AntsAntsArachnidsArachnids
BeesBeesBeetlesBeetles
BookliceBookliceBugsBugs
BumblebeesBumblebeesButterfliesButterflies
CicadasCicadasCrane fliesCrane flies
DragonfliesDragonfliesEarwigsEarwigs
FliesFliesIsopodsIsopods
LocustsLocustsMantisesMantises
MayfliesMayfliesMosquitoesMosquitoes
MyriapodsMyriapodsNet-wingedNet-winged
Plant-parasiticPlant-parasiticScorpion fliesScorpion flies
TermitesTermitesWaspsWasps
INSECTOIDGERENGDARK LIGHT
Home Arthropoda Imprint Contact
TAXFAMGENSPESUB
⌕
Checklist for Subfamily Harpalinae Bonelli, 1810
and region Croatia (Yugoslavia)


Class
 		Insecta
Linnaeus, 1758
Insects, True insects		409
3849
10823		Families
Genera
Species		 
Taxom
 

Subclass
 		Pterygota
Lang, 1888
Winged insects		408
3848
10822		Families
Genera
Species		 
Taxom
 

Infraclass
 		Neoptera
van der Wulp, 1890
Modern Wing-folding insects		395
3811
10763		Families
Genera
Species		 
Taxom
 

Superorder
 		Holometabola
Heider, 1889
Holometabolous Insects		262
3111
9355		Families
Genera
Species		 
Taxom
 

Order
 		Coleoptera
Linnaeus, 1758
Beetles, Coleopterans		79
1143
3526		Families
Genera
Species		 
Taxom
 

Suborder
 		Adephaga
Schellenberg, 1806
Adephagans, Ground Beetles and Water Beetles		5
150
826		Families
Genera
Species		 
Taxom
 

Superfamily
 		Caraboidea
Latreille, 1802
1
132
788		Family
Genera
Species		 
Taxom
 
FamilyCarabidae
Latreille, 1802
Carabid beetles, Carabids, Ground Beetles, Tiger Beetles		132
788		Genera
Species		 
Taxom
 
SubfamilyHarpalinae
Bonelli, 1810
73
441		Genera
Species		 
Taxom
 


Genera for Croatia
73 Genera with 441 Species
#ABCDGHLMOPSTZ
Abax Bonelli, 181211SpeciesTaxom
Acinopus Dejean, 18295SpeciesTaxom
Acupalpus Latreille, 188414SpeciesTaxom
Agonum Bonelli, 181231SpeciesTaxom
Amara Bonelli, 1831Seed-Eating Ground Beetles59SpeciesTaxom
Amblystomus Erichson, 18836SpeciesTaxom
Anchomenus Bonelli, 18473SpeciesTaxom
Anisodactylus Dejean, 19338SpeciesTaxom
Anomotarus Chaudoir, 18601SpeciesTaxom
Anthracus Motschulsky, 18624SpeciesTaxom
Apristus Chaudoir, 18461SpeciesTaxom
Badister Clairville, 179610SpeciesTaxom
Bradycellus Erichson, 18128SpeciesTaxom
Calathus Bonelli, 180220SpeciesTaxom
Callistus Bonelli, 17752SpeciesTaxom
Calodromius Reitter, 17981SpeciesTaxom
Carterus Dana, 17921SpeciesTaxom
Chlaenius Bonelli, 1820Vivid Metallic Ground Beetle19SpeciesTaxom
Cryptophonus Brandmayr & ..., 18293SpeciesTaxom
Cymindis Latreille, 18207SpeciesTaxom
Daptus Fischer von ..., 18231SpeciesTaxom
Demetrias Bonelli, 18193SpeciesTaxom
Diachromus Erichson, 17581SpeciesTaxom
Dicheirotrichus Jacquelin du..., 18292SpeciesTaxom
Dinodes Bonet, 18201SpeciesTaxom
Ditomus Dana, 17923SpeciesTaxom
Dixus Billberg, 17901SpeciesTaxom
Dolichus Bonelli, 17831SpeciesTaxom
Dromius Bonelli, 17584SpeciesTaxom
Drypta Latreille, 17902SpeciesTaxom
Graniger Motschulsky, 18211SpeciesTaxom
Gynandromorphus Dejean, 18062SpeciesTaxom
Harpalus Latreille, 184767SpeciesTaxom
Henrotiochoromus Busulini, 19141SpeciesTaxom
Hololeius LaFerté-Séne..., 18561SpeciesTaxom
Laemostenus Bonelli, 178426SpeciesTaxom
Lebia Latreille, 1966colorful foliage ground beetle13SpeciesTaxom
Licinus Latreille, 18036SpeciesTaxom
Limodromus Motschulsky, 18352SpeciesTaxom
Lionychus Wissmann, 18121SpeciesTaxom
Microlestes Schmidt-Goebel, 17776SpeciesTaxom
Molops Bonelli, 180133SpeciesTaxom
Odacantha Paykull, 17661SpeciesTaxom
