AntsAntsArachnidsArachnids
BeesBeesBeetlesBeetles
BookliceBookliceBugsBugs
BumblebeesBumblebeesButterfliesButterflies
CicadasCicadasCrane fliesCrane flies
DragonfliesDragonfliesEarwigsEarwigs
FliesFliesIsopodsIsopods
LocustsLocustsMantisesMantises
MayfliesMayfliesMosquitoesMosquitoes
MyriapodsMyriapodsNet-wingedNet-winged
Plant-parasiticPlant-parasiticScorpion fliesScorpion flies
TermitesTermitesWaspsWasps
INSECTOIDGERENGDARK LIGHT
Home Arthropoda Imprint Contact
TAXFAMGENSPESUB
Checklist for Subfamily Harpalinae Bonelli, 1810
and region Portugal (Iberian Peninsula)


Class
 
Insecta
Linnaeus, 1758
Insects, True insects
546
5554
13778
Families
Genera
Species
 
Taxom
 

Subclass
 
Pterygota
Lang, 1888
Winged insects
542
5540
13730
Families
Genera
Species
 
Taxom
 

Infraclass
 
Neoptera
van der Wulp, 1890
Modern Wing-folding insects
512
5443
13568
Families
Genera
Species
 
Taxom
 

Superorder
 
Holometabola
Heider, 1889
Holometabolous Insects
333
4250
11221
Families
Genera
Species
 
Taxom
 

Order
 
Coleoptera
Linnaeus, 1758
Beetles, Coleopterans
86
1550
4587
Families
Genera
Species
 
Taxom
 

Suborder
 
Adephaga
Schellenberg, 1806
Adephagans, Ground Beetles and Water Beetles
6
196
938
Families
Genera
Species
 
Taxom
 

Superfamily
 
Caraboidea
Latreille, 1802
1
157
793
Family
Genera
Species
 
Taxom
 
FamilyCarabidae
Latreille, 1802
Carabid beetles, Carabids, Ground Beetles, Tiger Beetles
157
793
Genera
Species
 
Taxom
 
SubfamilyHarpalinae
Bonelli, 1810
94
426
Genera
Species
 
Taxom
 


Genera for Portugal
94 Genera with 426 Species
#ABCDEGHLMNOPSTZ
Abacetus Dejean, 18241SpeciesTaxom
Acinopus Dejean, 18403SpeciesTaxom
Acupalpus Latreille, 182512SpeciesTaxom
Agonum Bonelli, 175824SpeciesTaxom
Agra Fabricius, 2000Elegant canopy beetles1SpeciesTaxom
Amara Bonelli, 1831Seed-Eating Ground Beetles47SpeciesTaxom
Amblystomus Erichson, 18617SpeciesTaxom
Anchomenus Bonelli, 03SpeciesTaxom
Anisodactylus Dejean, 19318SpeciesTaxom
Anthia Weber, 17931SpeciesTaxom
Apristus Chaudoir, 19801SpeciesTaxom
Badister Clairville, 18443SpeciesTaxom
Barrymooreana Baehr, 1997
0SpeciesTaxom
Bradycellus Erichson, 181213SpeciesTaxom
Calathus Bonelli, 187332SpeciesTaxom
Calleida Latreille & ..., 18481SpeciesTaxom
Callistus Bonelli, 17752SpeciesTaxom
Calodromius Reitter, 18762SpeciesTaxom
Carterus Dana, 18376SpeciesTaxom
Cedrorum Borges & Ser..., 19932SpeciesTaxom
Chlaenius Bonelli, 1932Vivid Metallic Ground Beetle10SpeciesTaxom
Cryptophonus Brandmayr & ..., 18291SpeciesTaxom
Cymindis Latreille, 175819SpeciesTaxom
Daptus Fischer von ..., 18231SpeciesTaxom
Demetrias Bonelli, 18244SpeciesTaxom
Diachromus Erichson, 17581SpeciesTaxom
Dicheirotrichus Jacquelin du..., 18292SpeciesTaxom
Dicrochile Guérin-Ménev..., 19311SpeciesTaxom
Dinodes Bonet, 19326SpeciesTaxom
Ditomus Dana, 17923SpeciesTaxom
Dixus Billberg, 17954SpeciesTaxom
Dromius Bonelli, 18254SpeciesTaxom
Drypta Latreille, 17902SpeciesTaxom
Eurygnathus Wollaston, 19223SpeciesTaxom
Graniger Motschulsky, 18592SpeciesTaxom
Gynandromorphus Dejean, 18062SpeciesTaxom
Harpalus Latreille, 181250SpeciesTaxom
Laemostenus Bonelli, 17843SpeciesTaxom
Lebia Latreille, 1966colorful foliage ground beetle16SpeciesTaxom
Licinus Latreille, 07SpeciesTaxom
Lionychus Wissmann, 18122SpeciesTaxom
Lonchosternus LaFerté-Séne..., 18261SpeciesTaxom
Machozetus Chaudoir, 1850
0SpeciesTaxom
Masoreus Dejean, 18132SpeciesTaxom
Mesolestes Schatzmayr, 18292SpeciesTaxom
Metadromius Bedel, 18682SpeciesTaxom
Microlestes Schmidt-Goebel, 181212SpeciesTaxom
Neocolpodes Jeannel, 19351SpeciesTaxom
Nesacinopus Tschitschérine, 18601SpeciesTaxom
Nesarpalus Bedel, 18974SpeciesTaxom
Odacantha Paykull, 17661SpeciesTaxom
Odontocarus Solier, 18261SpeciesTaxom
Odotoncarus Solier, 18261SpeciesTaxom
Oedesis Motschulsky, 18591SpeciesTaxom
Olisthopus Dejean, 179811SpeciesTaxom
Oodes Bonelli, 18881SpeciesTaxom
Ophonus Dejean, 182819SpeciesTaxom
Orthomus Chaudoir, 195514SpeciesTaxom
Panagaeus Latreille, 17581SpeciesTaxom
Pangus Dejean, 18181SpeciesTaxom
Paradromius Fowler, 17955SpeciesTaxom
Paranchus Lindroth, 17961SpeciesTaxom
Parazuphium Jeannel, 19634SpeciesTaxom
Parhypates Motschulsky, 19001SpeciesTaxom
Parophonus Ganglbauer, 17906SpeciesTaxom
Percus Bonelli, 18203SpeciesTaxom
Perigona Laporte, 18311SpeciesTaxom
Philorhizus Hope, 19059SpeciesTaxom
Platyderus Stephens, 186811SpeciesTaxom
Platytarus Fairmaire, 18263SpeciesTaxom
Poecilus Bonelli, 182813SpeciesTaxom
Polistichus Bonelli, 17851SpeciesTaxom
Pseudanchomenus Tarnier, 18601SpeciesTaxom
Pseudomasoreus Desbrochers ..., 18541SpeciesTaxom
Pseudoophonus Motschulsky, 17743SpeciesTaxom
Pterostichus Bonelli, 1874Woodland Ground Beetle24SpeciesTaxom
Scybalicus Schaum, 18291SpeciesTaxom
Singilis Rambur, 19612SpeciesTaxom
Siopelus Murray, 19261SpeciesTaxom
Sphodrus Clairville, 17581SpeciesTaxom
Stenolophus Dejean, 1781Seedcorn Beetle9SpeciesTaxom
Stomis Clairville, 17962SpeciesTaxom
Styracoderus Chaudoir, 18651SpeciesTaxom
Syntomus Hope, 17618SpeciesTaxom
Synuchus Gyllenhal, 17982SpeciesTaxom
Trechicus LeConte, 18311SpeciesTaxom
Trichotichnus A. Morawitz, 18121SpeciesTaxom
Trymosternus Chaudoir, 18251SpeciesTaxom
Tschitscherinellus Csiki, 18252SpeciesTaxom
Typsiharpalus Tschitschérine, 18661SpeciesTaxom
Zabrus Clairville, 177710SpeciesTaxom
Zargus Wollaston, 18544SpeciesTaxom
Zuphioides Ball & Shpeley, 18791SpeciesTaxom
Zuphium Latreille, 17902SpeciesTaxom


