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Checklist for Subfamily Harpalinae Bonelli, 1810
and region Bhutan (South Asia)


Class
 
Insecta
Linnaeus, 1758
Insects, True insects
205
1532
2897
Families
Genera
Species
 
Taxom
 

Subclass
 
Pterygota
Lang, 1888
Winged insects
205
1532
2897
Families
Genera
Species
 
Taxom
 

Infraclass
 
Neoptera
van der Wulp, 1890
Modern Wing-folding insects
198
1516
2881
Families
Genera
Species
 
Taxom
 

Superorder
 
Holometabola
Heider, 1889
Holometabolous Insects
138
1363
2673
Families
Genera
Species
 
Taxom
 

Order
 
Coleoptera
Linnaeus, 1758
Beetles, Coleopterans
50
546
1154
Families
Genera
Species
 
Taxom
 

Suborder
 
Adephaga
Schellenberg, 1806
Adephagans, Ground Beetles and Water Beetles
2
89
213
Families
Genera
Species
 
Taxom
 

Superfamily
 
Caraboidea
Latreille, 1802
1
87
211
Family
Genera
Species
 
Taxom
 
FamilyCarabidae
Latreille, 1802
Carabid beetles, Carabids, Ground Beetles, Tiger Beetles
87
211
Genera
Species
 
Taxom
 
SubfamilyHarpalinae
Bonelli, 1810
46
118
Genera
Species
 
Taxom
 


Genera for Bhutan
46 Genera with 118 Species
#ABCDHLMNOPSTX
Abacetus Dejean, 18691SpeciesTaxom
Agonum Bonelli, 18581SpeciesTaxom
Amara Bonelli, 2011Seed-Eating Ground Beetles7SpeciesTaxom
Amblops Andrewes, 19311SpeciesTaxom
Amphimenes Bates, 19311SpeciesTaxom
Anthracus Motschulsky, 20151SpeciesTaxom
Bradycellus Erichson, 18731SpeciesTaxom
Calleida Latreille & ..., 18921SpeciesTaxom
Chlaenius Bonelli, 1856Vivid Metallic Ground Beetle20SpeciesTaxom
Chydaeus Chaudoir, 18996SpeciesTaxom
Deltocolpodes Morvan, 20021SpeciesTaxom
Dendrocellus Schmidt-Goebel, 18313SpeciesTaxom
Dicranoncus Chaudoir, 19351SpeciesTaxom
Dinodes Bonet, 18562SpeciesTaxom
Dioryche W.S. Macleay, 19223SpeciesTaxom
Dromius Bonelli, 19773SpeciesTaxom
Harpalus Latreille, 18913SpeciesTaxom
Lebia Latreille, 1871colorful foliage ground beetle3SpeciesTaxom
Lepcha Andrewes, 19821SpeciesTaxom
Lesticus Dejean, 18944SpeciesTaxom
Lioptera Chaudoir, 19032SpeciesTaxom
Meleagros Kirschenhofer, 19231SpeciesTaxom
Microlestes Schmidt-Goebel, 19233SpeciesTaxom
Nepalorthogonius Habu, 20161SpeciesTaxom
Nirmala Andrewes, 19301SpeciesTaxom
Onycholabis Bates, 18921SpeciesTaxom
Orthogonius MacLeay, 18465SpeciesTaxom
Oxycentrus Chaudoir, 20131SpeciesTaxom
Parena Motschulsky, 19772SpeciesTaxom
Philorhizus Hope, 19231SpeciesTaxom
Physodera Eschscholtz, 18291SpeciesTaxom
Platymetopus Dana, 17981SpeciesTaxom
Platynus Bonelli, 19962SpeciesTaxom
Prakasha Andrewes, 18921SpeciesTaxom
Pristosia Motschulsky, 18771SpeciesTaxom
Psychristus Andrewes, 19303SpeciesTaxom
Pterostichus Bonelli, 1937Woodland Ground Beetle16SpeciesTaxom
Scalidion Schmidt-Gobel, 18461SpeciesTaxom
Sericoda Kirby, 17741SpeciesTaxom
Stenolophus Dejean, 1938Seedcorn Beetle3SpeciesTaxom
Steropanus Fairmaire, 19371SpeciesTaxom
Syntomus Hope, 18441SpeciesTaxom
Trichotichnus A. Morawitz, 19302SpeciesTaxom
Trigonotoma Dejean, 18941SpeciesTaxom
Xestagonum Habu, 199913SpeciesTaxom
Xestopus Andrewes, 19783SpeciesTaxom


