterhouse, 1891
| | |
| Species | Copris exigua Boheman, 1860
| | |
| Species | Copris exserta Sturm, 1843
| | |
| Species | Copris fagellatus Olivier, 1790
| | |
| Species | Copris fallaciosus Gillet, 1907
Kenyan dung beetle | | |
| Species | Copris fallax Felsche, 1910
| | |
| Species | Copris fasciatus Olivier, 1790
| | |
| Species | Copris fastidiosus Thomson, 1858
| | |
| Species | Copris felschei Reitter, 1893 ⇒ Copris armeniacus
| | |
| Species | Copris femoratus Illiger, 1800 ⇒ Liatongus femoratus
| | |
| Species | Copris ferreirae Frey, 1974 ⇒ Copris megaceratoides
| | |
| Species | Copris ferrugineus Olivier, 1790
| | |
| Species | Copris fidius Olivier, 1789
| | |
| Species | Copris fissicornis Steven, 1809
| | |
| Species | Copris fissifrons
| | |
| Species | Copris flavipellis Voet, 1769 ⇒ Canthon triangularis
| | |
| Species | Copris floriger Kirby, 1818 ⇒ Phanaeus (Notiophanaeus) splendidulus
| | |
| Species | Copris formosanus Gillet, 1922 ⇒ Copris (Copris) punctatus formosanus
| | |
| Species | Copris frankebergeri Balthasar, 1934
| | |
| Species | Copris frankenbergeri Balthasar, 1934 ⇒ Copris acutidens
| | |
| Species | Copris frankenbergerianus Balthasar, 1958
| | |
| Species | Copris freyi Ferreira, 1961
| | |
| Species | Copris fricator Fabricius, 1787
| 2 | Subspecies |
| Species | Copris fujiokai Ochi & Kon, 1996
| 1 | Subspecies |
| Species | Copris fulgidus Olivier, 1790
| | |
| Species | Copris fulliensis
| | |
| Species | Copris fulvus Fourcroy & Geoffroy, 1785
| | |
| Species | Copris furciceps Felsche, 1910
| | |
| Species | Copris furcillatus Felsche, 1910 ⇒ Copris costaricensis
| | |
| Species | Copris furculus Fabricius, 1798 ⇒ Onthophagus furculus
| | |
| Species | Copris fuscopunctatus Fabricius, 1798
| | |
| Species | Copris gagates Olivier, 1790
| | |
| Species | Copris garambae Cambefort, 1992
| | |
| Species | Copris gazellarum Gillet, 1918
| | |
| Species | Copris gibbosus Scriba, 1790 ⇒ Euonthophagus gibbosus
| | |
| Species | Copris gibbulus Lansberge, 1886
| 2 | Subspecies |
| Species | Copris gigas Olivier, 1789 ⇒ Heliocopris colossus
| | |
| Species | Copris gilleti Kolbe, 1907
| | |
| Species | Copris giraffa Hausmann, 1807
| | |
| Species | Copris gistelianus Gistel, 1857 ⇒ Copris lunaris
| | |
| Species | Copris glabriceps Felsche, 1910
| | |
| Species | Copris glabricollis Hope, 1842 ⇒ Coptodactyla glabricollis
| | |
| Species | Copris gladiator Arrow, 1933
| | |
| Species | Copris globulipennis Waterhouse, 1891
| | |
| Species | Copris gopheri Hubbard, 1894
Copris tortoise commensal scarab | | |
| Species | Copris gorillus Thomson, 1858
| | |
| Species | Copris gracilicornis Germar, 1813 ⇒ Onthophagus mopsus
| | |
| Species | Copris gracilis Waterhouse, 1891
| | |
| Species | Copris granulatus Olivier, 1790
| | |
| Species | Copris graueri Kolbe, 1914 ⇒ Copris camerunus
Madla Cave meshweaver | | |
| Species | Copris haemorrhoidalis Geoffroy, 1785 ⇒ Caccobius schreberi
| | |
| Species | Copris halffteri Matthews, 1959
| | |
| Species | Copris hamadryas Fabricius, 1775 ⇒ Heliocopris hamadryas
| | |
| Species | Copris haroldi Lansberge, 1886 ⇒ Copris spinator