Species list for Portugal
426 Species and 113 Subspecies
Abacetus (Astigis) salzmanni Germar, 1824
Acinopus (Acinopus) picipes Olivier, 1795
Acinopus (Oedemalicus) gutturosus Buquet, 1840
Acinopus (Oedematicus) gutturosus Buquet, 1840
Acupalpus (Acupalpus) brunnipes Sturm, 1825
Acupalpus (Acupalpus) dubius Schilsky, 1888
Acupalpus (Acupalpus) elegans Dejean, 1829
Acupalpus (Acupalpus) flavicollis Sturm, 1825
Acupalpus (Acupalpus) maculatus Schaum, 1860
Acupalpus (Acupalpus) meridianus Linnaeus, 1761Midday harp ground beetle
Acupalpus (Acupalpus) parvulus Sturm, 1825
Acupalpus (Tachistodes) cantabricus Brulerie, 1868
Acupalpus (Tachistodes) ibericus Jaeger, 1988
Acupalpus (Tachistodes) notatus Mulsant & Rey, 1861
Acupalpus (Tachistodes) oliveirae Reitter, 1884
Acupalpus brunneipes Sturm, 1825
Agonum (Agonum) carbonarium Dejean, 1828
Agonum carbonarium carbonarium Dejean, 1828
Agonum carbonarium jeannei Aubry, 1970
Agonum (Agonum) marginatum Linnaeus, 1758
Agonum (Agonum) muelleri Herbst, 1784Müller'S Harp Ground Beetle
Agonum muelleri muelleri Herbst, 1784
Agonum (Agonum) nigrum (Dejean, 1828)⇒ Agonum fuliginosum Panzer, 1809
Agonum (Agonum) numidicum Lucas, 1846
Agonum (Europhilus) anchomenoides Randall, 1838
Agonum (Europhilus) fuliginosum Panzer, 1809
Agonum gracile gracile Sturm, 1824
Agonum (Europhilus) thoreyi Dejean, 1828
Agonum (Olisares) dolens C.R. Sahlberg, 1827
Agonum (Olisares) duftschmidi J. Schmidt, 1994
Agonum (Olisares) emarginatum (Gyllenhal, 1827)⇒ Agonum thoreyi Dejean, 1828
Agonum (Olisares) ericeti Panzer, 1809
Agonum (Olisares) lugens Duftschmid, 1812
Agonum (Olisares) permoestum Puel, 1938
Agonum (Olisares) sexpunctatum (Linnaeus, 1758)⇒ Agonum muelleri Herbst, 1784
Agonum (Olisares) versutum Sturm, 1824
Agonum (Olisares) viduum Panzer, 1796
Agonum (Olisares) viridicupreum Goeze, 1777
Agonum viridicupreum viridicupreum Goeze, 1777
Agonum (Punctagonum) sexpunctatum Linnaeus, 1758
Agra yola Erwin, 2000
Amara (Acorius) metallescens C. Zimmermann, 1831
Amara (Amara) aenea De Geer, 1774Common sun beetle
Amara (Amara) anthobia A. Villa & G..., 1833Italy harp ground beetle
Amara (Amara) convexior Stephens, 1828
Amara (Amara) eurynota Panzer, 1796
Amara (Amara) famelica C. Zimmermann, 1832Early sunshiner
Amara (Amara) familiaris Duftschmid, 1812Bareyed goby
Amara (Amara) impuncticollis Say, 1823
Amara (Amara) littorea C.C. Thomson, 1857
Amara (Amara) lucida Duftschmid, 1812
Amara (Amara) ovata (Fabricius, 1792)⇒ Amara tricuspidata Dejean, 1831
Amara (Amara) similata Gyllenhal, 1810
Amara (Amara) subconvexa Putzeys, 1865
Amara (Amarocelia) erratica Duftschmid, 1812
Amara (Bradytus) apricaria Paykull, 1790Exposed sun beetle
Amara (Bradytus) consularis Duftschmid, 1812
Amara (Bradytus) fulva (O.F. Muller, 1776)⇒ Amara apricaria Paykull, 1790
Amara (Camptocelia) affinis (Dejean, 1828)⇒ Amara erratica Duftschmid, 1812
Amara (Camptocelia) arcuata (Putzeys, 1865)⇒ Amara impuncticollis Say, 1823
Amara arcuata castiliana Hieke, 1983
Amara (Camptocelia) brevis Dejean, 1828
Amara (Camptocelia) corpulenta Putzeys, 1866
Amara cottyi cottyi Coquerel, 1859
Amara (Camptocelia) eximia Dejean, 1828
Amara (Camptocelia) rotundata Dejean, 1828
Amara (Celia) bifrons Gyllenhal, 1810Two-faced harp ground beetle
Amara (Celia) brevis Dejean, 1828
Amara (Celia) eximia Dejean, 1828
Amara (Celia) fervida Coquerel, 1859
Amara fervida fervida Coquerel, 1859
Amara (Celia) montana (Dejean, 1828)⇒ Amara spectabilis Schaum, 1858
Amara (Celia) pseudobrunnea Lindroth, 1968Leopard-spotted goby
Amara (Curtonotus) jacobina LeConte, 1855Calabura
Amara (Curtonotus) pennsylvanica Hayward, 1908
Amara (Leironotus) glabrata (Dejean, 1828)⇒ Amara longula LeConte, 1855
Amara (Paracelia) superans Wollaston, 1854