Olisthopus Dejean, 18125SpeciesTaxom
Omphreus Dejean, 19305SpeciesTaxom
Oodes Bonelli, 17923SpeciesTaxom
Ophonus Dejean, 182822SpeciesTaxom
Oxypselaphus Chaudoir, 17841SpeciesTaxom
Panagaeus Latreille, 17581SpeciesTaxom
Paradromius Fowler, 17952SpeciesTaxom
Paranchus Lindroth, 17961SpeciesTaxom
Parazuphium Jeannel, 18335SpeciesTaxom
Parophonus Ganglbauer, 18296SpeciesTaxom
Pedius Motschulsky, 18241SpeciesTaxom
Perigona Laporte, 18311SpeciesTaxom
Philorhizus Hope, 17906SpeciesTaxom
Platyderus Stephens, 18125SpeciesTaxom
Platynus Bonelli, 18017SpeciesTaxom
Poecilus Bonelli, 178316SpeciesTaxom
Polistichus Bonelli, 17902SpeciesTaxom
Pseudoophonus Motschulsky, 17743SpeciesTaxom
Pterostichus Bonelli, 1812Woodland Ground Beetle60SpeciesTaxom
Scybalicus Schaum, 18291SpeciesTaxom
Speluncarius Reitter, 20162SpeciesTaxom
Sphodrus Clairville, 17581SpeciesTaxom
Stenolophus Dejean, 1781Seedcorn Beetle10SpeciesTaxom
Stomis Clairville, 18122SpeciesTaxom
Syntomus Hope, 17616SpeciesTaxom
Synuchus Gyllenhal, 17982SpeciesTaxom
Trechicus LeConte, 18311SpeciesTaxom
Trichotichnus A. Morawitz, 18121SpeciesTaxom
Zabrus Clairville, 17773SpeciesTaxom
Zuphium Latreille, 17901SpeciesTaxom


Species list for Croatia
441 Species and 146 Subspecies
Abax (Abacopercus) carinatus Duftschmid, 1812
Abax carinatus carinatus Duftschmid, 1812
Abax (Abax) ovalis Duftschmid, 1812
Abax (Abax) parallelepipedus (Piller & Mit..., 1783)⇒ Abax pilleri Csiki, 1916
Abax parallelepipedus parallelepipedus Piller & Mit..., 1783
Abax parallelepipedus subpunctatus Dejean, 1828
Abax (Abax) parallelus Duftschmid, 1812
Abax parallelus parallelus Duftschmid, 1812
Abax (Abax) pilleri Csiki, 1916
Abax (Pterostichoabax) beckenhaupti (Duftschmid, 1812)⇒ Abax beckenhauptii Duftschmid, 1812
Abax (Pterostichoabax) beckenhauptii Duftschmid, 1812
Acinopus (Acinopus) laevigatus Ménétriés, 1832
Acinopus laevigatus laevigatus Ménétriés..., 1832
Acinopus (Acinopus) picipes Olivier, 1795
Acinopus (Oedematicus) megacephalus (P. Rossi, 1794)⇒ Acinopus picipes Olivier, 1795
Acinopus (Osimus) ammophilus Dejean, 1829
Acupalpus (Acupalpus) brunnipes Sturm, 1825
Acupalpus (Acupalpus) dubius Schilsky, 1888
Acupalpus (Acupalpus) elegans Dejean, 1829
Acupalpus (Acupalpus) exiguus Dejean, 1829
Acupalpus (Acupalpus) flavicollis Sturm, 1825
Acupalpus (Acupalpus) luteatus (Duftschmid, 1812)⇒ Acupalpus dubius Schilsky, 1888
Acupalpus (Acupalpus) maculatus Schaum, 1860
Acupalpus (Acupalpus) meridianus Linnaeus, 1761Midday harp ground beetle
Acupalpus (Acupalpus) parvulus Sturm, 1825
Acupalpus (Acupalpus) suturalis (Dejean, 1829)⇒ Acupalpus meridianus Linnaeus, 1761
Acupalpus suturalis suturalis Dejean, 1829
Acupalpus (Ancylostria) interstitialis Reitter, 1884
Acupalpus (Tachistodes) notatus Mulsant & Rey, 1861
Acupalpus (Tachistodes) paludicola Reitter, 1884
Agonum (Agonum) marginatum Linnaeus, 1758
Agonum (Agonum) monachum Duftschmid, 1812
Agonum monachum monachum Duftschmid, 1812
Agonum (Agonum) muelleri Herbst, 1784Müller'S Harp Ground Beetle
Agonum muelleri muelleri Herbst, 1784
Agonum (Agonum) nigrum (Dejean, 1828)⇒ Agonum fuliginosum Panzer, 1809
Agonum (Agonum) sordidum Dejean, 1828
Agonum sordidum sordidum
Agonum (Europhilus) antennarium Duftschmid, 1812
Agonum (Europhilus) fuliginosum Panzer, 1809
Agonum gracile gracile Sturm, 1824
Agonum (Europhilus) micans Nicolai, 1822