| | |
| Species | Copris harpax Fabricius, 1801
| | |
| Species | Copris harrisi Waterhouse, 1891
| | |
| Species | Copris hartli Frey, 1975
| | |
| Species | Copris hastator Fabricius, 1798 ⇒
| | |
| Species | Copris hatayamai Ochi & Araya, 1992
| | |
| Species | Copris helwigi Fabricius, 1798 ⇒ Sisyphus longipes
| | |
| Species | Copris helwigii Fabricius, 1798
| | |
| Species | Copris hidakai Ochi & Kon, 1996
| | |
| Species | Copris hintoni Saylor, 1933 ⇒ Copris klugi
| | |
| Species | Copris hircus Wiedemann, 1823 ⇒ Tibiodrepanus hircus
| | |
| Species | Copris hirta Illiger, 1803 ⇒ Trichonthophagus hirtus
| | |
| Species | Copris hirtus Illiger, 1803
| | |
| Species | Copris hispanicus Poiret, 1789 ⇒ Copris hispanus hispanus
| | |
| Species | Copris hispanus Linnaeus, 1764
| 3 | Subspecies |
| Species | Copris hosakai Ochi & Kon, 2014
| | |
| Species | Copris howdeni Matthews & Halffter, 1959
| | |
| Species | Copris humilis Gillet, 1908
| | |
| Species | Copris hyaena Fabricius, 1801 ⇒ Onthophagus hyaena
| | |
| Species | Copris hybridus Gillet, 1910
| | |
| Species | Copris hypocrita Lucas, 1857 ⇒ Isocopris inhiatus
| | |
| Species | Copris icarus Olivier, 1790
| | |
| Species | Copris igualensis Warner, 1990
| | |
| Species | Copris illotus Balthasar, 1942
| | |
| Species | Copris imitans Felsche, 1910
| | |
| Species | Copris inaequabilis Zhang, 1997
| | |
| Species | Copris incertus Say, 1835
Black dung beetle, Mexican dung beetle, Poisonbean | 1 | Subspecies |
| Species | Copris indicus Gillet, 1910 ⇒ Copris fricator
| | |
| Species | Copris inemarginatus Blatchley, 1918
| | |
| Species | Copris inermis Castelnau, 1840 ⇒ Metacatharsius inermis
| | |
| Species | Copris inhalatus Quedenfeldt, 1884
| 5 | Subspecies |
| Species | Copris inhiata Germar, 1823
| | |
| Species | Copris insidiosus Péringuey, 1901
| | |
| Species | Copris integer Reiche, 1847 ⇒ Copris fallaciosus
| | |
| Species | Copris integra Reiche, 1850
| | |
| Species | Copris interioris Kolbe, 1897
| | |
| Species | Copris iris Sharp, 1875
| | |
| Species | Copris isidis Latreille, 1819 ⇒ Heliocopris gigas
| | |
| Species | Copris jacchoides Nguyen-phung & Cambefort, 1987
| | |
| Species | Copris jacchus Lang, 1971
| | |
| Species | Copris jahi Nguyen-phung & Cambefort, 1986
| | |
| Species | Copris janus Olivier, 1790
| | |
| Species | Copris japetus Klug, 1855
| | |
| Species | Copris jasius Olivier, 1790
| | |
| Species | Copris javana Fabricius, 1801
| | |
| Species | Copris javanensis Bui, Ziegler & Bonkowski, 2019
| | |
| Species | Copris jenisonianus Gistel, 1857 ⇒ Copris lunaris
| | |
| Species | Copris johkii Ochi & Kon, 2014
| | |
| Species | Copris juanmorai Mora-Aguilar & Delgado, 2015
| | |
| Species | Copris jucundus Gillet, 1932
| | |
| Species | Copris juvencus Scriba, 1790 ⇒ Euonthophagus gibbosus
| | |
| Species | Copris kasagii Ochi & Kon, 1996
| | |
| Species | Copris kasanka Josso & Prévost, 2016
| | |
| Species | Copris katangae Cambefort & Nguyen-phung, 1996
| | |
| Species | Copris keralensis Gill, 1986
| | |
| Species | Copris kimioi Ochi & Kon, 2021
| | |
| Species | Copris kiriromensis Ochi & Kon, 2021