Species list for Bhutan
118 Species and 16 Subspecies
Abacetus (Triaenabacetus) quadrinotatus Chaudoir, 1869
Agonum (Agonum) scintillans Boheman, 1858
Amara (Amara) darjelingensis Putzeys, 1877
Amara (Amara) nila Andrewes, 1924
Amara (Bradytus) batesi Csiki, 1929
Amara (Bradytus) chalciope Bates, 1891
Amara (Bradytus) elegantula Tschitschérine, 1899
Amara (Xenocelia) sikkimensis Andrewes, 1930
Amara bhutanensis Hieke, 2011
Amblops piceus Andrewes, 1931
Amphimenes piceus Andrewes, 1931
Anthracus skalei Jaeger, 2015
Bradycellus (Tachycellus) anchomenoides Bates, 1873
Calleida (Callidiola) doriae Bates, 1892
Chlaenius (Amblygenius) bengalensis Chaudoir, 1856
Chlaenius (Amblygenius) ducalis Chaudoir, 1876
Chlaenius praefectus ducalis Chaudoir, 1876
Chlaenius (Amblygenius) quadricolor (Olivier, 1790)⇒ Chlaenius bengalensis Chaudoir, 1856
Chlaenius (Amblygenius) trachys Andrewes, 1923
Chlaenius (Amblygenius) wittmerianus Mandl, 1978
Chlaenius (Chlaeniellus) fraterculus Maindron, 1899
Chlaenius fraterculus atupus Andrewes, 1923
Chlaenius (Chlaeniostenus) circumdatus Brullé, 1835
Chlaenius circumdatus circumdatus Brullé, 1835
Chlaenius circumdatus subviridulus
Chlaenius (Chlaeniostenus) punctatostriatus Chaudoir, 1856
Chlaenius (Lissauchenius) rufifemoratus MacLeay, 1825
Chlaenius rufifemoratus bhutanensis Mandl, 1978
Chlaenius (Nectochlaenius) dureli Maindron, 1899
Chlaenius (Ocybatus) bioculatus Chaudoir, 1856
Chlaenius (Paralissauchenius) jaegeri (Kirschenhofer, 2002)⇒ Chlaenius berndjaegeri Kirschenhofer, 2008
Chlaenius (Wilmernstus) laetus Fabricius, 1794
Chlaenius hamatus Dejean, 1831
Chlaenius hamifer Chaudoir, 1856
Chydaeus (Chydaeus) andrewesi Schauberger, 1932
Chydaeus andrewesi andrewesi Schauberger, 1932
Chydaeus (Chydaeus) bedeli Tschitschérine, 1897
Chydaeus bedeli interjectus Kataev & Sch..., 2001
Chydaeus (Chydaeus) bhutanensis Kataev & J.S..., 2001
Chydaeus (Chydaeus) semenowi Tschitschérine, 1899
Deltocolpodes bhutanensis Morvan, 2002
Dendrocellus coelestinus Klug, 1834
Dendrocellus geniculata Klug, 1834
Dendrocellus nepalensis Hope, 1831
Dicranoncus philippinensis Jedlička, 1935
Dinodes (Pachydinodes) hamatus Eschscholtz, 1829
Dinodes (Pachydinodes) hamifer Chaudoir, 1856
Dioryche indochinensis Bates, 1889
Dioryche lucidula Dejean, 1829
Dioryche melanauges Andrewes, 1922
Dromius (Klepterus) bhutanensis Mateu, 1977
Dromius (Klepterus) indicus Andrewes, 1923
Dromius (Klepterus) wittmeri Mateu, 1977
Harpalus (Pseudoophonus) indicus Bates, 1891
Harpalus indicus indicus
Harpalus (Zangoharpalus) praticola Bates, 1891
Lebia (Lebia) andrewesi Jedlička, 1933
Lebia (Lebia) karenia Bates, 1892
Lebia (Poecilothais) gressoria Chaudoir, 1871
Lepcha gogonasensis Morvan, 1982
Lesticus (Lesticus) cupricollis Pouillaude, 1914
Lesticus (Lesticus) harmandi Tschitschérine, 1900
Lesticus (Lesticus) wittmeri Morvan, 1980
Lesticus (Triplogenius) desgodinsi Tschitschérine, 1894
Lioptera brevicornis Heller, 1903
Lioptera oberthueri Heller, 1903
Meleagros sikkimensis Andrewes, 1923
Microlestes asiaticus Mateu, 1971
Microlestes demessus Andrewes, 1923
Microlestes demessus demessus Andrewes, 1923
Nepalorthogonius micheli Tian & Deuve, 2016
Nirmala odelli Andrewes, 1930
Onycholabis melitopus Bates, 1892
Orthogonius dureli Tian & Deuve, 2005