Agonum (Europhilus) scitulum Dejean, 1828
Agonum (Europhilus) thoreyi Dejean, 1828
Agonum (Melanagonum) gisellae Csiki, 1931
Agonum (Olisares) afrum (Duftschmid, 1812)⇒ 
Agonum (Olisares) angustatum Dejean, 1828
Agonum (Olisares) dolens C.R. Sahlberg, 1827
Agonum (Olisares) duftschmidi J. Schmidt, 1994
Agonum (Olisares) emarginatum (Gyllenhal, 1827)⇒ Agonum thoreyi Dejean, 1828
Agonum (Olisares) hypocrita Apfelbeck, 1904
Agonum (Olisares) longicorne Chaudoir, 1846
Agonum (Olisares) lugens Duftschmid, 1812
Agonum (Olisares) permoestum Puel, 1938
Agonum (Olisares) sexpunctatum (Linnaeus, 1758)⇒ Agonum muelleri Herbst, 1784
Agonum (Olisares) versutum Sturm, 1824
Agonum (Olisares) viduum Panzer, 1796
Agonum (Olisares) viridicupreum Goeze, 1777
Agonum viridicupreum viridicupreum Goeze, 1777
Agonum (Punctagonum) sexpunctatum Linnaeus, 1758
Agonum (Stictanchus) gracilipes Duftschmid, 1812
Amara (Amara) aenea De Geer, 1774Common sun beetle
Amara (Amara) anthobia A. Villa & G..., 1833Italy harp ground beetle
Amara (Amara) communis Panzer, 1797Common harp ground beetle
Amara (Amara) convexior Stephens, 1828
Amara (Amara) curta Dejean, 1828
Amara (Amara) eurynota Panzer, 1796
Amara (Amara) familiaris Duftschmid, 1812Bareyed goby
Amara (Amara) littorea C.C. Thomson, 1857
Amara (Amara) lucida Duftschmid, 1812
Amara (Amara) lunicollis Schiodte, 1837Golden-banded goby
Amara (Amara) montivaga Sturm, 1825
Amara (Amara) nitida (Sturm, 1825)⇒ Amara sinuosa Casey, 1918
Amara (Amara) ovata (Fabricius, 1792)⇒ Amara tricuspidata Dejean, 1831
Amara (Amara) proxima Putzeys, 1866
Amara (Amara) pulpani (Kult, 1949)⇒ Amara communis Panzer, 1797
Amara (Amara) saphyrea Dejean, 1828
Amara (Amara) similata Gyllenhal, 1810
Amara (Amara) tibialis (Paykull, 1798)⇒ Amara tumida tumida Morawitz, 1862
Amara (Amarocelia) erratica Duftschmid, 1812
Amara (Bradytus) apricaria Paykull, 1790Exposed sun beetle
Amara (Bradytus) consularis Duftschmid, 1812
Amara (Bradytus) crenata Dejean, 1828
Amara (Bradytus) fulva (O.F. Muller, 1776)⇒ Amara apricaria Paykull, 1790
Amara (Celia) arenaria Putzeys, 1865
Amara (Celia) bifrons Gyllenhal, 1810Two-faced harp ground beetle
Amara (Celia) cursitans Zimmermann, 1832
Amara (Celia) dalmatina Dejean, 1828
Amara (Celia) erratica Duftschmid, 1812Bareyed goby
Amara (Celia) montana (Dejean, 1828)⇒ Amara spectabilis Schaum, 1858
Amara municipalis municipalis Duftschmid, 1812
Amara (Celia) praetermissa C.R. Sahlberg, 1827
Amara quenseli silvicola Zimm., 1832
Amara (Celia) sabulosa Audinet-Serv..., 1821
Amara (Celia) sinuosa Casey, 1918
Amara (Curtonotus) aulica Panzer, 1796Princely harp ground beetle
Amara (Curtonotus) convexiuscula Marsham, 1802
Amara (Curtonotus) gebleri Dejean, 1831
Amara (Curtonotus) pennsylvanica Hayward, 1908
Amara tumida tumida Morawitz, 1862
Amara (Leirides) spectabilis Schaum, 1858
Amara (Paracelia) dalmatina Dejean, 1828
Amara (Parapercosia) taurica Motschulsky, 1844
Amara (Percosia) equestris Duftschmid, 1812
Amara equestris equestris Duftschmid, 1812
Amara (Xenocelia) bischoffi Jedlička, 1946
Amara (Xenocelia) cursitans C. Zimmermann, 1832
Amara (Xenocelia) fusca (Dejean, 1828)⇒ Amara quenseli silvicola Zimm., 1832
Amara (Xenocelia) ingenua Duftschmid, 1812
Amara (Xenocelia) municipalis Duftschmid, 1812
Amara (Zezea) chaudoiri Schaum, 1858
Amara chaudoiri incognita Fassati, 1946
