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Checklist for Subfamily Harpalinae Bonelli, 1810
and region Morocco (North Africa)


Class
 		Insecta
Linnaeus, 1758
Insects, True insects		414
3689
10159		Families
Genera
Species		 
Taxom
 

Subclass
 		Pterygota
Lang, 1888
Winged insects		411
3685
10153		Families
Genera
Species		 
Taxom
 

Infraclass
 		Neoptera
van der Wulp, 1890
Modern Wing-folding insects		400
3638
10055		Families
Genera
Species		 
Taxom
 

Superorder
 		Holometabola
Heider, 1889
Holometabolous Insects		251
2823
8311		Families
Genera
Species		 
Taxom
 

Order
 		Coleoptera
Linnaeus, 1758
Beetles, Coleopterans		73
1340
4610		Families
Genera
Species		 
Taxom
 

Suborder
 		Adephaga
Schellenberg, 1806
Adephagans, Ground Beetles and Water Beetles		5
177
844		Families
Genera
Species		 
Taxom
 

Superfamily
 		Caraboidea
Latreille, 1802
1
154
798		Family
Genera
Species		 
Taxom
 
FamilyCarabidae
Latreille, 1802
Carabid beetles, Carabids, Ground Beetles, Tiger Beetles		154
798		Genera
Species		 
Taxom
 
SubfamilyHarpalinae
Bonelli, 1810
89
448		Genera
Species		 
Taxom
 


Genera for Morocco
89 Genera with 448 Species
#ABCDEGHILMOPSTZ
Abacetus Dejean, 18242SpeciesTaxom
Acinopus Dejean, 184013SpeciesTaxom
Acupalpus Latreille, 188412SpeciesTaxom
Afrotarus Jeannel, 19491SpeciesTaxom
Agonum Bonelli, 184110SpeciesTaxom
Amara Bonelli, 1828Seed-Eating Ground Beetles53SpeciesTaxom
Amblycoleus Chaudoir, 18611SpeciesTaxom
Amblystomus Erichson, 18415SpeciesTaxom
Anchomenus Bonelli, 02SpeciesTaxom
Anisodactylus Dejean, 18017SpeciesTaxom
Anthia Weber, 17924SpeciesTaxom
Anthracus Motschulsky, 18121SpeciesTaxom
Apristus Chaudoir, 18461SpeciesTaxom
Atlantomasoreus Mateu, 19142SpeciesTaxom
Badister Clairville, 18443SpeciesTaxom
Bleusea Bedel, 18961SpeciesTaxom
Bradycellus Erichson, 18127SpeciesTaxom
Calathus Bonelli, 199917SpeciesTaxom
Calodromius Reitter, 19333SpeciesTaxom
Cardiomera Bassi, 18341SpeciesTaxom
Carterus Dana, 18377SpeciesTaxom
Chlaenius Bonelli, 1820Vivid Metallic Ground Beetle19SpeciesTaxom
Cryptophonus Brandmayr & ..., 18291SpeciesTaxom
Cymindis Latreille, 182531SpeciesTaxom
Daptus Fischer von ..., 18231SpeciesTaxom
Demetrias Bonelli, 17581SpeciesTaxom
Diachromus Erichson, 17581SpeciesTaxom
Dicheirotrichus Jacquelin du..., 18293SpeciesTaxom
Dinodes Bonet, 18312SpeciesTaxom
Diplocheila Brullé, 18311SpeciesTaxom
Ditomus Dana, 17921SpeciesTaxom
Dixus Billberg, 17954SpeciesTaxom
Dregus Motschulsky, 18581SpeciesTaxom
Dromius Bonelli, 19634SpeciesTaxom
Drypta Latreille, 17902SpeciesTaxom
Eocarterus Stichel, 19593SpeciesTaxom
Eucolliuris Liebke, 18643SpeciesTaxom
Graniger Motschulsky, 18592SpeciesTaxom
Graphipterus Latreille, 18599SpeciesTaxom
Harpalus Latreille, 192541SpeciesTaxom
Heteracantha Brullé, 18341SpeciesTaxom
Idiomelas Tschitschérine, 18291SpeciesTaxom
Ifridytes Deuve & Quei..., 19941SpeciesTaxom
Laemostenus Bonelli, 201235SpeciesTaxom
Lebia Latreille, 1966colorful foliage ground beetle14SpeciesTaxom
Licinus Latreille, 17904SpeciesTaxom
Lionychus Wissmann, 18251SpeciesTaxom
Lonchosternus LaFerté-Séne..., 18462SpeciesTaxom
Masoreus Dejean, 18139SpeciesTaxom
Mesolestes Schatzmayr, 18296SpeciesTaxom
Metadromius Bedel, 18684SpeciesTaxom
Microlestes Schmidt-Goebel, 194116SpeciesTaxom
Odontocarus Solier, 18261SpeciesTaxom
Odotoncarus Solier, 18261SpeciesTaxom
Oedesis Motschulsky, 18592SpeciesTaxom
Olisthopus Dejean, 18283SpeciesTaxom
Oodes Bonelli, 17922SpeciesTaxom
Ophonus Dejean, 183117SpeciesTaxom
Orthomus Chaudoir, 194119SpeciesTaxom
Panagaeus Latreille, 17581SpeciesTaxom
Pangus Dejean, 18181SpeciesTaxom
Paradromius Fowler, 190010SpeciesTaxom
Paranchus Lindroth, 17961SpeciesTaxom
Parazuphium Jeannel, 198211SpeciesTaxom
Parophonus Ganglbauer, 18384SpeciesTaxom
Philorhizus Hope, 19637SpeciesTaxom
Platyderus Stephens, 201711SpeciesTaxom
Platytarus Fairmaire, 19336SpeciesTaxom
Plochionus Dejean, 17751SpeciesTaxom
Poecilus Bonelli, 185821SpeciesTaxom
Polistichus Bonelli, 17851SpeciesTaxom
Pseudomasoreus Desbrochers ..., 18541SpeciesTaxom
Pseudoophonus Motschulsky, 17742SpeciesTaxom
Pseudotrechus Rosenhauer, 18561SpeciesTaxom
Pterostichus Bonelli, 1933Woodland Ground Beetle19SpeciesTaxom
Scybalicus Schaum, 18291SpeciesTaxom
Selina Motschulsky, 18581SpeciesTaxom
Singilis Rambur, 195410SpeciesTaxom
Somotrichus Seidlitz, 18311SpeciesTaxom
Sphodrus Clairville, 17581SpeciesTaxom
Stenolophus Dejean, 1781Seedcorn Beetle7SpeciesTaxom
Stomis Clairville, 17962SpeciesTaxom
Syntomus Hope, 182514SpeciesTaxom
Tetragonoderus Dejean, 19051SpeciesTaxom
Trymosternus Chaudoir, 19636SpeciesTaxom
Tschitscherinellus Csiki, 18252SpeciesTaxom
Typsiharpalus Tschitschérine, 19251SpeciesTaxom
Zabrus Clairville, 190712SpeciesTaxom
Zuphium Latreille, 18747SpeciesTaxom