Amara (Percosia) equestris Duftschmid, 1812
Amara equestris equestris Duftschmid, 1812
Amara equestris zabroides Dejean, 1828
Amara (Zezea) fulvipes (Audinet-Serv..., 1821)⇒ Amara pennsylvanica Hayward, 1908
Amara (Zezea) kulti Fassati, 1947
Amara (Zezea) longula LeConte, 1855Calabura
Amara (Zezea) plebeja (Gyllenhal, 1810)⇒ Amara equestris equestris Duftschmid, 1812
Amara reflexicollis reflexicollis Motschulsky, 1845
Amara (Zezea) rufipes (Dejean, 1828)⇒ Amara erratica Duftschmid, 1812
Amara (Zezea) scitula Zimmermann, 1832
Amara (Zezea) tricuspidata Dejean, 1831
Amblystomus algirinus Reitter, 1887
Amblystomus escorialensis Gautier Des ..., 1866
Amblystomus mauritanicus Dejean, 1829
Amblystomus metallescens Dejean, 1829
Amblystomus niger (Heer, 1841)⇒ Amblystomus nigrinus Csiki, 1931
Amblystomus nigrinus Csiki, 1931
Amblystomus raymondi Gautier Des ..., 1861
Anchomenus (Anchodemus) cyaneus Dejean, 1828
Anchomenus (Anchomenus) dorsalis Pontoppidan, 1763
Anchomenus dorsalis dorsalis
Anisodactylus (Anisodactylus) binotatus Fabricius, 1787Two-marked harp ground beetle
Anisodactylus (Anisodactylus) californicus Dejean, 1829Checked goby
Anisodactylus (Anisodactylus) nemorivagus Duftschmid, 1812Heath Short-spur
Anisodactylus (Hexatrichus) virens Dejean, 1829
Anisodactylus virens virens Dejean, 1829
Anisodactylus (Pseudanisodactylus) signatus Panzer, 1796
Anisodactylus (Pseudhexatrichus) heros Fabricius, 1801
Anisodactylus (Spongopus) hispanus Puel, 1931
Anthia nimrod nimrod Fabricius, 1793
Apristus europaeus Mateu, 1980
Badister (Badister) meridionalis Puel, 1925
Badister peltatus peltatus Panzer, 1796
Badister (Baudia) collaris (Motschulsky, 1844)⇒ Badister peltatus peltatus Panzer, 1796
Bradycellus (Atlantocellus) assingi Wrase & Jaeger, 1996
Bradycellus (Atlantocellus) excultus Wollaston, 1854
Bradycellus (Atlantocellus) maderensis Mateu, 1958
Bradycellus (Atlantocellus) wollastoni Wrase & Jaeger, 1996Sensabush cave beetle
Bradycellus (Stenocellus) californicus LeConte, 1857
Bradycellus (Stenocellus) rupestris Say, 1823Rich Mountain cave beetle
Bradycellus (Triliarthrus) chavesi Alluaud, 1919Ground-beetle
Bradycellus (Triliarthrus) distinctus Dejean, 1829
Bradycellus (Triliarthrus) harpalinus Audinet-Serv..., 1821European harp ground beetle
Bradycellus (Triliarthrus) lusitanicus Dejean, 1829
Bradycellus (Triliarthrus) ruficollis Stephens, 1828
Bradycellus (Triliarthrus) sharpi Joy, 1912
Bradycellus (Triliarthrus) verbasci Duftschmid, 1812
Calathus (Amphigynus) rotundicollis Dejean, 1828
Calathus (Amphyginus) rotundatus Jaequelin du..., 1857
Calathus (Amphyginus) rotundicollis Dejean, 1828
Calathus (Bedelinus) circumseptus Germar, 1824
Calathus (Calathus) brevis Gautier Des ..., 1866
Calathus (Calathus) hispanicus Gautier Des ..., 1866
Calathus hispanicus dejeani Ganglbauer, 1891
Calathus (Calathus) luctuosus (Latreille, 1804)⇒ Calathus hispanicus dejeani Ganglbauer, 1891
Calathus (Iberocalathus) rotundatus Jaequelin du..., 1857
Calathus rotundatus estrelensis Jeanne, 1976
Calathus rotundatus rotundatus Jacquelin du..., 1857
Calathus (Lauricalathus) carvalhoi Serrano & Bo..., 1986Ground Beetle
Calathus (Lauricalathus) colasianus Mateu, 1970
Calathus (Lauricalathus) complanatus Dejean, 1828
Calathus complanatus complanatus Dejean, 1828
Calathus complanatus vandeli Colas & Mateu, 1958
Calathus (Lauricalathus) fimbriatus Wollaston, 1858
Calathus (Lauricalathus) pecoudi Colas, 1938
Calathus (Lauricalathus) vividus Fabricius, 1801
Calathus (Neocalathus) asturiensis Vuillefroy, 1866
Calathus (Neocalathus) cinctus Motschulsky, 1850
Calathus (Neocalathus) erratus C.R.Sahlberg, 1827
Calathus erratus erratus C.R. Sahlberg, 1827
Calathus (Neocalathus) extensicollis Putzeys, 1873Ground beetle