Amara (Zezea) concinna C. Zimmermann, 1832
Amara (Zezea) fulvipes (Audinet-Serv..., 1821)⇒ Amara pennsylvanica Hayward, 1908
Amara (Zezea) kulti Fassati, 1947
Amara (Zezea) plebeja (Gyllenhal, 1810)⇒ Amara equestris equestris Duftschmid, 1812
Amara (Zezea) reflexicollis Motschulsky, 1844
Amara reflexicollis reflexicollis Motschulsky, 1845
Amara (Zezea) strandi Lutshnik, 1933
Amara (Zezea) tricuspidata Dejean, 1831
Amblystomus levantinus Reitter, 1883
Amblystomus levantinus levantinus Reitter, 1883
Amblystomus metallescens Dejean, 1829
Amblystomus niger (Heer, 1841)⇒ Amblystomus nigrinus Csiki, 1931
Amblystomus nigrinus Csiki, 1931
Amblystomus rectangulus Reitter, 1883
Anchomenus (Anchomenus) dorsalis Pontoppidan, 1763
Anchomenus dorsalis dorsalis
Anchomenus orphanus ♰ O. Heer, 1847
Anisodactylus (Anisodactylus) binotatus Fabricius, 1787Two-marked harp ground beetle
Anisodactylus (Anisodactylus) nemorivagus Duftschmid, 1812Heath Short-spur
Anisodactylus (Hexatrichus) poeciloides Stephens, 1828Saltmarsh Short-spur
Anisodactylus poeciloides poeciloides Stephens, 1828
Anisodactylus (Pseudanisodactylus) signatus Panzer, 1796
Anisodactylus (Pseudodichirus) intermedius Dejean, 1829
Anisodactylus (Spongopus) pueli Schauberger, 1933
Anisodactylus pueli pueli Schauberger, 1933
Anomotarus geophilus geophilus Montrouzier, 1860
Anthracus consputus Duftschmid, 1812
Anthracus longicornis Schaum, 1857
Anthracus quarnerensis Reitter, 1884
Anthracus transversalis Schaum, 1862
Apristus subaeneus Chaudoir, 1846
Badister (Badister) bullatus Schrank, 1798
Badister (Badister) lacertosus Sturm, 1815
Badister (Badister) meridionalis Puel, 1925
Badister (Badister) unipustulatus Bonelli, 1813
Badister (Bandia) dilatatus Chaudoir, 1837
Badister (Bandia) peltatus Panzer, 1796
Badister peltatus peltatus Panzer, 1796
Badister (Baudia) collaris (Motschulsky, 1844)⇒ Badister peltatus peltatus Panzer, 1796
Badister (Baudia) dilatatus Chaudoir, 1837
Badister (Baudia) peltatus Panzer, 1796
Bradycellus (Stenocellus) nebulosus LeConte, 1853
Bradycellus (Triliarthrus) caucasicus Chaudoir, 1846
Bradycellus (Triliarthrus) csikii Laczó, 1912
Bradycellus (Triliarthrus) distinctus Dejean, 1829
Bradycellus (Triliarthrus) ganglbaueri Apfelbeck, 1904
Bradycellus (Triliarthrus) harpalinus Audinet-Serv..., 1821European harp ground beetle
Bradycellus (Triliarthrus) ruficollis Stephens, 1828
Bradycellus (Triliarthrus) verbasci Duftschmid, 1812
Calathus (Bedelinus) circumseptus Germar, 1824
Calathus fuscipes fuscipes Goeze, 1777
Calathus fuscipes graecus (Dejean, 1831)⇒ Calathus fuscipes punctipennis Germar, 1824
Calathus fuscipes punctipennis Germar, 1824
Calathus (Calathus) glabricollis Dejean, 1828
Calathus (Neocalathus) ambiguus Paykull, 1790
Calathus ambiguus ambiguus Paykull, 1790
Calathus (Neocalathus) erratus C.R.Sahlberg, 1827
Calathus erratus erratus C.R. Sahlberg, 1827
Calathus (Neocalathus) melanocephalus Linnaeus, 1758
Calathus melanocephalus antoinei Puel, 1939
Calathus melanocephalus furvus Tschitschérine, 1895
Calathus melanocephalus melanocephalus Linné, 1758
Calathus melanocephalus paphlagonicus Maran, 1935
Calathus melanocephalus purkynei Maran, 1935
Calathus (Neocalathus) metallicus Dejean, 1828
Calathus metallicus metallicus Dejean, 1828
Calathus (Neocalathus) micropterus Dufischmid, 1812
Calathus (Neocalathus) mollis Marsham, 1802