Species list for Morocco
448 Species and 147 Subspecies
Abacetus (Abacetillus) antoinei Straneo, 1951
Abacetus (Astigis) salzmanni Germar, 1824
Acinopus (Acinopus) boiteli Alluaud, 1930
Acinopus (Acinopus) dianae Schatzmayr, 1943
Acinopus (Acinopus) grassator Coquerel, 1859
Acinopus (Acinopus) haroldi Schaum, 1863
Acinopus (Acinopus) laevipennis Fairmaire, 1859
Acinopus (Acinopus) liouvillei Puel, 1934
Acinopus (Acinopus) sabulosus Fabricius, 1792
Acinopus sabulosus lucidus Alluaud, 1930
Acinopus sabulosus peyerimhoffi Antoine, 1959
Acinopus sabulosus sabulosus Fabricius, 1792
Acinopus (Acmastes) haroldii Schaum, 1863
Acinopus (Oedemalicus) gutturosus Buquet, 1840
Acinopus (Oedematicus) gutturosus Buquet, 1840
Acupalpus (Acupalpus) brunnipes Sturm, 1825
Acupalpus (Acupalpus) dubius Schilsky, 1888
Acupalpus (Acupalpus) elegans Dejean, 1829
Acupalpus (Acupalpus) luteatus (Duftschmid, 1812)⇒ Acupalpus dubius Schilsky, 1888
Acupalpus (Acupalpus) maculatus Schaum, 1860
Acupalpus (Acupalpus) zaerensis Antoine, 1922
Acupalpus (Ancylostria) mediterraneus Csiki, 1932
Acupalpus (Tachistodes) cantabricus Brulerie, 1868
Acupalpus (Tachistodes) ibericus Jaeger, 1988
Acupalpus (Tachistodes) lucasi Gaubil, 1849
Acupalpus (Tachistodes) notatus Mulsant & Rey, 1861
Acupalpus (Tachistodes) oliveirae Reitter, 1884
Afrotarus alluaudi Jeannel, 1949
Agonum (Agonum) atlantis Antoine, 1957
Agonum (Agonum) marginatum Linnaeus, 1758
Agonum (Agonum) nigrum (Dejean, 1828)⇒ Agonum fuliginosum Panzer, 1809
Agonum (Agonum) numidicum Lucas, 1846
Agonum (Europhilus) fuliginosum Panzer, 1809
Agonum (Europhilus) persecretum Antoine, 1941
Agonum (Olisares) fulgidicolle Erichson, 1841
Agonum (Olisares) lugens Duftschmid, 1812
Agonum (Olisares) viridicupreum Goeze, 1777
Agonum viridicupreum fulgidicolle Erichson, 1841
Amara (Acorius) metallescens C. Zimmermann, 1831
Amara (Amara) aenea De Geer, 1774Common sun beetle
Amara (Amara) anthobia A. Villa & G..., 1833Italy harp ground beetle
Amara (Amara) eurynota Panzer, 1796
Amara (Amara) famelica C. Zimmermann, 1832Early sunshiner
Amara (Amara) familiaris Duftschmid, 1812Bareyed goby
Amara (Amara) lucida Duftschmid, 1812
Amara (Amara) similata Gyllenhal, 1810
Amara (Amara) subconvexa Putzeys, 1865
Amara (Amarocelia) erratica Duftschmid, 1812
Amara (Amathitis) rufescens (Dejean, 1829)⇒ Amara aulica Panzer, 1796
Amara rufescens schismatica Antoine, 1957
Amara rufescens shismatica Antoine, 1953
Amara (Atlantocnemis) colasi Paulian & Vi..., 1939
Amara (Atlantocnemis) liouvillei Antoine, 1936
Amara (Atlantocnemis) mairei Peyerimhoff, 1922
Amara (Atlantocnemis) notha Antoine, 1936
Amara (Camptocelia) affinis (Dejean, 1828)⇒ Amara erratica Duftschmid, 1812
Amara (Camptocelia) brevis Dejean, 1828
Amara (Camptocelia) coiffaiti Jeanne, 1981
Amara (Camptocelia) corpulenta Putzeys, 1866
Amara (Camptocelia) cottyi Coquerel, 1859
Amara cottyi cottyi Coquerel, 1859
Amara cottyi schrammi Antoine, 1941
Amara (Camptocelia) gottelandi Antoine, 1931
Amara gottelandi gottelandi Antoine, 1931
Amara gottelandi rhatica Antoine, 1936
Amara (Camptocelia) kocheri Antoine, 1933
Amara (Camptocelia) otini Antoine, 1938
Amara (Camptocelia) pueli Antoine, 1923
Amara (Camptocelia) rotundata Dejean, 1828
Amara (Celia) brevis Dejean, 1828
Amara (Celia) cottyi Coquerel, 1859
Amara (Celia) cursitans Zimmermann, 1832
Amara (Celia) fervida Coquerel, 1859
Amara fervida perobtecta Antoine, 1953
Amara (Celia) montana (Dejean, 1828)⇒ Amara spectabilis Schaum, 1858
Amara quenseli silvicola Zimm., 1832
Amara (Celia) rufoaenea (Dejean, 1828)⇒ Amara cursitans Zimmermann, 1832
Amara (Celia) simplex Dejean, 1828
Amara (Celia) sollicita Pantel, 1888
Amara (Curtonotus) aulica Panzer, 1796Princely harp ground beetle
Amara (Harpalodema) maindroni Bedel, 1907
Amara (Paracelia) rufoaenea Dejean, 1828
Amara (Paracelia) simplex Dejean, 1828
Amara simplex simplex Dejean, 1828
Amara (Percosia) perabdita Antoine, 1941
Amara (Xenocelia) atlantis Antoine, 1925
Amara (Xenocelia) fusca (Dejean, 1828)⇒ Amara quenseli silvicola Zimm., 1832