| | |
| Species | Copris kiuchii Masumoto, 1989
| | |
| Species | Copris klapperichi Balthasar, 1942
| | |
| Species | Copris klugi Harold, 1869
| 1 | Subspecies |
| Species | Copris kusakabei Ochi, Kon & Kawahara, 2005
| | |
| Species | Copris laeviceps Harold, 1869
| | |
| Species | Copris laevicollis Mulsant, 1842 ⇒ Copris hispanus hispanus
| | |
| Species | Copris laevigatus Gillet, 1927 ⇒ Onthophagus laevigatus
| | |
| Species | Copris laiiformis Nguyen-phung & Cambefort, 1987
| | |
| Species | Copris laioides Boucomont, 1932
| | |
| Species | Copris laius Harold, 1868
| 2 | Subspecies |
| Species | Copris lamina Wiedemann, 1823
| | |
| Species | Copris lannathai Hanboonsong & Masumoto & Ochi, 2003
| | |
| Species | Copris laosensis Ochi & Kon, 2021
| | |
| Species | Copris latebrosis Fabricius, 1801 ⇒
| | |
| Species | Copris latebrosus Fabricius, 1801
| | |
| Species | Copris laticornis Boheman, 1857 ⇒ Copris fallaciosus
| 1 | Subspecies |
| Species | Copris latifrons Harold, 1868
| | |
| Species | Copris leakeyorum ♰ R. Paulian, 1976
| | |
| Species | Copris lecontei Matthews, 1961
| 2 | Subspecies |
| Species | Copris leruei Josso & Prévost, 2017
| | |
| Species | Copris leucostigma Steven, 1812
| | |
| Species | Copris lineata Wiedemann, 1823
| | |
| Species | Copris lineatus Wiedeman, 1823
| | |
| Species | Copris lividus Olivier, 1790
| | |
| Species | Copris longiceps Taschenberg, 1870
| | |
| Species | Copris longipes Olivier, 1790
| | |
| Species | Copris lucida Sturm, 1800 ⇒ Onthophagus (Palaeonthophagus) lucidus
| | |
| Species | Copris lucidus Illiger, 1800 ⇒ Onthophagus (Palaeonthophagus) lucidus
| | |
| Species | Copris lugubris Boheman, 1858
| | |
| Species | Copris lunarioides Waterhouse, 1891 ⇒ Copris fallaciosus
| 1 | Subspecies |
| Species | Copris lunaris Linnaeus, 1758
Horned Dung Beetle | 4 | Subspecies |
| Species | Copris lunaroides Waterhouse, 1891
| | |
| Species | Copris lunaticeps Gillet, 1910 ⇒ Pseudocopris lunaticeps
| | |
| Species | Copris lunus Schrank, 1798
| | |
| Species | Copris luteipennis Wiedemann, 1823 ⇒ Onthophagus centricornis
| | |
| Species | Copris macacus Lansberge, 1886
| | |
| Species | Copris macclevei Warner, 1990
| | |
| Species | Copris macer Péringuey, 1901
| | |
| Species | Copris maculosus Voet, 1769 ⇒ Canthon triangularis
| | |
| Species | Copris maesi Ratcliffe, 1998
| | |
| Species | Copris magicus Harold, 1881
| | |
| Species | Copris maki Illiger, 1803 ⇒ Trichonthophagus maki
| | |
| Species | Copris mannerheimi Harold, 1859
| | |
| Species | Copris manni Ochi, Kon & Bai, 2009
| | |
| Species | Copris marcoberrai Ochi, Kon & Pham, 2019
| | |
| Species | Copris marcus Gillet, 1921
| | |
| Species | Copris marginalis Gebler, 1817 ⇒ Onthophagus (Palaeonthophagus) marginalis
| | |
| Species | Copris marianus Petiver, 1764
| | |
| Species | Copris marsyas Olivier, 1790
| | |
| Species | Copris martinae Nguyen-phung, 1988
| 2 | Subspecies |
| Species | Copris martinezi Matthews & Halffter, 1968
Streaked blenny | | |
| Species | Copris masumotoi Ochi, Kon & Bai, 2020
| | |
| Species | Copris matthewsi Delgado & Kohlmann, 2001