Calathus mollis mollis Marsham, 1802
Callistus lunatus Fabricius, 1775
Callistus lunatus lunatus Fabricius, 1775
Calodromius spilotus Illiger, 1798
Carterus (Pristocarterus) dama P.Rossi, 1792
Chlaenius (Chlaeniellus) flavipes Ménétriés, 1832
Chlaenius (Chlaeniellus) nigricornis Fabricius, 1787
Chlaenius (Chlaeniellus) nitidulus (Schrank, 1781)⇒ Chlaenius nigricornis Fabricius, 1787
Chlaenius (Chlaeniellus) olivieri Crotch, 1871
Chlaenius (Chlaeniellus) terminatus Dejean, 1826
Chlaenius (Chlaeniellus) tristis (Schaller, 1783)⇒ Chlaenius moestus Csiki, 1931
Chlaenius tristis tristis Schaller, 1783
Chlaenius (Chlaeniellus) vestitus Paykull, 1790
Chlaenius (Chlaenites) spoliatus P. Rossi, 1792
Chlaenius spoliatus spoliatus P. Rossi, 1792
Chlaenius (Chlaenius) festivus Panzer, 1796
Chlaenius festivus festivus Panzer, 1796
Chlaenius festivus velutinus Duftschmid, 1812
Chlaenius (Chlaenius) velutinus Duftschmid, 1812
Chlaenius (Epomis) circumscriptus Duftschmid, 1812
Chlaenius (Homalolachnus) moestus Csiki, 1931
Chlaenius (Trichochlaenius) aeneocephalus Dejean, 1826
Chlaenius aeneocephalus aeneocephalus Dejean, 1826
Chlaenius decipiens L.Dufour, 1820
Cryptophonus litigiosus Dejean, 1829
Cryptophonus melancholicus Dejean, 1829
Cryptophonus tenebrosus Dejean, 1829
Cymindis (Cymindis) angularis Gyllenhal, 1810
Cymindis (Cymindis) axillaris (Fabricius, 1794)⇒ Cymindis scapularis Schaum, 1857
Cymindis (Cymindis) scapularis Schaum, 1857
Cymindis (Tarsostinus) macularis Fischer von ..., 1824
Cymindis (Tarulus) coadunata Dejean, 1825
Cymindis coadunata coadunata Dejean, 1825
Cymindis (Tarulus) lineola L. Dufour, 1820
Daptus vittatus Fischer von ..., 1823
Demetrias (Aetophorus) imperialis Germar, 1824
Demetrias (Demetrias) atricapillus Linnaeus, 1758
Demetrias (Demetrias) monostigma Samouelle, 1819
Diachromus germanus Linnaeus, 1758
Dicheirotrichus (Trichocellus) lacustris L.Redtenbacher, 1858
Dicheirotrichus (Trichocellus) obsoletus Dejean, 1829
Dinodes (Dinodes) decipiens (L. Dufour, 1820)⇒ 
Ditomus calydonius P. Rossi, 1790
Ditomus calydonius calydonius P. Rossi, 1790
Ditomus tricuspidatus Fabricius, 1792
Dixus clypeatus P. Rossi, 1790
Dolichus halensis Schaller, 1783
Dromius (Dromius) agilis Fabricius, 1787
Dromius (Dromius) fenestratus Fabricius, 1794
Dromius (Dromius) meridionalis Dejean, 1825
Dromius (Dromius) quadrimaculatus Linnaeus, 1758
Drypta (Deserida) distincta P.Rossi, 1792
Drypta (Drypta) dentata P.Rossi, 1790
Graniger cordicollis Audinet-Serv..., 1821
Gynandromorphus etruscus Quensel, 1806
Gynandromorphus etruscus etruscus Quensel, 1806
Harpalus (Acardystus) atratus Latreille, 1804
Harpalus (Acardystus) attenuatus Stephens, 1828
Harpalus (Acardystus) flavescens Piller &Mitt..., 1783
Harpalus (Acardystus) pygmaeus Dejean, 1829
Harpalus (Actephilus) albanicus Reitter, 1900
Harpalus (Amblystus) latus Linnaeus, 1758
Harpalus (Amblystus) luteicornis Dufischmid, 1812
Harpalus (Amblystus) marginellus Gyllenhal, 1827
Harpalus (Amblystus) rubripes Dufischmid, 1812Bareyed goby
Harpalus (Amblystus) solitaris Dejean, 1829Checked goby
Harpalus (Artabas) punctatostriatus (Dejean, 1829)⇒ Harpalus rubefactus rubefactus Bates, 1873
Harpalus (Artabas) rumelicus Apfelbeck, 1904
Harpalus (Cryptophonus) litigiosus (Dejean, 1829)⇒ Cryptophonus litigiosus Dejean, 1829
Harpalus litigiosus litigiosus Dejean, 1829