Amara (Zezea) hypsophila Antoine, 1953
Amara (Zezea) kulti Fassati, 1947
Amara reflexicollis reflexicollis Motschulsky, 1845
Amara (Zezea) rufipes (Dejean, 1828)⇒ Amara erratica Duftschmid, 1812
Amblycoleus platyderus Chaudoir, 1861
Amblystomus algirinus Reitter, 1887
Amblystomus escorialensis Gautier Des ..., 1866
Amblystomus mauritanicus Dejean, 1829
Amblystomus metallescens Dejean, 1829
Amblystomus niger (Heer, 1841)⇒ Amblystomus nigrinus Csiki, 1931
Anchomenus (Anchomenus) dorsalis Pontoppidan, 1763
Anchomenus dorsalis dorsalis
Anisodactylus (Anisodactylus) antoinei Puel, 1931
Anisodactylus (Anisodactylus) binotatus Fabricius, 1787Two-marked harp ground beetle
Anisodactylus (Hexatrichus) poeciloides Stephens, 1828Saltmarsh Short-spur
Anisodactylus poeciloides poeciloides Stephens, 1828
Anisodactylus (Hexatrichus) virens Dejean, 1829
Anisodactylus virens winthemi Dejean, 1831
Anisodactylus (Pseudhexatrichus) heros Fabricius, 1801
Anthia (Thermophilum) sexmaculata Fabricius, 1787
Anthia sexmaculata marginata Latreille, 1823
Anthia sexmaculata sexmaculata Fabricius, 1787
Anthia (Thermophilum) venator Fabricius, 1792
Anthracus consputus Duftschmid, 1812
Apristus striatipennis Lucas, 1846
Atlantomasoreus (Groneri) orbipennis Bedel, 1904
Atlantomasoreus desertorum Escalera, 1914
Badister (Badister) meridionalis Puel, 1925
Badister peltatus peltatus Panzer, 1796
Badister (Baudia) collaris (Motschulsky, 1844)⇒ Badister peltatus peltatus Panzer, 1796
Bleusea deserticola Bedel, 1896
Bradycellus (Bradycellus) otini Antoine, 1959Stone-dwelling cave beetle
Bradycellus (Triliarthrus) brevitarsis Normand, 1946
Bradycellus (Triliarthrus) distinctus Dejean, 1829
Bradycellus (Triliarthrus) lusitanicus Dejean, 1829
Bradycellus (Triliarthrus) ruficollis Stephens, 1828
Bradycellus (Triliarthrus) sharpi Joy, 1912
Bradycellus (Triliarthrus) verbasci Duftschmid, 1812
Calathus (Bedelinus) circumseptus Germar, 1824
Calathus (Calathus) fuscipes Goeze, 1777Dark-footed harp ground beetle
Calathus fuscipes algiricus Gautier Des ..., 1866
Calathus (Calathus) opacus Lucas, 1846
Calathus (Neocalathus) melanocephalus Linnaeus, 1758
Calathus melanocephalus antoinei Puel, 1939
Calathus melanocephalus furvus Tschitschérine, 1895
Calathus melanocephalus melanocephalus Linné, 1758
Calathus melanocephalus paphlagonicus Maran, 1935
Calathus melanocephalus purkynei Maran, 1935
Calathus (Neocalathus) mollis Marsham, 1802
Calathus mollis atticus Gautier Des ..., 1867
Calathus (Neocalathus) opaculus LeConte, 1854
Calathus (Neocalathus) rhaticus Antoine, 1941
Calathus (Neocalathus) semisericeus Fairmaire, 1879
Calathus (Neocalathus) solieri Bassi, 1834
Calathus atitari Novoa, 1999
Calodromius bifasciatus Dejean, 1825
Calodromius mayeti Bedel, 1907
Calodromius sphex Andrewes, 1933
Cardiomera genei Bassi, 1834
Carterus (Microcarterus) gracilis Rambur, 1837
Carterus (Microcarterus) microcephalus Rambur, 1837Saint Paul cave beetle
Carterus (Pristocarterus) dama P.Rossi, 1792
Carterus (Pristocarterus) fulvipes Latreille, 1817
Carterus (Pristocarterus) gilvipes Piochard de ..., 1873
Carterus (Pristocarterus) interceptus Dejean, 1830
Carterus (Pristocarterus) rotundicollis Rambur, 1837
Chlaenius lithophilus lithophilus Say, 1823
Chlaenius (Chlaeniellus) circumcinctus Say, 1830Prickly scorpion's-tail
Chlaenius (Chlaeniellus) olivieri Crotch, 1871
Chlaenius (Chlaeniostenus) angustatus (Dejean, 1831)⇒ Chlaenius gotschii Chaudoir, 1846
Chlaenius (Chlaenites) spoliatus P. Rossi, 1792
Chlaenius spoliatus spoliatus P. Rossi, 1792
Chlaenius festivus velutinus Duftschmid, 1812
Chlaenius (Chlaenius) velutinus Duftschmid, 1812
Chlaenius (Nectochlaenius) canariensis Dejean, 1831
Chlaenius canariensis seminitidus Chaudoir, 1856
Chlaenius (Trichochlaenius) chrysocephalus P. Rossi, 1790
Chlaenius (Trichochlaenius) cyaneus Brullé, 1835
Chlaenius (Trichochlaenius) gotschii Chaudoir, 1846
Chlaenius (Trichochlaenius) infantulus Chaudoir, 1876
Chlaenius (Trichochlaenius) milleti Antoine, 1932