Calathus (Neocalathus) lundbladi Colas, 1938Ground beetle
Calathus (Neocalathus) mollis Marsham, 1802
Calathus mollis mollis Marsham, 1802
Calathus (Neocalathus) subfuscus Wollaston, 1865
Calathus (Neocalathus) vicenteorum Schatzmayr, 1937Ground beetle
Calathus (Trichocalathus) minutus Gautier Des ..., 1866
Calathus (Trichocalathus) obliteratus Wollaston, 1865
Calathus heydeni (Putzeys, 1873)⇒ Calathus brevis Gautier Des ..., 1866
Calleida (Calleida) punctata LeConte, 1848Goldspot goby
Callistus lunatus Fabricius, 1775
Callistus lunatus lunatus Fabricius, 1775
Calodromius bifasciatus Dejean, 1825
Calodromius putzeysi Paulino de O..., 1876
Carterus (Microcarterus) gracilis Rambur, 1837
Carterus (Microcarterus) microcephalus Rambur, 1837Saint Paul cave beetle
Carterus (Pristocarterus) fulvipes Latreille, 1817
Carterus (Pristocarterus) gilvipes Piochard de ..., 1873
Carterus (Pristocarterus) interceptus Dejean, 1830
Carterus (Pristocarterus) rotundicollis Rambur, 1837
Cedrorum azoricus azoricus Borges & Ser..., 1993
Cedrorum azoricus caveirensis Borges & Ser..., 1993
Chlaenius (Chlaeniellus) olivieri Crotch, 1871
Chlaenius (Epomis) circumscriptus Duftschmid, 1812
Chlaenius (Lithochlaenius) solitarius Say, 1823Chlaenius Ground Beetle
Chlaenius (Trichochlaenius) chrysocephalus P. Rossi, 1790
Chlaenius (Trichochlaenius) infantulus Chaudoir, 1876
Chlaenius decipiens L.Dufour, 1820
Chlaenius dives Dejean, 1826
Chlaenius dives dives
Chlaenius dives gallaecianus Chaudoir, 1876
Chlaenius dives lusitanicus H.Wagner, 1932
Cryptophonus tenebrosus Dejean, 1829
Cymindis (Cymindis) axillaris (Fabricius, 1794)⇒ Cymindis scapularis Schaum, 1857
Cymindis axillaris axillaris Fabricius, 1794
Cymindis axillaris meridionalis Dejean, 1825
Cymindis (Cymindis) humeralis (Geoffroy, 1785)⇒ 
Cymindis (Cymindis) scapularis Schaum, 1857
Cymindis scapularis scapularis Schaum, 1857
Cymindis (Cymindis) setifeensis Lucas, 1842
Cymindis (Menas) miliaris Fabricius, 1801
Cymindis (Tarulus) alternans Rambur, 1837
Cymindis alternans alternans Rambur, 1837
Cymindis alternans vogelii Schaufuss, 1862
Cymindis (Tarulus) discoidea Dejean, 1824
Cymindis (Tarulus) lineata Quensel In S..., 1806
Cymindis (Tarulus) maderae Wollaston, 1857
Cymindis (Tarulus) paivana Wollaston, 1860
Cymindis (Tarulus) portugalica Jedlička, 1946
Cymindis suturalis pseudosuturalis Bedel, 1906
Cymindis (Tarulus) vaporariorum Linnaeus, 1758
Cymindis vaporarium Linné, 1758
Daptus vittatus Fischer von ..., 1823
Demetrias (Aetophorus) imperialis Germar, 1824
Demetrias (Demetrias) atricapillus Linnaeus, 1758
Demetrias (Demetrias) monostigma Samouelle, 1819
Demetrias (Risophilus) imperialis Germar, 1824
Diachromus germanus Linnaeus, 1758
Dicheirotrichus (Trichocellus) obsoletus Dejean, 1829
Dicheirotrichus (Trichocellus) pallidus Dejean, 1829
Dicrochile whitei Csiki, 1931
Dinodes (Dinodes) decipiens (L. Dufour, 1820)⇒ 
Dinodes (Dinodes) dives Dejean, 1826
Dinodes dives dives Dejean, 1826
Dinodes dives galaecianus Chaudoir, 1876
Dinodes dives gallaecianus Chaudoir, 1876
Dinodes dives lusitanicus Wagner, 1932
Ditomus calydonius P. Rossi, 1790
Ditomus calydonius calydonius P. Rossi, 1790
Ditomus tricuspidatus Fabricius, 1792
Dixus capito Audinet-Serv..., 1821
Dixus clypeatus P. Rossi, 1790
Dixus interruptus Fabricius, 1775
Dixus sphaerocephalus (Olivier, 1795)⇒ Dixus interruptus Fabricius, 1775
Dromius (Dromius) angustus (Brullé, 1834)⇒ Microlestes linearis LeConte, 1851
Dromius angustus alutaceus Wollaston, 1857
Dromius (Dromius) fenestratus Fabricius, 1794
Dromius (Dromius) meridionalis Dejean, 1825
Drypta (Deserida) distincta P.Rossi, 1792
Drypta (Drypta) dentata P.Rossi, 1790
Eurygnathus latreillei Laporte, 1834
Eurygnathus latreillei latreillei Laporte de C..., 1834