Harpalus (Cryptophonus) melancholicus (Dejean, 1829)⇒ Cryptophonus melancholicus Dejean, 1829
Harpalus melancholicus melancholicus
Harpalus melancholicus reicheianus Kataev, 2012
Harpalus (Cryptophonus) tenebrosus (Dejean, 1829)⇒ 
Harpalus tenebrosus tenebrosus
Harpalus (Euharpalops) fraternus LeConte, 1852
Harpalus (Harpalobius) autumnalis Duftschmid, 1812
Harpalus (Harpalobius) hirtipes Panzer, 1796
Harpalus (Harpaloderus) honestus Dufischmid, 1812St. Bees Seed-eater
Harpalus (Harpaloderus) neglectus Serville, 1821
Harpalus neglectus neglectus Audinet-Serv..., 1821
Harpalus (Harpaloderus) rufipalpis Sturm, 1818
Harpalus rufipalpis rufipalpis Sturm, 1818
Harpalus (Harpaloderus) sulphuripes Germar, 1824
Harpalus sulphuripes sulphuripes Germar, 1824
Harpalus (Harpalus) affinis (Schrank, 1781)⇒ Harpalus serripes ernsti Kataev, 1995
Harpalus (Harpalus) brevicornis Germar, 1824
Harpalus (Harpalus) caspius Steven, 1806
Harpalus (Harpalus) oblitus (Dejean, 1829)⇒ Harpalus fraternus LeConte, 1852
Harpalus oblitus oblitus Dejean, 1829
Harpalus (Harpalus) saxicola Dejean, 1829
Harpalus (Harpalus) subcylindricus Dejean, 1829
Harpalus (Harpalus) xanthopus Gemminger & ..., 1868
Harpalus xanthopus winkleri Schauberger, 1923
Harpalus (Lasioharpalus) cupreus Dejean, 1829
Harpalus cupreus fastuosus Faldermann, 1835
Harpalus (Lasioharpalus) dimidiatus P. Rossi, 1790
Harpalus (Lasioharpalus) distinguendus Duftschmid, 1812
Harpalus distinguendus distinguendus Duftschmid, 1812
Harpalus (Lasioharpalus) smaragdinus Duftschmid, 1812
Harpalus rubefactus rubefactus Bates, 1873
Harpalus (Opadius) laevipes Zetterstedt, 1828
Harpalus (Pheuginus) anxius Duftschmid, 1812
Harpalus (Pheuginus) flavicornis Dejean, 1829
Harpalus flavicornis flavicornis Dejean, 1829
Harpalus (Pheuginus) modestus Dejean, 1829
Harpalus (Pheuginus) politus Dejean, 1829
Harpalus politus politus Dejean, 1829
Harpalus (Pheuginus) serripes Quensel, 1806
Harpalus serripes ernsti Kataev, 1995
Harpalus serripes serripes Quensel, 1806
Harpalus (Pheuginus) taciturnus Dejean, 1829
Harpalus (Pheuginus) tardus Panzer, 1796
Harpalus (Plectralidus) euchlorus Ménétriés, 1839
Harpalus (Plectralidus) karamani Apfelbeck, 1902
Harpalus karamani hoffmanni Schauberger, 1928
Harpalus karamani karamani Apfelbeck, 1902
Harpalus (Pseudoophonus) calceatus (Duftschmid, 1812)⇒ Pseudoophonus calceatus Duftschmid, 1812
Harpalus (Pseudoophonus) griseus (Panzer, 1796)⇒ Pseudoophonus griseus Panzer, 1797
Harpalus (Pseudoophonus) rufipes (DeGeer, 1774)⇒ 
Harpalus (Semiophonus) signaticornis Duftschmid, 1812
Harpalus reitteri Wrase & Kataev, 2011
Harpalus tabidus ♰ O. Heer, 1847
Henrotiochoromus winkleri Breit, 1914
Hololeius ceylanicus Nietner, 1856
Laemostenus caussolensis caussolensis Ochs, 1937
Laemostenus (Antisphodrus) cavicola Schaum, 1858
Laemostenus cavicola aeacus L. Miller, 1861
Laemostenus cavicola cavicola Schaum, 1858
Laemostenus cavicola erberii Schaufuss, 1863
Laemostenus cavicola modestus Schaufuss, 1862
Laemostenus cavicola muelleri Ganglbauer, 1903
Laemostenus cavicola redtenbacheri
Laemostenus cavicola redtenbacherii L.Schaufuss, 1865
Laemostenus cavicola romualdi G.Müller, 1905
Laemostenus cavicola sinjensis G.Müller, 1906
Laemostenus (Antisphodrus) elongatus (Dejean, 1828)⇒ Laemostenus caussolensis caussolensis Ochs, 1937