Chlaenius (Trichochlaenius) nebrioides Antoine, 1961
Chlaenius (Trichochlaenius) virens (Rambur, 1837)⇒ Chlaenius circumcinctus Say, 1830
Chlaenius conformis Dejean, 1831
Chlaenius decipiens L.Dufour, 1820
Cryptophonus tenebrosus Dejean, 1829
Cymindis (Cymindis) alluaudi (Antoine, 1939)⇒ Afrotarus alluaudi Jeannel, 1949
Cymindis alluaudi alluaudi Antoine, 1939
Cymindis alluaudi brevepilosa Antoine, 1941
Cymindis alluaudi lecerfi Antoine, 1963
Cymindis alluaudi tisselitensis Antoine, 1963
Cymindis (Cymindis) cobosi Mateu, 1965
Cymindis (Cymindis) ephippium Escalera, 1914
Cymindis (Cymindis) favieri Fairmaire, 1859
Cymindis favieri atlasica Escalera, 1914
Cymindis favieri debduensis Antoine, 1941
Cymindis favieri favieri Lucas, 1859
Cymindis favieri mediatlantis Antoine, 1939
Cymindis favieri tazekensis Antoine, 1941
Cymindis (Cymindis) gottelandi Paulian & Vi..., 1939
Cymindis (Cymindis) hookeri Bates, 1875
Cymindis hookeri hookeri Bates, 1875
Cymindis hookeri merdelensis Antoine, 1963
Cymindis (Cymindis) kocheri Antoine, 1939
Cymindis (Cymindis) pilosipennis Escalera, 1922
Cymindis (Cymindis) rhatica Antoine, 1936
Cymindis (Cymindis) setifeensis Lucas, 1842
Cymindis setifeensis agoniothorax Bedel, 1906
Cymindis setifeensis antoinei Schatzmayr, 1944
Cymindis setifeensis laevistriata Lucas, 1846
Cymindis setifeensis leucophthalma Lucas, 1842
Cymindis setifeensis setifeensis Lucas, 1842
Cymindis (Menas) bedeli Tschitschérine, 1897
Cymindis (Tarulus) bedeli Tschitschérine, 1897
Cymindis (Tarulus) discophora Chaudoir, 1873
Cymindis (Tarulus) suturalis Dejean, 1825
Cymindis suturalis suturalis Dejean, 1825
Daptus vittatus Fischer von ..., 1823
Demetrias (Demetrias) atricapillus Linnaeus, 1758
Diachromus germanus Linnaeus, 1758
Dicheirotrichus (Pelagophilus) punicus Bedel, 1899
Dicheirotrichus punicus punicus Bedel, 1899
Dicheirotrichus (Trichocellus) obsoletus Dejean, 1829
Dinodes (Dinodes) decipiens (L. Dufour, 1820)⇒ 
Dinodes (Pachydinodes) conformis Dejean, 1831
Diplocheila (Isorembus) aegyptiaca Dejean, 1831
Ditomus tricuspidatus Fabricius, 1792
Dixus capito Audinet-Serv..., 1821
Dixus clypeatus P. Rossi, 1790
Dixus interruptus Fabricius, 1775
Dixus sphaerocephalus (Olivier, 1795)⇒ Dixus interruptus Fabricius, 1775
Dregus glebalis Coquerel, 1858
Dromius (Dromius) angustus (Brullé, 1834)⇒ Microlestes linearis LeConte, 1851
Dromius (Dromius) chobauti Puel, 1924
Dromius (Dromius) meridionalis Dejean, 1825
Dromius (Dromius) simplicior Antoine, 1963
Drypta (Deserida) distincta P.Rossi, 1792
Drypta (Drypta) dentata P.Rossi, 1790
Eocarterus (Baeticocarus) tazekensis Antoine, 1959
Eocarterus tazekensis rifensis Cobos, 1961
Eocarterus tazekensis tazekensis Antoine, 1959
Eucolliuris oliverii Buquet, 1864
Eucolliuris olivieri Buquet, 1864
Eucolliuris olivierii Buquet, 1864
Graniger cordicollis Audinet-Serv..., 1821
Graniger femoralis Coquerel, 1859
Graphipterus exclamationis Fabricius, 1792
Graphipterus exclamationis exclamationis Fabricius, 1792
Graphipterus exclamationis rolphii Fairmaire, 1867
Graphipterus luctuosus Dejean, 1825
Graphipterus luctuosus peletieri Laporte, 1840
Graphipterus peletieri Laporte, 1840
Graphipterus reymondi (Antoine, 1953)⇒ Graphipterus serrator valdanii Guérin-Mén..., 1859
Graphipterus serrator Forskal, 1775
Graphipterus serrator valdanii Guérin-Mén..., 1859
Harpalus (Acardystus) attenuatus Stephens, 1828
Harpalus (Amblystus) latus Linnaeus, 1758
Harpalus (Amblystus) rubripes Dufischmid, 1812Bareyed goby
Harpalus (Artabas) punctatostriatus (Dejean, 1829)⇒ Harpalus rubefactus rubefactus Bates, 1873
Harpalus (Cryptophonus) fulvus Dejean, 1829
Harpalus (Cryptophonus) tenebrosus (Dejean, 1829)⇒ 
Harpalus tenebrosus tenebrosus
Harpalus (Euharpalops) fraternus LeConte, 1852
Harpalus (Harpaloderus) neglectus Serville, 1821
Harpalus neglectus alluaudi Antoine, 1922
Harpalus (Harpaloderus) sulphuripes Germar, 1824
Harpalus sulphuripes goudotii Dejean, 1829
Harpalus (Harpalus) angustitarsis Reitter, 1887
Harpalus (Harpalus) dissitus Antoine, 1931