| 2 | Subspecies |
| Species | Copris media Kugelann, 1792 ⇒ Onthophagus (Palaeonthophagus) vacca
| | |
| Species | Copris medius Kugelan, 1792 ⇒ Palaeonthophagus vacca
| | |
| Species | Copris medogensis Y.-W. Zhang, 1988
| | |
| Species | Copris megaceratoides Waterhouse, 1891
| | |
| Species | Copris megasoma Matthews & Halffter, 1959
| | |
| Species | Copris melanocephalus Olivier, 1790
| | |
| Species | Copris melitaeus Fabricius, 1798 ⇒ Amphionthophagus melitaeus
| | |
| Species | Copris mesacanthus Harold, 1878
| 2 | Subspecies |
| Species | Copris metallicus Fabricius, 1798 ⇒ Digitonthophagus gazella
| | |
| Species | Copris mexicanus Matthews & Halffter, 1959
| | |
| Species | Copris micros Ochi & Kon, 2021
| | |
| Species | Copris minator Harold, 1880 ⇒ Copris anceus
| | |
| Species | Copris mindorensis Ochi, 1992
| | |
| Species | Copris minuta Hausmann, 1807
| | |
| Species | Copris minutissimus Kryzhanovskii & Medvedev, 1966 ⇒ Copris (Microcopris) propinquus
| | |
| Species | Copris minutus Drury, 1773
Small Black Dung Beetle | | |
| Species | Copris misellus Péringuey, 1901
| | |
| Species | Copris miyakei Ochi & Kon, 2004
| | |
| Species | Copris modesta Boheman, 1857
| | |
| Species | Copris modestus Boheman, 1857 ⇒ Copris evanidus
| | |
| Species | Copris moecha LeConte, 1854
| | |
| Species | Copris moechus LeConte, 1854
| | |
| Species | Copris moffartsi Gillet, 1907
| | |
| Species | Copris mongkhoni Hanboonsong & Masumoto & Ochi, 2003
| | |
| Species | Copris montivagus Gillet, 1908 ⇒ Copris harrisi
| | |
| Species | Copris montreuili Ochi, Kon & Pham, 2019
| | |
| Species | Copris morettoi Josso & Prévost, 2020
| | |
| Species | Copris morgani Waterhouse, 1891 ⇒ Copris carmelita
| | |
| Species | Copris moroni Darling & Génier, 2018
| | |
| Species | Copris morphaeus Gillet, 1932
| | |
| Species | Copris mourgliai Zelenka, 1993
| | |
| Species | Copris muelleri Balthasar, 1939 ⇒ Copris acutidens
| | |
| Species | Copris mutica Boheman, 1857
| | |
| Species | Copris muticus Boheman, 1857 ⇒ Litocopris muticus
| | |
| Species | Copris nasuta Castelnau, 1840
| | |
| Species | Copris neglecta ⇒ Litocopris simplex
| | |
| Species | Copris neglectus Moxey, 1963
| | |
| Species | Copris neotridens Génier & Prévost, 2017
| | |
| Species | Copris nepos Gillet, 1908
| | |
| Species | Copris nepticulus Kolbe, 1897 ⇒ Copris diversus
| | |
| Species | Copris neptis Reiche, 1847
| | |
| Species | Copris nevinsoni Waterhouse, 1891
Warton meshweaver | | |
| Species | Copris nigella Illiger, 1803
| | |
| Species | Copris nigellus Illiger, 1803 ⇒ Parentius nigellus
| | |
| Species | Copris nimrod Fabricius, 1801
| | |
| Species | Copris nisus Olivier, 1789 ⇒ Luederwaldtinia nisus
| | |
| Species | Copris nitens Olivier, 1790
| | |
| Species | Copris nitida ⇒ Litocopris simplex
| | |
| Species | Copris nonarmatus Ochi & Kon, 2021
| | |
| Species | Copris novaki Zelenka, 1992
| | |
| Species | Copris nubilosus Kohlmann & Cano & Delgado, 2003
| | |
| Species | Copris nuchidens Fabricius, 1798 ⇒ Gibbonthophagus limbatus
| | |
| Species | Copris numa Lansberge, 1886
Braken Bat Cave meshweaver | | |