Orthogonius himalaya Tian & Deuve, 2006
Orthogonius himalayicus Tian & Deuve, 2005
Orthogonius longiphallus Tian & Deuve, 2005
Orthogonius opacus Schmidt-Gobel, 1846
Oxycentrus scabericollis N.Ito, 2013
Parena obscura Mateu, 1977
Parena quadrisignata Mateu, 1977
Philorhizus adoxus Andrewes, 1923
Physodera dejeani Eschscholtz, 1829
Platymetopus flavilabris Fabricius, 1798
Platynus (Batenus) praedator Andrewes, 1930
Platynus (Batenus) rougemonti Morvan, 1996
Prakasha amariformis Bates, 1892
Pristosia amaroides Putzeys, 1877
Psychristus (Psychristus) amicorum Jaeger, 1997
Psychristus (Psychristus) andrewesi Jaeger, 1997
Psychristus (Psychristus) discretus Andrewes, 1930
Pterostichus (Bothriopterus) aeneocupreus Fairmaire, 1887
Pterostichus (Bothriopterus) bhutanensis (Morvan, 1980)⇒ Pterostichus bhunetansis Davies, 2004
Pterostichus (Unitrichus) brevilama Straneo, 1989
Pterostichus (Unitrichus) championi Andrewes, 1926
Pterostichus (Unitrichus) dorjulensis Morvan, 1978
Pterostichus (Unitrichus) gagates (Hope, 1831)⇒ Pterostichus strenuus Panzer, 1797
Pterostichus gagates gagates Hope, 1831
Pterostichus (Unitrichus) gwervaenus Morvan, 1995
Pterostichus (Unitrichus) hedkeinekus Morvan, 1995
Pterostichus (Unitrichus) imitatus Morvan, 1978
Pterostichus (Unitrichus) lugi Morvan, 1982
Pterostichus (Unitrichus) nepalensis Straneo, 1977
Pterostichus (Unitrichus) pseudoplatysma Straneo, 1982
Pterostichus (Unitrichus) wittmeri Morvan, 1978
Pterostichus bhunetansis Davies, 2004
Pterostichus infissus Andrewes, 1937
Scalidion hilare Schmidt-Goebel, 1846
Sericoda quadripunctata DeGeer, 1774
Stenolophus (Egadroma) lucidus Dejean, 1829
Stenolophus (Egadroma) smaragdulus (Fabricius, 1798)⇒ Egadroma subrobustus Schauberger, 1938
Stenolophus subrobustus Schauberger, 1938
Steropanus infissus Andrewes, 1937
Syntomus fuscomaculatus Motschulsky, 1844
Trichotichnus (Bottchrus) birmanicus (Bates, 1892)⇒ Trichotichnus batesi Csiki, 1932
Trichotichnus (Bottchrus) glabellus Andrewes, 1930
Trigonotoma oberthueri Tschitschérine, 1894
Xestagonum bhutanense Morvan, 1982
Xestagonum bhutanense bhutanense Morvan, 1982
Xestagonum brancuccii J. Schmidt, 1997
Xestagonum brancuccii brancuccii
Xestagonum brancuccii connexum J.Schmidt, 1997
Xestagonum connexum J.Schmidt, 1997
Xestagonum convexicolle J. Schmidt, 1997
Xestagonum immarginatum J. Schmidt, 1997
Xestagonum immarginatum immarginatum J.Schmidt, 1997
Xestagonum immarginatum pelelaense J.Schmidt, 1997
Xestagonum immarginatum pelelaensis
Xestagonum marginale Morvan, 1999
Xestagonum marginalis Morvan, 1999
Xestopus bhutanensis Morvan, 1979
Xestopus cordicollis Morvan, 1979
Xestopus walteri Morvan, 1978

Subnodes of Harpalinae with occurrences in Bhutan
SupertribeChlaeniitae
Bonelli, 1810
2
18
Genera
Species

Taxom

SupertribeDryptitae
Bonelli, 1810
1
3
Genus
Species

Taxom

SupertribeHarpalitae
Erwin, 1985
11
22
Genera
Species

Taxom

SupertribeLebiitae
Bonelli, 1810
12
19
Genera
Species

Taxom

SupertribePlatynitae
Bonelli, 1810
11
20
Genera
Species

Taxom

SupertribePterostichitae
Bonelli, 1810
7
30
Genera
Species

Taxom

TribeOrthogoniini
Schaum, 1857
2
6
Genera
Species

Taxom



Links, sources and further information
Harpalinae at artfakta.se
Harpalinae at dyntaxa.se
Harpalinae at itis.gov