Harpalus (Harpalus) mairei Peyerimhoff, 1928
Harpalus (Harpalus) oblitus (Dejean, 1829)⇒ Harpalus fraternus LeConte, 1852
Harpalus oblitus patruelis Dejean, 1829
Harpalus (Harpalus) wohlberedti Emden & Scha..., 1932
Harpalus (Lasioharpalus) cupreus Dejean, 1829
Harpalus cupreus asaphus Antoine, 1941
Harpalus (Lasioharpalus) dimidiatus P. Rossi, 1790
Harpalus (Lasioharpalus) distinguendus Duftschmid, 1812
Harpalus distinguendus distinguendus Duftschmid, 1812
Harpalus rubefactus rubefactus Bates, 1873
Harpalus (Pheuginus) serripes Quensel, 1806
Harpalus serripes serripes Quensel, 1806
Harpalus (Pheuginus) tardus Panzer, 1796
Harpalus (Plectralidus) cardoni Antoine, 1922
Harpalus (Plectralidus) fuscicornis Ménétriés, 1832
Harpalus (Plectralidus) lateralis Dejean, 1829
Harpalus (Plectralidus) lethierryi Reiche, 1860
Harpalus lethierryi azrouanus Emden & Scha..., 1932
Harpalus (Plectralidus) microthorax Motschulsky, 1849
Harpalus (Plectralidus) numidicus Bedel, 1893
Harpalus (Plectralidus) rufitarsoides Schauberger, 1934
Harpalus rufitarsoides diversipennis Schatzmayr, 1943
Harpalus rufitarsoides rufitarsoides Schauberger, 1934
Harpalus (Plectralidus) siculus Dejean, 1829
Harpalus (Pseudoophonus) griseus (Panzer, 1796)⇒ Pseudoophonus griseus Panzer, 1797
Harpalus (Pseudoophonus) rufipes (DeGeer, 1774)⇒ 
Harpalus azruensis Antoine, 1925
Heteracantha depressa Brullé, 1834
Idiomelas (Egaploa) crenulatus Dejean, 1829
Ifridytes mateui Deuve & Quei..., 1994
Laemostenus (Antisphodrus) ambroggii Antoine, 1961
Laemostenus (Antisphodrus) malhommei Antoine, 1954
Laemostenus malhommei fongondi Casale, 1988
Laemostenus malhommei malhommei Antoine, 1954
Laemostenus (Antisphodrus) vagabundus Antoine, 1961
Laemostenus (Antisphodrus) villardi Antoine, 1948
Laemostenus (Atlantosphodrus) jeannelianus Paulian & Vi..., 1939
Laemostenus (Ceuthosthenes) ajmasianus Bolivar y Pi..., 1922
Laemostenus (Ceuthosthenes) berberus Mateu, 1975
Laemostenus (Ceuthosthenes) kolbi Coiffait, 1972
Laemostenus (Ceuthosthenes) lyauteyi Alluaud, 1923
Laemostenus (Ceuthosthenes) mauritanicus Dejean, 1828
Laemostenus mauritanicus mauritanicus Dejean, 1828
Laemostenus (Ceuthosthenes) mogadoricus Escalera, 1922
Laemostenus (Ceuthosthenes) schrammi Antoine, 1938
Laemostenus (Laemostenus) barbarus Lucas, 1846
Laemostenus (Laemostenus) complanatus Dejean, 1828Bynoe's goby
Laemostenus (Pristonychus) algerinus Gory, 1833
Laemostenus algerinus algerinus (Gory, 1833)⇒ Laemostenus algerinus Gory, 1833
Laemostenus (Pristonychus) cadilhaci Antoine, 1951
Laemostenus (Pristonychus) sericeus Fischer von ..., 1824
Laemostenus (Rhysosphodrus) deneveui Fairmaire, 1859
Laemostenus (Sphodroides) aelleni Antoine, 1952
Laemostenus (Sphodroides) anomostriatus Antoine, 1954
Laemostenus (Sphodroides) atlanticus Escalera, 1913
Laemostenus atlanticus atlanticus Escalera, 1913
Laemostenus atlanticus deserticus Mateu, 1975
Laemostenus atlanticus macrops Antoine, 1951
Laemostenus (Sphodroides) favieri Fairmaire, 1868
Laemostenus (Sphodroides) gomezi Bolivar y Pi..., 1922
Laemostenus (Sphodroides) melillensis Escalera, 1922
Laemostenus (Sphodroides) punctatostriatus Fairmaire, 1859
Laemostenus (Sphodroides) susicus Antoine, 1941
Laemostenus (Sphodroides) vidali Antoine, 1938
Laemostenus aurouxi Casale & Comas, 2012
Lebia (Lamprias) cyanocephala Linnaeus, 1758Blue plunderer
Lebia cyanocephala numidica Lucas, 1846
Lebia (Lamprias) fulvicollis (Fabricius, 1792)⇒ Lebia pubipennis Dufour, 1820
Lebia fulvicollis fulvicollis Fabricius, 1792
Lebia (Lamprias) pubipennis Dufour, 1820
Lebia (Lamprias) rufipes Dejean, 1825
Lebia (Lebia) cruxminor Linnaeus, 1758
Lebia cruxminor cruxminor Linnaeus, 1758
Lebia cruxminor pilosula Antoine, 1963
Lebia (Lebia) scapularis (Geoffroy, 1785)⇒ Lebia solea Hentz, 1830
Lebia scapularis scapularis Geoffroy, 1785
Lebia (Lebia) solea Hentz, 1830
Lebia (Lebia) trimaculata (Villers, 1789)⇒ 
Lebia (Stephana) turkestanica Jedlička, 1966
Licinus (Licinus) lindbergi Antoine, 1936