Eurygnathus latreillei wollastoni Cockerell, 1922
Graniger cordicollis Audinet-Serv..., 1821
Graniger femoralis Coquerel, 1859
Gynandromorphus etruscus Quensel, 1806
Gynandromorphus etruscus etruscus Quensel, 1806
Harpalus (Acardystus) attenuatus Stephens, 1828
Harpalus (Acardystus) pygmaeus Dejean, 1829
Harpalus (Amblystus) rubripes Dufischmid, 1812Bareyed goby
Harpalus (Artabas) dispar Dejean, 1829
Harpalus dispar dispar Dejean, 1829
Harpalus (Artabas) punctatostriatus (Dejean, 1829)⇒ Harpalus rubefactus rubefactus Bates, 1873
Harpalus (Cryptophonus) fulvus Dejean, 1829
Harpalus (Cryptophonus) litigiosus (Dejean, 1829)⇒ Cryptophonus litigiosus Dejean, 1829
Harpalus (Cryptophonus) melancholicus (Dejean, 1829)⇒ Cryptophonus melancholicus Dejean, 1829
Harpalus melancholicus melancholicus
Harpalus (Cryptophonus) tenebrosus (Dejean, 1829)⇒ 
Harpalus tenebrosus tenebrosus
Harpalus (Euharpalops) fraternus LeConte, 1852
Harpalus (Harpaloderus) honestus Dufischmid, 1812St. Bees Seed-eater
Harpalus (Harpaloderus) neglectus Serville, 1821
Harpalus neglectus neglectus Audinet-Serv..., 1821
Harpalus (Harpaloderus) rufipalpis Sturm, 1818
Harpalus rufipalpis lusitanicus Schatzmayr, 1943
Harpalus rufipalpis machadoi Jeanne, 1970
Harpalus rufipalpis montanellus Mateu, 1953
Harpalus rufipalpis rufipalpis Sturm, 1818
Harpalus rufipalpis wagneri Schauberger, 1926
Harpalus (Harpaloderus) sulphuripes Germar, 1824
Harpalus sulphuripes sulphuripes Germar, 1824
Harpalus (Harpalus) affinis (Schrank, 1781)⇒ Harpalus serripes ernsti Kataev, 1995
Harpalus (Harpalus) angustitarsis Reitter, 1887
Harpalus (Harpalus) oblitus (Dejean, 1829)⇒ Harpalus fraternus LeConte, 1852
Harpalus oblitus patruelis Dejean, 1829
Harpalus (Lasioharpalus) cupreus Dejean, 1829
Harpalus cupreus asaphus Antoine, 1941
Harpalus cupreus cupreus Dejean, 1829
Harpalus (Lasioharpalus) dimidiatus P. Rossi, 1790
Harpalus (Lasioharpalus) distinguendus Duftschmid, 1812
Harpalus distinguendus distinguendus Duftschmid, 1812
Harpalus rubefactus rubefactus Bates, 1873
Harpalus (Pheuginus) serripes Quensel, 1806
Harpalus serripes ernsti Kataev, 1995
Harpalus serripes serripes Quensel, 1806
Harpalus (Pheuginus) tardus Panzer, 1796
Harpalus (Plectralidus) contemptus Dejean, 1829
Harpalus (Plectralidus) decipiens Dejean, 1829
Harpalus decipiens correiroi Schatzmayr, 1943
Harpalus decipiens decipiens Dejean, 1829
Harpalus (Plectralidus) lateralis Dejean, 1829
Harpalus (Plectralidus) rufitarsoides Schauberger, 1934
Harpalus rufitarsoides diversipennis Schatzmayr, 1943
Harpalus (Pseudoophonus) calceatus (Duftschmid, 1812)⇒ Pseudoophonus calceatus Duftschmid, 1812
Harpalus (Pseudoophonus) griseus (Panzer, 1796)⇒ Pseudoophonus griseus Panzer, 1797
Harpalus (Pseudoophonus) rufipes (DeGeer, 1774)⇒ 
Harpalus (Semiophonus) signaticornis Duftschmid, 1812
Laemostenus (Laemostenus) complanatus Dejean, 1828Bynoe's goby
Laemostenus (Pristonychus) terricola Herbst, 1784Bandy-legged burrow-dweller beetle
Laemostenus terricola terricola Herbst, 1784
Lebia (Lamprias) cyanocephala Linnaeus, 1758Blue plunderer
Lebia cyanocephala cyanocephala Linnaeus, 1758
Lebia (Lamprias) fulvicollis (Fabricius, 1792)⇒ Lebia pubipennis Dufour, 1820
Lebia fulvicollis fulvicollis Fabricius, 1792
Lebia (Lamprias) pubipennis Dufour, 1820
Lebia (Lamprias) rufipes Dejean, 1825
Lebia (Lebia) cruxminor Linnaeus, 1758
Lebia cruxminor cruxminor Linnaeus, 1758
Lebia (Lebia) cyanipennis Dejean, 1831
Lebia (Lebia) marginata Geoffroy, 1785
Lebia (Lebia) scapularis (Geoffroy, 1785)⇒ Lebia solea Hentz, 1830
Lebia scapularis scapularis Geoffroy, 1785
Lebia (Lebia) solea Hentz, 1830
Lebia (Lebia) trimaculata (Villers, 1789)⇒ 
Lebia (Lebia) viridis Say, 1823Flower Lebia Beetle