| Species | Copris obenbergeri Balthasar, 1933
Government Canyon Bat Cave meshweaver | | |
| Species | Copris obesus Boheman, 1857
| | |
| Species | Copris obliquatus Voet, 1806 ⇒ Canthon quadriguttatus
| | |
| Species | Copris obliquus Olivier, 1790
| | |
| Species | Copris obliteratus Mulsant, 1842 ⇒ Copris lunaris
| | |
| Species | Copris obtecta Palisot de Beauvois, 1805
| | |
| Species | Copris obtectus Palisot de Beauvois, 1805 ⇒ Onthophagus (Onthophagus) hecate
| | |
| Species | Copris obtusa Wiedemann, 1819
| | |
| Species | Copris obtusus Wiedeman, 1819 ⇒ Onthophagus (Serrophorus) sagittarius
| | |
| Species | Copris ochus Motschulsky, 1860
| | |
| Species | Copris oedipus Fabricius, 1775
| | |
| Species | Copris olivieri Illiger, 1800
| 5 | Subspecies |
| Species | Copris opacus Blanchard, 1846 ⇒ Dichotomius (Luederwaldtinia) carbonarius
| | |
| Species | Copris orientalis Fabricius, 1792
| | |
| Species | Copris orion Klug, 1835
| 4 | Subspecies |
| Species | Copris orphanus Guérin-Méneville, 1847 ⇒ Copris evanidus
| | |
| Species | Copris ovalipennis Blanchard, 1845 ⇒ Ontherus sulcator
| | |
| Species | Copris overlaeti Balthasar, 1961
| | |
| Species | Copris palemo Olivier, 1790
| | |
| Species | Copris pallens Olivier, 1790
| | |
| Species | Copris paniscus Fabricius, 1775
| 4 | Subspecies |
| Species | Copris paramaribous
| 5 | Subspecies |
| Species | Copris parapecuarius Ochi, Kon & Kawahara, 2008
| | |
| Species | Copris parasonensis Ochi, Kon & Pham, 2019
| | |
| Species | Copris paraszechouanicus Ochi, Kon & Pham, 2018
| | |
| Species | Copris pardalis Fabricius, 1798 ⇒ Onthophagus pardalis
| | |
| Species | Copris parvulus Fabricius, 1798
| | |
| Species | Copris pauliani Nguyen-phung & Cambefort, 1986
| | |
| Species | Copris pauperata Germar, 1823
| | |
| Species | Copris pectoralis Thunberg, 1818
| | |
| Species | Copris peculiariprotibialis Ochi & Kon, 2023
| | |
| Species | Copris pedarioides Lansberge, 1886
| | |
| Species | Copris peleus Olivier, 1790
| | |
| Species | Copris perseus
| | |
| Species | Copris persianus Miksic, 1958 ⇒ Copris hispanus hispanus
| | |
| Species | Copris persimilis Felsche, 1908 ⇒ Copris fallaciosus
| | |
| Species | Copris perturbator Péringuey, 1901 ⇒ Copris inhalatus perturbator
| | |
| Species | Copris phidias Olivier, 1790
| | |
| Species | Copris phungae Cambefort, 1992
| 2 | Subspecies |
| Species | Copris phylax Gillet, 1908
| | |
| Species | Copris pirmal Fabricius, 1798 ⇒ Heliocopris pirmal
| | |
| Species | Copris pithecius Olivier, 1790 ⇒ Catharsius sesostris
| | |
| Species | Copris platycera Wiedemann, 1823
| | |
| Species | Copris pluridens Gillet, 1908 ⇒ Copris interioris
| | |
| Species | Copris plutus Péringuey, 1901 ⇒ Copris fidius
| | |
| Species | Copris poggii Ochi & Kon, 2005
| | |
| Species | Copris politus Fabricius, 1798 ⇒ Onthophagus (Parascatonomus) politus
| | |
| Species | Copris polynice Blanchard, 1845 ⇒ Ontherus appendiculatus
| | |
| Species | Copris pompilius Waterhouse, 1875 ⇒ Copris sarpedon
| | |
| Species | Copris popaensis Ochi, Kon & Kawahara, 2019
| | |
| Species | Copris potanini Semenov, 1891
| | |