Licinus (Licinus) punctatulus Fabricius, 1792
Licinus punctatulus punctatulus
Licinus (Licinus) silphoides (P. Rossi, 1790)⇒ Licinus punctatulus Fabricius, 1792
Lionychus (Paralionychus) albonotatus Dejean, 1825
Lonchosternus hispanicus Dejean, 1826
Lonchosternus mauritanicus Lucas, 1846
Masoreus (Masoreus) affinis Chaudoir, 1843
Masoreus affinis affinis Chaudoir, 1843
Masoreus orientalis orientalis Dejean, 1828
Masoreus (Masoreus) saharensis Mateu, 1984
Masoreus (Masoreus) wetterhallii Gyllenhal, 1813
Masoreus wetterhallii antoinei Mateu, 1984
Masoreus wetterhallii axillaris Küster, 1852
Masoreus wetterhallii testaceus Lucas, 1846
Masoreus wetterhallii wetterhallii Gyllenhal, 1813
Mesolestes (Mesolestes) accentifer Raffray, 1873
Mesolestes (Neomesolestes) caecus Antoine, 1941
Mesolestes (Neomesolestes) pueli Antoine, 1923
Mesolestes (Neomesolestes) sicardi Bedel, 1918
Mesolestes (Pseudomesolestes) scapularis Dejean, 1830
Mesolestes scapularis scapularis Dejean & Boi..., 1829
Metadromius ephippiatus Fairmaire, 1884
Metadromius lateplagiatus Fairmaire, 1873
Metadromius myrmidon Fairmaire, 1859
Metadromius ramburii Piochard de ..., 1868
Microlestes abeillei Brisout, 1885
Microlestes abeillei brisouti Holdhaus, 1912
Microlestes angusteforcipatus Antoine, 1941
Microlestes corticalis L. Dufour, 1820
Microlestes gallicus Holdhaus, 1912
Microlestes gomerensis Harold Lindberg, 1953
Microlestes ibericus Holdhaus, 1912
Microlestes levipennis Lucas, 1846
Microlestes levipennis levipennis Lucas, 1846
Microlestes luctuosus Holdhaus, 1904
Microlestes luctuosus chobauti Jeannel, 1942
Microlestes mauritanicus Lucas, 1846
Microlestes mirei Mateu, 1953
Microlestes negrita Wollaston, 1854
Microlestes numidicus Normand, 1941
Microlestes phenax Antoine, 1941
Odontocarus cephalotes Dejean, 1826
Odotoncarus cephalotes Dejean, 1826
Oedesis tomentosus Dejean, 1831
Oedesis villosulus Reiche, 1859
Olisthopus elongatus Wollaston, 1854
Olisthopus fuscatus Dejean, 1828
Olisthopus hispanicus Dejean, 1828
Oodes helopioides Fabricius, 1792
Oodes helopioides helopioides Fabricius, 1792
Ophonus (Hesperophonus) pumilio Dejean, 1829
Ophonus (Hesperophonus) rotundatus Dejean, 1829
Ophonus (Hesperophonus) subquadratus (Dejean, 1829)⇒ 
Ophonus (Incisophonus) incisus Dejean, 1829
Ophonus (Metophonus) berberus Antoine, 1925
Ophonus (Metophonus) cordatus Duftschmid, 1812
Ophonus (Metophonus) cunii Fairmaire, 1880
Ophonus (Metophonus) ferrugatus Reitter, 1902
Ophonus (Metophonus) puncticeps (Stephens, 1828)⇒ Ophonus puncticollis Paykull, 1798
Ophonus (Metophonus) puncticollis Paykull, 1798
Ophonus (Metophonus) rufibarbis Fabricius, 1792
Ophonus (Metophonus) sahlbergianus Lutshnik, 1922
Ophonus (Metophonus) subsinuatus Rey, 1886
Ophonus (Metophonus) syriacus Dejean, 1829
Ophonus (Ophonus) ardosiacus Lutshnik, 1922
Ophonus (Ophonus) opacus Dejean, 1829
Ophonus (Ophonus) quadricollis Dejean, 1831
Orthomus (Orthomus) barbarus (Dejean, 1828)⇒ Orthomus berytensis Reiche & Saulcy, 1855
Orthomus barbarus kocheri Mateu, 1955
Orthomus barbarus lacouri Antoine, 1941
Orthomus (Orthomus) berytensis Reiche & Saulcy, 1855
Orthomus (Orthomus) dimorphus Antoine, 1933
Orthomus dimorphus antoinei Mateu, 1955
Orthomus dimorphus dimorphus Antoine, 1933
Orthomus (Orthomus) lacouri Antoine, 1941
Orthomus lacouri haroldi Pupier & Coulon, 2013
Orthomus lacouri kocheri Mateu, 1955
Orthomus lacouri lacouri Antoine, 1941
Orthomus (Orthomus) maroccanus Chaudoir, 1873
Orthomus (Orthomus) rubicundus Coquerel, 1859
Orthomus rubicundus rubicundus Coquerel, 1859
Orthomus rubicundus tazekensis Antoine, 1941
Orthomus (Orthomus) starkei Wrase & Jeanne, 2005
Orthomus (Orthomus) tazekensis Antoine, 1941
Orthomus tazekensis rifensis Wrase & Jeanne, 2005
Orthomus tazekensis tazekensis Antoine, 1941
Panagaeus (Panagaeus) cruxmajor Linnaeus, 1758Crucifix ground beetle
Pangus scaritides Sturm, 1818
Paradromius (Manodromius) bermejoi Mateu, 1956
Paradromius (Manodromius) kocheri Antoine, 1963
Paradromius (Manodromius) linearis Olivier, 1795