Lebia (Stephana) turkestanica Jedlička, 1966
Licinus (Licinus) aequatus Audinet-Serv..., 1821
Licinus aequatus aequatus Audinet-Serv..., 1821
Licinus aequatus reymondi Colas, 1949
Licinus (Licinus) peltoides Dejean, 1826
Licinus (Licinus) punctatulus Fabricius, 1792
Licinus punctatulus granulatus Dejean, 1826
Licinus punctatulus punctatulus
Lionychus (Paralionychus) albonotatus Dejean, 1825
Lionychus (Paralionychus) quadrillum Duftschmid, 1812
Lonchosternus hispanicus Dejean, 1826
Masoreus orientalis nobilis Wollaston, 1864
Masoreus (Masoreus) wetterhallii Gyllenhal, 1813
Mesolestes (Pseudomesolestes) scapularis Dejean, 1830
Mesolestes scapularis scapularis Dejean & Boi..., 1829
Metadromius myrmidon Fairmaire, 1859
Metadromius ramburii Piochard de ..., 1868
Microlestes abeillei Brisout, 1885
Microlestes abeillei abeillei Brisout de B..., 1885
Microlestes aljezurensis Ortuño & Oli..., 2012
Microlestes corticalis L. Dufour, 1820
Microlestes linearis LeConte, 1851
Microlestes luctuosus Holdhaus, 1904
Microlestes luctuosus chobauti Jeannel, 1942
Microlestes luctuosus luctuosus Holdhaus, 1904
Microlestes maurus Sturm, 1827
Microlestes maurus maurus Sturm, 1827
Microlestes negrita Wollaston, 1854
Microlestes plagiatus (Duftschmid, 1812)⇒ Microlestes corticalis L. Dufour, 1820
Neocolpodes hylobius Alluaud, 1935
Nesacinopus pelagicus Wollaston, 1860
Nesarpalus (Nesarpalus) cimensis Cockerell, 1922
Nesarpalus cimensis cimensis Cockerell, 1922
Nesarpalus cimensis maderae Schauberger, 1932
Nesarpalus (Nesarpalus) gregarius Fauvel, 1897
Odacantha (Odacantha) melanura Linnaeus, 1766
Odontocarus cephalotes Dejean, 1826
Odotoncarus cephalotes Dejean, 1826
Oedesis villosulus Reiche, 1859
Olisthopus elongatus Wollaston, 1854
Olisthopus ericae Wollaston, 1854
Olisthopus fuscatus Dejean, 1828
Olisthopus hispanicus Dejean, 1828
Olisthopus humerosus Wollaston, 1858
Olisthopus inclavatus Israelson, 1983Ground Beetle
Olisthopus maderensis Wollaston, 1854
Olisthopus maderensis acutangulus Wollaston, 1857
Olisthopus maderensis maderensis Wollaston, 1854
Olisthopus rotundatus Paykull, 1790
Olisthopus rotundatus rotundatus Paykull, 1798
Oodes froggatti W.J.Macleay, 1888
Ophonus (Hesperophonus) azureus Fabricius, 1775
Ophonus (Hesperophonus) longicollis (Rambur, 1838)⇒ Ophonus incisus Dejean, 1829
Ophonus (Hesperophonus) pumilio Dejean, 1829
Ophonus (Hesperophonus) rotundatus Dejean, 1829
Ophonus (Hesperophonus) similis Dejean, 1829
Ophonus (Hesperophonus) subquadratus (Dejean, 1829)⇒ 
Ophonus (Incisophonus) incisus Dejean, 1829
Ophonus (Metophonus) brevicollis (Audinet-Serv..., 1821)⇒ Ophonus melletii Heer, 1837
Ophonus (Metophonus) cordatus Duftschmid, 1812
Ophonus (Metophonus) ferrugatus Reitter, 1902
Ophonus (Metophonus) parallelus (Dejean, 1829)⇒ Ophonus xaxarsi Schauberger, 1928
Ophonus (Metophonus) rufibarbis Fabricius, 1792
Ophonus (Metophonus) sahlbergianus Lutshnik, 1922
Ophonus (Metophonus) subsinuatus Rey, 1886
Ophonus (Ophonus) ardosiacus Lutshnik, 1922
Ophonus (Ophonus) diffinis (Dejean, 1829)⇒ Ophonus ardosiacus Lutshnik, 1922
Ophonus (Ophonus) opacus Dejean, 1829
Ophonus (Ophonus) sabulicola Panzer, 1796
Ophonus (Ophonus) stictus Stephens, 1828Oolite Downy-back
Orthomus (Nesorthomus) bedelianus Lutshnik, 1915
Orthomus (Nesorthomus) berrai F. Battoni, 1987
Orthomus (Nesorthomus) curtus Wollaston, 1854
Orthomus (Nesorthomus) dilaticollis Wollaston, 1854
Orthomus (Nesorthomus) gracilipes Wollaston, 1854
Orthomus (Nesorthomus) lundbladi Jeannel, 1938
Orthomus (Nesorthomus) pecoudi Jeannel, 1943
Orthomus (Nesorthomus) susanae (Serrano & Bo..., 2009)⇒ Orthomus annae Donabauer, 2008
Orthomus (Orthomus) barbarus (Dejean, 1828)⇒ Orthomus berytensis Reiche & Saulcy, 1855
Orthomus barbarus barbarus Dejean, 1828