| Species | Copris prevosti Josso, 2011
| | |
| Species | Copris pristinus ♰ W. D. Pierce, 1946
| | |
| Species | Copris procidua Say, 1835 ⇒ Copris incertus
| | |
| Species | Copris prociduus Say, 1835 ⇒ Copris incertus
| | |
| Species | Copris pronus Gerstaecker, 1884
| | |
| Species | Copris protensa Perty, 1830
| | |
| Species | Copris pseudobootes Ferreira, 1962
| | |
| Species | Copris pseudochus Ochi & Kon, 2004
| 3 | Subspecies |
| Species | Copris pseudolunaris Ferreira, 1969 ⇒ Copris hispanus
| | |
| Species | Copris pseudomoffartsi Moretto, 2011
| | |
| Species | Copris pseudosinicus Balthasar, 1958
| | |
| Species | Copris pubescens Olivier, 1790
| | |
| Species | Copris pueli Mollandin De Boissy, 1905
| | |
| Species | Copris pumilionis Wallengren, 1881
| | |
| Species | Copris punctata Illiger, 1803
| | |
| Species | Copris punctatissima Curtis, 1844
| | |
| Species | Copris puncticollis Boheman, 1857
| | |
| Species | Copris punctipennis Boucomont, 1914
| | |
| Species | Copris punctiventris Waterhouse, 1891
| | |
| Species | Copris punctulatus Wiedemann, 1823
| 2 | Subspecies |
| Species | Copris punjabensis Gillet, 1921
| | |
| Species | Copris pusillus Fabricius, 1798
| | |
| Species | Copris quadrata Erichson, 1848
| | |
| Species | Copris quadricornis Fabricius, 1801
| | |
| Species | Copris quadridens Say, 1835 ⇒ Phanaeus (Phanaeus) quadridens
| | |
| Species | Copris quadridentatus Fabricius, 1798 ⇒ Copris lunaris
| | |
| Species | Copris quadriguttatus Olivier, 1790
| | |
| Species | Copris quadripunctatus Olivier, 1790
| | |
| Species | Copris quadrum Kugelann, 1792 ⇒ Onthophagus (Onthophagus) taurus
| | |
| Species | Copris quasilaevigatus Ochi, Kon & Bai, 2007
| | |
| Species | Copris ramosa Wiedemann, 1823
| | |
| Species | Copris ramosiceps Gillet, 1921
| | |
| Species | Copris ramosus Wiedemann, 1823 ⇒ Onthophagus ramosus
| | |
| Species | Copris rebouchei Harold, 1869
| | |
| Species | Copris reflexus Fabricius, 1787 ⇒ Copris minutus
| | |
| Species | Copris remotus LeConte, 1866
| 2 | Subspecies |
| Species | Copris renwarti Nguyen-phung & Cambefort, 1986
| | |
| Species | Copris repertus Walker, 1858
| | |
| Species | Copris retusus Mulsant, 1842 ⇒ Copris hispanus hispanus
| | |
| Species | Copris rhinoceros Klug, 1855
| | |
| Species | Copris ribbei Lansberge, 1886
| | |
| Species | Copris riekoae Ochi & Kon, 2003
| | |
| Species | Copris ritsemae Harold, 1875
| | |
| Species | Copris rotundatus Blanchard, 1845 ⇒ Ontherus appendiculatus
| | |
| Species | Copris ruandanus Balthasar, 1961
| | |
| Species | Copris rubidus Olivier, 1790
| | |
| Species | Copris ruficollis Germar, 1823
| | |
| Species | Copris rugosus Gillet, 1908 ⇒ Copris inhalatus
| | |
| Species | Copris rupicapra Gravenhorst, 1807
| | |
| Species | Copris ruricola Balthasar, 1933
| | |
| Species | Copris sabinus Gillet, 1910
| | |
| Species | Copris sacontala Redtenbacher, 1848
| | |
| Species | Copris sagax Quenstedt, 1806 ⇒ Catharsius sagax
| | |
| Species | Copris sallei Harold, 1869
| | |
| Species | Copris sanctus Fabricius, 1798 ⇒ Scarabaeus sanctus
| | |
| Species | Copris saphirinus Sturm, 1826
| | |
