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Checklist for Family Passalidae Leach, 1815 (bess beetle)
and region Borneo (Indonesia)


Class
 
Insecta
Linnaeus, 1758
Insects, True insects
420
4676
14052
Families
Genera
Species
 
Taxom
 

Subclass
 
Pterygota
Lang, 1888
Winged insects
420
4676
14052
Families
Genera
Species
 
Taxom
 

Infraclass
 
Neoptera
van der Wulp, 1890
Modern Wing-folding insects
400
4569
13774
Families
Genera
Species
 
Taxom
 

Superorder
 
Holometabola
Heider, 1889
Holometabolous Insects
256
3371
10525
Families
Genera
Species
 
Taxom
 

Order
 
Coleoptera
Linnaeus, 1758
Beetles, Coleopterans
77
1597
5463
Families
Genera
Species
 
Taxom
 

Suborder
 
Polyphaga
Emery, 1886
Polyphags
73
1425
4831
Families
Genera
Species
 
Taxom
 

Infraorder
 
Scarabaeiformia
Crowson, 1960
9
218
1268
Families
Genera
Species
 
Taxom
 

Superfamily
 
Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles
9
218
1268
Families
Genera
Species
 
Taxom
 
FamilyPassalidae
Leach, 1815
Bess beetle, Bess beetles, Betsy beetles, Passalid beetles
11
93
Genera
Species
 
Taxom
 


Genera for Borneo
11 Genera with 93 Species
Aceraius Kaup, 189827SpeciesTaxom
Aulacocyclus Kaup, 18681SpeciesTaxom
Basilianus Kaup, 199110SpeciesTaxom
Cacoius Boucher, 19931SpeciesTaxom
Comacupes Kaup, 19147SpeciesTaxom
Labienus Kaup, 18681SpeciesTaxom
Leptaulax Kaup, 184128SpeciesTaxom
Macrolinus Kaup, 18412SpeciesTaxom
Ophrygonius Zang, 199111SpeciesTaxom
Pelopides Kuwert, 19043SpeciesTaxom
Taeniocerus Kaup, 18684SpeciesTaxom