Orthomus barbarus haligena Wollaston, 1860
Orthomus (Orthomus) berytensis Reiche & Saulcy, 1855
Orthomus (Orthomus) velocissimus Waltl, 1835
Orthomus velocissimus akbensis Mateu, 1955
Panagaeus (Panagaeus) cruxmajor Linnaeus, 1758Crucifix ground beetle
Pangus scaritides Sturm, 1818
Paradromius (Manodromius) insularis Wollaston, 1854
Paradromius insularis insularis Wollaston, 1854
Paradromius insularis oceanicus Wollaston, 1865
Paradromius (Manodromius) linearis Olivier, 1795
Paradromius linearis linearis Olivier, 1795
Paranchus albipes Fabricius, 1796White-legged harp ground beetle
Parazuphium baeticum mauretaniae Hurka, 1982
Parazuphium (Parazuphium) chevrolatii Laporte, 1833
Parazuphium chevrolatii chevrolatii Laporte de C..., 1833
Parazuphium (Parazuphium) maroccanum Antoine, 1963
Parhypates (Parhypates) arctus Tschitschérine, 1900
Parophonus (Ophonomimus) hirsutulus Dejean, 1829
Parophonus (Parophonus) dejeani Csiki, 1932
Parophonus (Parophonus) hespericus Jeanne, 1985
Parophonus (Parophonus) iberiparcus Zaballos & G..., 1991
Parophonus (Parophonus) maculicornis Duftschmid, 1812
Parophonus (Parophonus) mendax P.Rossi, 1790
Percus (Pseudopercus) patruelis L. Dofour, 1820
Percus (Pseudopercus) politus Dejean, 1831
Percus (Pseudopercus) stultus L. Dufour, 1820
Perigona (Trechicus) nigriceps Dejean, 1831Tawny Harp Ground Beetle
Philorhizus (Philorhizus) conicipennis Fauvel, 1905
Philorhizus (Philorhizus) insignis Lucas, 1846
Philorhizus (Philorhizus) melanocephalus Dejean, 1825Frillfin gobies
Philorhizus (Philorhizus) umbratus Wollaston, 1865
Philorhizus (Philorhizus) vectensis Rye, 1873
Philorhizus (Philorhizus) vieirai Mateu, 1957
Philorhizus (Philorhizus) wollastoni Fauvel, 1905
Philorhizus wollastoni nitidus Mateu, 1957
Philorhizus wollastoni wollastoni Fauvel, 1905
Platyderus (Platyderus) barrosi Jeanne, 1996
Platyderus (Platyderus) beseanus Jeanne, 1970
Platyderus (Platyderus) coiffaiti Jeanne, 1996
Platyderus (Platyderus) gallaecus Jeanne, 1970
Platyderus (Platyderus) gregarius Reiche, 1862
Platyderus (Platyderus) lusitanicus Dejean, 1828
Platyderus lusitanicus herminius Jeanne, 1970
Platyderus lusitanicus lusitanicus Dejean, 1828
Platyderus (Platyderus) portalegrae Vuillefroy, 1868
Platyderus (Platyderus) preciosae Campos & Novoa, 2005
Platyderus (Platyderus) saezi Vuillefroy, 1868
Platytarus bufo Fabricius, 1801
Platytarus faminii Dejean, 1826
Platytarus faminii faminii Dejean, 1826
Poecilus (Ancholeus) crenatus (Dejean, 1828)⇒ Poecilus gisellae gisellae Csiki, 1930
Poecilus (Ancholeus) gisellae Csiki, 1930
Poecilus gisellae gisellae Csiki, 1930
Poecilus (Ancholeus) nitidus Dejean, 1828
Poecilus (Ancholeus) wollastoni Wollaston, 1854
Poecilus (Carenostylus) purpurascens Dejean, 1828
Poecilus purpurascens purpurascens Dejean, 1828
Poecilus (Macropoecilus) kugelanni Panzer, 1797Kugelann's Green Clock
Poecilus (Macropoecilus) striatopunctatus Duftschmid, 1812
Poecilus (Macropoecilus) subcoeruleus Quensel, 1806
Poecilus crenulatus crenulatus Dejean, 1828
Poecilus (Poecilus) prasinotinctus Csiki, 1930
Poecilus (Poecilus) quadricollis Dejean, 1828
Polistichus connexus Geoffroy, 1785
Pseudanchomenus aptinoides Tarnier, 1860Ground-beetle
Pseudomasoreus canigoulensis Fairmaire & ..., 1854
Pseudoophonus (Platus) calceatus Duftschmid, 1812
Pseudoophonus (Pseudoophonus) griseus Panzer, 1797
Pseudoophonus (Pseudoophonus) rufipes De Geer, 1774
Pterostichus (Argus) strenuus Panzer, 1797
Pterostichus (Argutor) vernalis Panzer, 1796Spring Blackclock
Pterostichus (Corax) ghilianii Putzeys, 1846
Pterostichus pumilio pumilio Dejean, 1828
Pterostichus (Haptoderus) pumilio Dejean, 1828
Pterostichus (Lagarus) vernalis Panzer, 1796
Pterostichus (Lamenius) caudicalis Say, 1823