| Species | Copris sarpedon Harold, 1868
| | |
| Species | Copris saundersi Harold, 1869
| | |
| Species | Copris scalpellum Taschenberg, 1870 ⇒ Dichotomius cotopaxi
| | |
| Species | Copris schoutedeni Boucomont, 1928 ⇒ Copris wiesei
| | |
| Species | Copris scorpio Zelenka, 1993
| | |
| Species | Copris semiaenea Germar, 1823
| | |
| Species | Copris semicuprea Germar, 1823
| | |
| Species | Copris semisquamosa Curtus, 1844
| | |
| Species | Copris senegalensis Gillet, 1910 ⇒ Ontherus sulcator
| | |
| Species | Copris serius Nguyen-phung, 1987
| | |
| Species | Copris serratus Fourcroy & Geoffroy, 1785 ⇒ Ateuchetus laticollis
| | |
| Species | Copris servius Harold, 1877 ⇒ Copris agnus
| | |
| Species | Copris setosus Wiedemann, 1823 ⇒ Tibiodrepanus setosus
| | |
| Species | Copris sexdentata Redtenbacher, 1848
| | |
| Species | Copris sexdentatus Thunberg, 1818 ⇒ Copris sarpedon
| | |
| Species | Copris sexpunctatus Olivier, 1790
| | |
| Species | Copris siamensis Gillet, 1910
| | |
| Species | Copris siangensis Biswas, 1980
| | |
| Species | Copris sierrensis Matthews, 1961
| | |
| Species | Copris signatus Walker, 1858
| | |
| Species | Copris similis Scriba, 1790 ⇒ Copris mesacanthus
| | |
| Species | Copris simillimus Felsche, 1910 ⇒ Copris minutus
| | |
| Species | Copris simonettai Cambefort, 1992
| | |
| Species | Copris simplex Harold, 1869 ⇒ Dichotomius (Selenocopris) simplex
| | |
| Species | Copris simulator Nguyen-phung & Cambefort, 1986
| | |
| Species | Copris sinensis Hope, 1842 ⇒ Catharsius pithecius
| | |
| Species | Copris singularis Nguyen-phung & Cambefort, 1986
| | |
| Species | Copris sinicus Hope, 1842
| | |
| Species | Copris sinon Olivier, 1789
| | |
| Species | Copris sinuatus Mulsant, 1842 ⇒ Copris hispanus hispanus
| | |
| Species | Copris smaragdifer Walker, 1858
| | |
| Species | Copris smaragdina Perty, 1830
| | |
| Species | Copris sodalis Walker, 1858
| | |
| Species | Copris sonensis Bui, Dumack & Bonkowski, 2018
| | |
| Species | Copris sorex Balthasar, 1942
| | |
| Species | Copris sphaeropterus Harold, 1877
| | |
| Species | Copris spinator Harold, 1881
| | |
| Species | Copris striatulus Palisot de Beauvois, 1809 ⇒ Onthophagus (Onthophagus) striatulus
| | |
| Species | Copris subaenea Castelnau, 1840
| | |
| Species | Copris subaeneus Palisot de Beauvois, 1805
| | |
| Species | Copris subdolus Balthasar, 1958
| | |
| Species | Copris subpunctatus Gillet, 1910
| | |
| Species | Copris subsidens Péringuey, 1901
| | |
| Species | Copris subterranea ♰ O. Heer, 1862
| | |
| Species | Copris subterraneus Heer, 1862
| | |
| Species | Copris sulcicollis Lansberge, 1886 ⇒ Copris sinicus
| | |
| Species | Copris sumatrensis Gillet, 1921
| | |
| Species | Copris surdus Arrow, 1931
| | |
| Species | Copris susanae Darling & Génier, 2018
| | |
| Species | Copris szechouanicus Balthasar, 1958
| | |
| Species | Copris tages Olivier, 1790
| | |
| Species | Copris takanoi Ochi, Kon & Bai, 2020
| | |
| Species | Copris taprobanus Gillet, 1910 ⇒ Copris sodalis
| | |
| Species | Copris teramachii Ochi, Kon & Bai, 2020
| | |
| Species | Copris terminata Eschscholtz, 1822
| | |