Paradromius linearis linearis Olivier, 1795
Paradromius (Manodromius) proderus Fairmaire, 1880
Paradromius (Manodromius) puncticeps Bedel, 1907
Paradromius (Manodromius) sublinearis Escalera, 1914
Paradromius (Manodromius) vagepictus Fairmaire, 1875
Paradromius vagepictus vagepictus Fairmaire, 1875
Paradromius (Trichodromius) pilifer Bedel, 1900
Paranchus albipes Fabricius, 1796White-legged harp ground beetle
Parazuphium (Neozuphium) damascenum Fairmaire, 1897
Parazuphium damascenum damascenum Fairmaire, 1897
Parazuphium (Parazuphium) aguilerai Andújar, He..., 2011
Parazuphium (Parazuphium) angustioculum (Hurka, 1982)⇒ Parazuphium angustioculus Hurka, 1982
Parazuphium (Parazuphium) baeticum K. Daniel & ..., 1898
Parazuphium baeticum baeticum K. Daniel & ..., 1898
Parazuphium baeticum mauretaniae Hurka, 1982
Parazuphium (Parazuphium) maroccanum Antoine, 1963
Parazuphium (Parazuphium) punicum K. Daniel & ..., 1898
Parazuphium (Parazuphium) vaucheri Vauloger De ..., 1898
Parazuphium angustioculus Hurka, 1982
Parophonus (Ophonomimus) hirsutulus Dejean, 1829
Parophonus (Parophonus) antoinei Schauberger, 1931
Parophonus (Parophonus) hespericus Jeanne, 1985
Parophonus (Parophonus) hispanus Rambur, 1838
Philorhizus (Philorhizus) crucifer Lucas, 1846
Philorhizus crucifer crucifer Lucas, 1846
Philorhizus (Philorhizus) insignis Lucas, 1846
Philorhizus (Philorhizus) melanocephalus Dejean, 1825Frillfin gobies
Philorhizus (Philorhizus) notatus Stephens, 1827
Philorhizus (Philorhizus) vectensis Rye, 1873
Philorhizus berberus Antoine, 1963
Platyderus (Eremoderus) insignitus Bedel, 1902
Platyderus insignitus insignitus
Platyderus insignitus presahariensis Lagar Mascaro, 1978
Platyderus (Eremoderus) languidus Reiche & Saulcy, 1855
Platyderus (Platyderus) calathoides Dejean, 1828
Platyderus calathoides calathoides Dejean, 1828
Platyderus (Platyderus) ellipticus Bedel, 1902
Platyderus (Platyderus) gregarius Reiche, 1862
Platyderus (Platyderus) otini Antoine, 1941
Platyderus (Platyderus) solissimus Antoine, 1939
Platyderus claudejeannei Machard, 2017
Platytarus bufo Fabricius, 1801
Platytarus faminii Dejean, 1826
Platytarus faminii faminii Dejean, 1826
Platytarus gracilis Dejean, 1831
Platytarus gracilis gracilis Dejean, 1831
Platytarus gracilis occidentalis Antoine, 1933
Plochionus (Plochionus) pallens Fabricius, 1775Ground beetle
Poecilus (Ancholeus) baeticus Rambur, 1838
Poecilus baeticus gharbensis Alluaud, 1927
Poecilus (Ancholeus) crenatus (Dejean, 1828)⇒ Poecilus gisellae gisellae Csiki, 1930
Poecilus (Ancholeus) gisellae Csiki, 1930
Poecilus gisellae freyanus Uyttenboogaart, 1940
Poecilus (Ancholeus) nitidus Dejean, 1828
Poecilus (Ancholeus) wollastoni Wollaston, 1854
Poecilus (Carenostylus) purpurascens Dejean, 1828
Poecilus purpurascens debduensis Antoine, 1962
Poecilus purpurascens purpurascens Dejean, 1828
Poecilus (Macropoecilus) lepidus Leske, 1785
Poecilus (Parapedius) decipiens Wallt, 1835
Poecilus decipiens baudoni Antoine, 1957
Poecilus decipiens cupripennis Fairmaire, 1852
Poecilus decipiens decipiens Waltl, 1835
Poecilus decipiens mediatlantis Antoine, 1951
Poecilus decipiens rotroui Antoine, 1957
Poecilus (Poecilus) crenulatus Dejean, 1828
Poecilus crenulatus mauritanicus Dejean, 1828
Poecilus (Poecilus) quadricollis Dejean, 1828
Poecilus (Poecilus) vicinus Levrat, 1858
Polistichus connexus Geoffroy, 1785
Pseudomasoreus canigoulensis Fairmaire & ..., 1854
Pseudoophonus (Pseudoophonus) griseus Panzer, 1797
Pseudoophonus (Pseudoophonus) rufipes De Geer, 1774
Pseudotrechus mutilatus Rosenhauer, 1856
Pterostichus aterrimus attenuatus Chevrolat, 1840
Pterostichus aterrimus surcoufi Antoine, 1933
Pterostichus (Argutor) vernalis Panzer, 1796Spring Blackclock
Pterostichus (Cheporus) transversalis Duftschmid, 1812
Pterostichus (Lagarus) vernalis Panzer, 1796
Pterostichus (Lamenius) caudicalis Say, 1823
Pterostichus (Leptoferonia) idahoae Csiki, 1930Key goby
Pterostichus anthracinus anthracinus Illiger, 1798
Pterostichus (Melanius) aterrimus Herbst, 1784