Species list for Borneo
93 Species and 3 Subspecies
Aceraius alpinus Kon, Ueda & ..., 1995
Aceraius alutaceosternus Kuwert, 1898
Aceraius borneanus (Kaup, 1871)⇒ Aceraius hikidai Kon, Ueda & ..., 1993
Aceraius boucheri Kon, Araya &..., 1993
Aceraius grandis Burmeister, 1847
Aceraius hassani Kon, Araya &..., 2006
Aceraius helferi Kuwert, 1891
Aceraius hidakai Kon, Araya &..., 1993
Aceraius hikidai Kon, Ueda & ..., 1993
Aceraius illegalis Kuwert, 1891
Aceraius jenisi Iwase, 1998
Aceraius katsurae Iwase, 1995
Aceraius kikutai Kon, Johki &..., 1995
Aceraius kinabalensis Kon & Johki, 1989
Aceraius kuwerti Zang, 1903
Aceraius laevicollis Illiger, 1800
Aceraius laevimargo Zang, 1905
Aceraius lamellatus Gravely, 1918
Aceraius laniger Zang, 1905
Aceraius moeschleri (Kon, Araya &..., 1992)⇒ Aceraius boucheri Kon, Araya &..., 1993
Aceraius oculidens Zang, 1905
Aceraius pilifer (Percheron, 1835)⇒ Aceraius helferi Kuwert, 1891
Aceraius riekoae Iwase, 1998
Aceraius sabanus Kon, Ueda & ..., 1995
Aceraius sawaii Kon, Araya &..., 2006
Aceraius tricornis Zang, 1903
Aceraius wallacei Kuwert, 1898
Aulacocyclus perlatus Kaup, 1868
Basilianus aequidens Gravely, 1914
Basilianus emas Iwase, 1998
Basilianus hirtimarginalis Iwase, 1993
Basilianus jacquesi Boucher, 1993
Basilianus koni Boucher, 1995
Basilianus pseudaequidens Boucher, 1995
Basilianus rajanus Doesburg, 1941
Basilianus singapurae Gravely, 1914
Basilianus sternibarbis Boucher, 1995
Basilianus uedai Kon & Johki, 1991
Cacoius jacquesi (Boucher, 1993)⇒ Basilianus jacquesi Boucher, 1993
Comacupes cavicornis Kaup, 1868
Comacupes cylindraceus (Perty, 1831)⇒ Comacupes masoni Stoliczka, 1873
Comacupes foveicollis Kuwert, 1891
Comacupes foveicollis foveicollis
Comacupes intermedius Dibb, 1937
Comacupes masoni Stoliczka, 1873
Comacupes stoliczkae Gravely, 1914
Labienus ptox Kaup, 1868
Leptaulax acutangularis Iwase, 1998
Leptaulax anibarbis Kuwert, 1898
Leptaulax arayai Johki & Kon, 2003
Leptaulax barbicauda Zang, 1905
Leptaulax beccarii Kuwert, 1891
Leptaulax bicolor (Fabricius, 1801)⇒ Leptaulax dentatus Fabricius, 1792
Leptaulax circulus Iwase, 1996
Leptaulax crockerensis Iwase, 1998
Leptaulax cyclotaenius Kuwert, 1891
Leptaulax datar Iwase, 1996
Leptaulax dentatus Fabricius, 1792
Leptaulax dentifrons Iwase, 1996
Leptaulax hidakai Johki, Araya..., 2003
Leptaulax humerosus (Kuwert, 1891)⇒ Leptaulax dentatus Fabricius, 1792
Leptaulax jenisi Iwase, 1998
Leptaulax kecil Iwase, 1996
Leptaulax malaccae Kuwert, 1891
Leptaulax manis Iwase, 1996
Leptaulax parvus balita Iwase, 1997
Leptaulax parvus parvus balita Iwase, 1998
Leptaulax planus (Illiger, 1800)⇒ Leptaulax roepstorffi Kuwert, 1898
Leptaulax roepstorffi Kuwert, 1898
Leptaulax sabahensis Iwase, 1996
Leptaulax sarawakus Iwase, 1996
Leptaulax schillhammeri Iwase, 1998
Leptaulax similis Iwase, 1996
Leptaulax strbai Iwase, 1998
Leptaulax timoriensis Percheron, 1841
Macrolinus johkii Boucher & Kon, 1998
Macrolinus latipennis Percheron, 1841
Ophrygonius aequidens Gravely, 1914
Ophrygonius emas Iwase, 1998
Ophrygonius hirtimarginalis Iwase, 1993
Ophrygonius inaequalis Burmeister, 1847
Ophrygonius koni (Boucher, 1995)⇒ Basilianus koni Boucher, 1995
Ophrygonius planus Iwase, 1995
Ophrygonius pseudaequidens (Boucher, 1995)⇒ Basilianus pseudaequidens Boucher, 1995
Ophrygonius rajanus (Doesburg, 1941)⇒ Basilianus rajanus Doesburg, 1941
Ophrygonius singapurae (Gravely, 1914)⇒ Basilianus singapurae Gravely, 1914
Ophrygonius sternibarbis (Boucher, 1995)⇒ Basilianus sternibarbis Boucher, 1995
Ophrygonius uedai (Kon & Johki, 1991)⇒ Basilianus uedai Kon & Johki, 1991
Pelopides burmeisteri Kaup, 1868
Pelopides monticulosus Smith, 1852
Pelopides symmetricus Zang, 1904
Taeniocerus bicanthatus Percheron, 1841
Taeniocerus bicuspis Kaup, 1868
Taeniocerus platypus Kaup, 1868
Taeniocerus pygmaeus Kaup, 1868

Subnodes of Passalidae with occurrences in Borneo
SubfamilyAulacocyclinae
Kaup, 1868
3
11
Genera
Species

Taxom

SubfamilyPassalinae
Leach, 1815
8
82
Genera
Species

Taxom



Links, sources and further information
Passalidae at gbif.org
Passalidae at irmng.org
Passalidae at itis.gov