Laemostenus elongatus elongatus Dejean, 1828
Laemostenus elongatus pentheri G.Müller, 1924
Laemostenus (Antisphodrus) puchneri Lohaj & Mlejnek, 2007
Laemostenus (Antisphodrus) schreibersii Kuster, 1846
Laemostenus (Cryptotrichus) janthinus Duftschmid, 1812
Laemostenus janthinus janthinus Duftschmid, 1812
Laemostenus (Laemostenus) complanatus Dejean, 1828Bynoe's goby
Laemostenus (Laemostenus) dalmatinus Dejean, 1828
Laemostenus (Laemostenus) venustus Dejean, 1828
Laemostenus (Pristonychus) algerinus Gory, 1833
Laemostenus algerinus algerinus (Gory, 1833)⇒ Laemostenus algerinus Gory, 1833
Laemostenus (Pristonychus) terricola Herbst, 1784Bandy-legged burrow-dweller beetle
Laemostenus terricola punctatus Dejean, 1828
Laemostenus terricola terricola Herbst, 1784
Lebia (Lamprias) cyanocephala Linnaeus, 1758Blue plunderer
Lebia cyanocephala cyanocephala Linnaeus, 1758
Lebia (Lamprias) fulvicollis (Fabricius, 1792)⇒ Lebia pubipennis Dufour, 1820
Lebia fulvicollis fulvicollis Fabricius, 1792
Lebia (Lebia) cruxminor Linnaeus, 1758
Lebia cruxminor cruxminor Linnaeus, 1758
Lebia (Lebia) humeralis Dejean, 1825
Lebia (Lebia) marginata Geoffroy, 1785
Lebia (Lebia) scapularis (Geoffroy, 1785)⇒ Lebia solea Hentz, 1830
Lebia scapularis scapularis Geoffroy, 1785
Lebia (Lebia) trimaculata (Villers, 1789)⇒ 
Lebia (Stephana) lepida Brullé, 1834
Lebia (Stephana) turkestanica Jedlička, 1966
Licinus (Licinus) cassideus Fabricius, 1792
Licinus cassideus cassideus Fabricius, 1792
Licinus (Licinus) depressus (Paykull, 1790)⇒ Licinus cassideus cassideus Fabricius, 1792
Licinus (Licinus) punctatulus Fabricius, 1792
Licinus (Licinus) silphoides (P. Rossi, 1790)⇒ Licinus punctatulus Fabricius, 1792
Licinus (Neorescius) hoffmannseggii Panzer, 1803
Limodromus assimilis (Pontoppidan, 1763)⇒ Platynus assimilis Paykull, 1790
Limodromus krynickii (Sperk, 1835)⇒ Platynus krynickii Sperk, 1835
Lionychus (Paralionychus) quadrillum Duftschmid, 1812
Microlestes abeillei Brisout, 1885
Microlestes abeillei abeillei Brisout de B..., 1885
Microlestes corticalis L. Dufour, 1820
Microlestes maurus Sturm, 1827
Microlestes maurus maurus Sturm, 1827
Microlestes minutulus Goeze, 1777
Molops (Henrotiochromus) winkleri (Breit, 1914)⇒ Molops ovipennis planipennis Apfelbeck, 1902
Molops (Molops) austriacus Ganglbauer, 1889
Molops austriacus austriacus Ganglbauer, 1889
Molops austriacus longulus G.Müller, 1918
Molops (Molops) biokovensis J. Muller, 1916
Molops (Molops) bucephalus Dejean, 1828
Molops (Molops) curtulus Ganglbauer, 1891
Molops curtulus troglavensis G. Müller, 1917
Molops (Molops) dalmatinus Dejean, 1828
Molops (Molops) dinaricus Apfelbeck, 1904
Molops (Molops) elatus Fabricius, 1801
Molops elatus elatus Fabricius, 1801
Molops elatus liburnicus G. Müller, 1908
Molops elatus livnensis G. Müller, 1930
Molops elatus plitvicensis Heyden, 1880
Molops (Molops) longipennis Dejean, 1828
Molops (Molops) obtusangulus Ganglbauer, 1889
Molops obtusangulus novaki G. Müller, 1930
Molops obtusangulus svilajensis G. Müller, 1917
Molops obtusangulus velebiticus G. Müller, 1917
Molops (Molops) ovipennis (Chaudoir, 1847)⇒ Molops austriacus austriacus Ganglbauer, 1889
Molops ovipennis istrianus G. Müller, 1916
Molops ovipennis latiusculus Kraatz, 1879
Molops ovipennis ovipennis Chaudoir, 1847
Molops ovipennis planipennis Apfelbeck, 1902
Molops ovipennis pseudoplanipennis Mlynar, 1977