Pterostichus anthracinus anthracinus Illiger, 1798
Pterostichus aterrimus aterrimus Herbst, 1784
Pterostichus (Melanius) nigrita (Paykull, 1790)⇒ Pterostichus anthracinus anthracinus Illiger, 1798
Pterostichus (Neohaptoderus) stictopleurus Fairmaire, 1888
Pterostichus (Oreophilus) paulini Vuillefroy, 1868
Pterostichus paulini paulini Vuillefroy, 1868
Pterostichus paulini vanvolxemi Putzeys, 1874
Pterostichus (Pseudomaseus) anthracinus Illiger, 1798
Pterostichus (Pseudomaseus) nigrita Paykull, 1790
Pterostichus nigrita nigrita
Pterostichus (Rhagadus) thorectes Bates, 1873
Pterostichus (Steropus) ebenus Quensel, 1806
Pterostichus (Steropus) ghilianii Putzeys, 1846
Pterostichus crenatus (Duftschmid, 1812)⇒ Pterostichus vernalis Panzer, 1796
Pterostichus paulinoi Vuillefroy, 1868
Pterostichus paulinoi paulinoi Vuillefroy, 1868
Pterostichus paulinoi vanvolxemi Putzeys, 1874
Scybalicus oblongiusculus Dejean, 1829
Singilis (Singilis) bicolor (Rambur, 1837)⇒ Singilis taschkensis Jedlicka, 1961
Singilis taschkensis Jedlicka, 1961
Siopelus (Neosiopelus) pediobius Alluaud, 1926
Sphodrus leucophthalmus Linnaeus, 1758
Stenolophus (Agonoderus) comma Fabricius, 1775
Stenolophus (Egadroma) marginatus Dejean, 1829
Stenolophus (Stenolophus) abdominalis Géné, 1836
Stenolophus abdominalis abdominalis Gené, 1836
Stenolophus (Stenolophus) mixtus Herbst, 1784
Stenolophus (Stenolophus) paulinoi Heyden, 1891
Stenolophus (Stenolophus) proximus Dejean, 1829
Stenolophus (Stenolophus) skrimshiranus Stephens, 1828
Stenolophus (Stenolophus) teutonus Schrank, 1781
Stomis (Stomis) pumicatus Panzer, 1796
Stomis pumicatus pumicatus Panzer, 1796
Styracoderus azarai Perez Arcas, 1865
Syntomus foveatus Geoffroy, 1785
Syntomus foveolatus Dejean, 1831
Syntomus fuscomaculatus Motschulsky, 1844
Syntomus lundbladi Jeannel, 1938
Syntomus nitidulus Piochard de ..., 1868
Syntomus obscuroguttatus Duftschmid, 1812
Syntomus paludicola Gistel, 1857
Syntomus truncatellus Linnaeus, 1761
Synuchus (Synuchus) vivalis Illiger, 1798
Synuchus vivalis vivalis (Illiger, 1798)⇒ Synuchus vivalis Illiger, 1798
Trechicus nigriceps (Dejean, 1831)⇒ Perigona nigriceps Dejean, 1831
Trichotichnus (Trichotichnus) laevicollis Duftschmid, 1812
Trymosternus onychinus Dejean, 1825
Tschitscherinellus cordatus Dejean, 1825
Tschitscherinellus cordatus cordatus Dejean, 1825
Typsiharpalus (Typsiderus) bonvouloiri Vuillefroy, 1866
Zabrus (Epomidozabrus) flavangulus Chevrolat, 1840
Zabrus (Epomidozabrus) humeralis Uhagon, 1904
Zabrus (Euryzabrus) pinguis Dejean, 1831
Zabrus (Iberozabrus) estrellanus Heyden, 1880
Zabrus (Iberozabrus) gravis Dejean, 1828
Zabrus (Iberozabrus) silphoides (Dejean, 1828)⇒ Zabrus flavangulus Chevrolat, 1840
Zabrus silphoides silphoides Dejean, 1828
Zabrus (Zabrus) ignavus Csiki, 1907
Zabrus ignavus ignavus (Csiki, 1907)⇒ Zabrus ignavus Csiki, 1907
Zabrus (Zabrus) tenebrioides Goeze, 1777Corn ground beetle
Zargus desertae Wollaston, 1854
Zargus monizii Wollaston, 1860
Zargus pellucidus Wollaston, 1854
Zargus schaumii Wollaston, 1854
Zuphioides longicolle LeConte, 1879
Zuphium (Zuphium) numidicum Lucas, 1846
Zuphium (Zuphium) olens P.Rossi, 1790

Subnodes of Harpalinae with occurrences in Portugal
SupertribeAnthiitae
Bonelli, 1813
1
1
Genus
Species

Taxom

SupertribeChlaeniitae
Bonelli, 1810
11
25
Genera
Species

Taxom

SupertribeDryptitae
Bonelli, 1810
5
9
Genera
Species

Taxom

SupertribeHarpalitae
Erwin, 1985
31
141
Genera
Species

Taxom

SupertribeLebiitae
Bonelli, 1810
24
79
Genera
Species

Taxom

SupertribePlatynitae
Bonelli, 1810
11
71
Genera
Species

Taxom

SupertribePterostichitae
Bonelli, 1810
11
100
Genera
Species

Taxom



Links, sources and further information
Harpalinae at artfakta.se
Harpalinae at dyntaxa.se
Harpalinae at itis.gov