Pterostichus (Melanius) elongatus (Duftschmid, 1812)⇒ Pterostichus idahoae Csiki, 1930
Pterostichus (Melanius) nigrita (Paykull, 1790)⇒ Pterostichus anthracinus anthracinus Illiger, 1798
Pterostichus (Omaseus) aterrimus (Herbst, 1784)⇒ Pterostichus transversalis Duftschmid, 1812
Pterostichus (Pseudomaseus) nigrita Paykull, 1790
Pterostichus nigrita nigrita
Pterostichus (Rhagadus) thorectes Bates, 1873
Pterostichus (Steropus) ebenus Quensel, 1806
Pterostichus (Unitrichus) carri Angus, Galla..., 2009
Pterostichus crenatus (Duftschmid, 1812)⇒ Pterostichus vernalis Panzer, 1796
Pterostichus rifensis Antoine, 1933
Scybalicus oblongiusculus Dejean, 1829
Selina westermanni Motschulsky, 1858
Singilis (Singilis) bedeli Escalera, 1913
Singilis (Singilis) bicolor (Rambur, 1837)⇒ Singilis taschkensis Jedlicka, 1961
Singilis (Singilis) centralis Antoine, 1963
Singilis (Singilis) mauritanicus Lucas, 1846
Singilis (Singilis) soror Rambur, 1837
Singilis soror melillensis Escalera, 1914
Singilis soror riffensis Mateu, 1954
Singilis soror soror Rambur, 1837
Singilis melillensis Escalera, 1914
Singilis pardoi Mateu, 1954
Somotrichus unifasciatus Dejean, 1831Ground beetle
Sphodrus leucophthalmus Linnaeus, 1758
Stenolophus (Egadroma) marginatus Dejean, 1829
Stenolophus (Stenolophus) abdominalis Géné, 1836
Stenolophus abdominalis abdominalis Gené, 1836
Stenolophus (Stenolophus) mixtus Herbst, 1784
Stenolophus (Stenolophus) proximus Dejean, 1829
Stenolophus (Stenolophus) skrimshiranus Stephens, 1828
Stenolophus (Stenolophus) teutonus Schrank, 1781
Stomis (Stomis) pumicatus Panzer, 1796
Stomis pumicatus pumicatus Panzer, 1796
Syntomus barbarus Puel, 1938
Syntomus bedeli Puel, 1924
Syntomus foveatus Geoffroy, 1785
Syntomus foveolatus Dejean, 1831
Syntomus fuscomaculatus Motschulsky, 1844
Syntomus impressus Dejean, 1825
Syntomus impressus decorus Bedel, 1913
Syntomus impressus impressus Dejean, 1825
Syntomus lateralis Motschulsky, 1855
Syntomus lateralis lateralis Motschulsky, 1855
Syntomus lateralis schrammi Antoine, 1941
Syntomus montanus Bedel, 1913
Syntomus obscuroguttatus Duftschmid, 1812
Syntomus pallipes Dejean, 1825
Tetragonoderus (Tetragonoderus) extremus Bedel, 1905
Trymosternus colombati Antoine, 1934
Trymosternus colombati colombati Antoine, 1934
Trymosternus colombati pecoudi Antoine, 1934
Trymosternus truncatus Rambur, 1837
Trymosternus truncatus antoinei Mateu, 1952
Trymosternus truncatus zaianus Antoine, 1963
Tschitscherinellus cordatus Dejean, 1825
Tschitscherinellus cordatus cordatus Dejean, 1825
Typsiharpalus azruensis Antoine, 1925
Zabrus (Aulacozabrus) distinctus Lucas, 1842
Zabrus (Craspedozabrus) tenuestriatus Fairmaire, 1884
Zabrus (Polysitus) deflexicollis Fairmaire, 1880
Zabrus (Polysitus) guildensis Alluaud, 1932
Zabrus guildensis alluaudi Antoine, 1957
Zabrus guildensis guildensis Alluaud, 1932
Zabrus guildensis lacouri Antoine, 1957
Zabrus guildensis rifensis Antoine, 1941
Zabrus (Polysitus) maroccanus Reiche, 1864
Zabrus (Zabrus) ignavus Csiki, 1907
Zabrus ignavus baudoni Antoine, 1957
Zabrus ignavus ignavus (Csiki, 1907)⇒ Zabrus ignavus Csiki, 1907
Zuphium (Zuphium) numidicum Lucas, 1846
Zuphium (Zuphium) olens P.Rossi, 1790
Zuphium (Zuphium) testaceum Klug, 1832
Zuphium ciliatum Vauloger De ..., 1897
Zuphium cilicium Peyron, 1858
Zuphium fuscum Gory, 1831
Zuphium microphthalmum Putzeys, 1874

Subnodes of Harpalinae with occurrences in Morocco
SupertribeAnthiitae
Bonelli, 1813
1
2		Genus
Species
Taxom

SupertribeChlaeniitae
Bonelli, 1810
8
30		Genera
Species
Taxom

SupertribeCtenodactylitae
Castelnau, 1834
1
1		Genus
Species
Taxom

SupertribeDryptitae
Bonelli, 1810
4
18		Genera
Species
Taxom

SupertribeHarpalitae
Erwin, 1985
31
132		Genera
Species
Taxom

SupertribeLebiitae
Bonelli, 1810
27
112		Genera
Species
Taxom

SupertribePlatynitae
Bonelli, 1810
10
63		Genera
Species
Taxom

SupertribePterostichitae
Bonelli, 1810
7
90		Genera
Species
Taxom



Links, sources and further information
Harpalinae at artfakta.se
Harpalinae at dyntaxa.se
Harpalinae at itis.gov