| Species | Copris tetraodon Gillet, 1910
| | |
| Species | Copris thoracicus Olivier, 1790
| | |
| Species | Copris tiberius Schaufuss, 1885
| | |
| Species | Copris tmolus Fischer von Waldheim, 1821 ⇒ Synapsis tmolus
| | |
| Species | Copris torulosus Eschscholtz, 1822
| | |
| Species | Copris triangulariceps Blanchard, 1846 ⇒ Dichotomius triangulariceps
| | |
| Species | Copris tricera Wiedemann, 1823
| | |
| Species | Copris tricorniprothoracicus Ochi, Kon & Kawahara, 2020
| | |
| Species | Copris tricornis Wiedemann, 1823
| | |
| Species | Copris tridens Felsche, 1901 ⇒ Copris hispanus hispanus
| | |
| Species | Copris tridentatus Solis & Kohlmann, 2003
| | |
| Species | Copris trituber Wiedemann, 1823 ⇒ Onthophagus (Onthophagus) trituber
| | |
| Species | Copris troglodyta Wiedemann, 1823 ⇒ Onthophagus troglodyta
| | |
| Species | Copris troglodytarum Roth, 1851
| | |
| Species | Copris truncatus Felsche, 1901
| | |
| Species | Copris tsukamatoi Ochi, Kon & Kawahara, 2008
| | |
| Species | Copris tsukamotoi Ochi, Kon & Kawahara, 2008
| | |
| Species | Copris tubericollis Felsche, 1910 ⇒ Pseudocopris tubericollis
| | |
| Species | Copris tullius Olivier, 1789
| | |
| Species | Copris typhoeus Gerstaecker, 1884
| | |
| Species | Copris tyrannus Thomson, 1859
| | |
| Species | Copris umbilicatus Abeille de Perrin, 1901
| | |
| Species | Copris undatus Olivier, 1790
| | |
| Species | Copris unguiculatus Olivier, 1790
| | |
| Species | Copris unicornis Fabricius, 1798 ⇒ Caccobius (Caccophilus) unicornis
| | |
| Species | Copris ursus Fabricius, 1801
| | |
| Species | Copris urus Boheman, 1857 ⇒ Catharsius molossus
| | |
| Species | Copris usambaricus Gillet, 1908
| | |
| Species | Copris valdiviana Philippi, 1859
| | |
| Species | Copris valdivianus Philippi, 1859 ⇒ Dichotomius torulosus valdivianus
| | |
| Species | Copris vankhaii Nguyen-phung, 1988
| 2 | Subspecies |
| Species | Copris vankhalii Nguyen-phung, 1988
| | |
| Species | Copris variegatus Fabricius, 1798
| | |
| Species | Copris venator Fabricius, 1801 ⇒ Liatongus venator
| | |
| Species | Copris versipellis Voet, 1769 ⇒ Canthon triangularis
| | |
| Species | Copris vertagus Fabricius, 1798 ⇒ Liatongus vertagus
| | |
| Species | Copris victorini Boheman, 1857 ⇒ Copris ritsemae
| | |
| Species | Copris vietnamicus Kabakov, 1994
| | |
| Species | Copris vietnamimontanus Ochi, Kon & Pham, 2018
| | |
| Species | Copris vilhenai Ferreira, 1962
| | |
| Species | Copris violaceus Olivier, 1790
| | |
| Species | Copris viridis Palisot de Beauvois, 1805
| | |
| Species | Copris vitalisi Gillet, 1921
| | |
| Species | Copris vitulus Fabricius, 1798
| | |
| Species | Copris vrydaghi Ferreira, 1962
| | |
| Species | Copris vulcanus Fabricius, 1801 ⇒ Caccobius vulcanus
| | |
| Species | Copris warneri Mccleve & Kohlmann, 2005
| | |
| Species | Copris wiesei Kolbe, 1914
| | |
| Species | Copris xingkeyangi Ochi, Kon & Bai, 2020
| | |
| Species | Copris yamamotoi Tesar, 1937 ⇒ Copris (Sinocopris) pecuarius
| | |
| Species | Copris yangi Ochi, Kon & Bai, 2007
| | |
| Species | Copris youngai Balthasar, 1967
| | |
| Species | Copris zhangi Ochi, Kon & Bai, 2009
| | |