Molops (Molops) piceus Panzer, 1793
Molops piceus longulus G. Müller, 1918
Molops piceus piceus Panzer, 1793
Molops (Molops) promissus Heyden, 1875
Molops (Molops) simplex Chaudoir, 1868
Molops simplex simplex Chaudoir, 1868
Molops (Molops) striolatus Fabricius, 1801
Odacantha (Odacantha) melanura Linnaeus, 1766
Olisthopus fuscatus Dejean, 1828
Olisthopus glabricollis Germar, 1817
Olisthopus rotundatus Paykull, 1790
Olisthopus rotundatus rotundatus Paykull, 1798
Olisthopus sturmii Duftschmid, 1812
Omphreus (Neomphreus) apfelbecki Reitter, 1893
Omphreus apfelbecki apfelbecki Reitter, 1893
Omphreus apfelbecki biokovensis Csiki, 1908
Omphreus apfelbecki dinaricus Apfelbeck, 1904
Omphreus apfelbecki mosorensis G. Müller, 1930
Oodes gracilis A. Villa & G..., 1833
Oodes helopioides Fabricius, 1792
Oodes helopioides helopioides Fabricius, 1792
Ophonus (Hesperophonus) azureus Fabricius, 1775
Ophonus (Hesperophonus) cribricollis Dejean, 1829
Ophonus (Hesperophonus) rotundatus Dejean, 1829
Ophonus (Hesperophonus) similis Dejean, 1829
Ophonus (Hesperophonus) subquadratus (Dejean, 1829)⇒ 
Ophonus (Incisophonus) incisus Dejean, 1829
Ophonus (Metophonus) brevicollis (Audinet-Serv..., 1821)⇒ Ophonus melletii Heer, 1837
Ophonus (Metophonus) cordatus Duftschmid, 1812
Ophonus (Metophonus) gammeli Schauberger, 1932
Ophonus (Metophonus) laticollis (Mannerheim, 1825)⇒ Ophonus puncticollis Paykull, 1798
Ophonus (Metophonus) melletii Heer, 1837Mellet's Downy-back
Ophonus (Metophonus) parallelus (Dejean, 1829)⇒ Ophonus xaxarsi Schauberger, 1928
Ophonus (Metophonus) puncticeps (Stephens, 1828)⇒ Ophonus puncticollis Paykull, 1798
Ophonus (Metophonus) puncticollis Paykull, 1798
Ophonus (Metophonus) rufibarbis Fabricius, 1792
Ophonus (Metophonus) rupicola Sturm, 1818
Ophonus (Metophonus) schaubergerianus (Puel, 1937)⇒ Ophonus melletii Heer, 1837
Ophonus (Metophonus) xaxarsi Schauberger, 1928
Ophonus (Ophonus) ardosiacus Lutshnik, 1922
Ophonus (Ophonus) diffinis (Dejean, 1829)⇒ Ophonus ardosiacus Lutshnik, 1922
Ophonus (Ophonus) sabulicola Panzer, 1796
Ophonus (Ophonus) stictus Stephens, 1828Oolite Downy-back
Oxypselaphus obscurus Herbst, 1784
Panagaeus (Panagaeus) cruxmajor Linnaeus, 1758Crucifix ground beetle
Paradromius (Manodromius) linearis Olivier, 1795
Paradromius linearis linearis Olivier, 1795
Paranchus albipes Fabricius, 1796White-legged harp ground beetle
Parazuphium (Neozuphium) damascenum Fairmaire, 1897
Parazuphium damascenum damascenum Fairmaire, 1897
Parazuphium (Neozuphium) novaki (J. Muller, 1934)⇒ Parazuphium damascenum damascenum Fairmaire, 1897
Parazuphium (Parazuphium) chevrolatii Laporte, 1833
Parazuphium chevrolatii chevrolatii Laporte de C..., 1833
Parophonus (Ophonomimus) hirsutulus Dejean, 1829
Parophonus (Parophonus) dejeani Csiki, 1932
Parophonus (Parophonus) laeviceps Ménétriés, 1832
Parophonus (Parophonus) maculicornis Duftschmid, 1812
Parophonus (Parophonus) mendax P.Rossi, 1790
Parophonus (Parophonus) planicollis (Dejean, 1829)⇒ Parophonus laeviceps Ménétriés, 1832
Pedius inquinatus Sturm, 1824
Perigona (Trechicus) nigriceps Dejean, 1831Tawny Harp Ground Beetle
Philorhizus (Philorhizus) crucifer Lucas, 1846
Philorhizus crucifer crucifer Lucas, 1846
Philorhizus (Philorhizus) melanocephalus Dejean, 1825Frillfin gobies
Philorhizus (Philorhizus) notatus Stephens, 1827
Philorhizus (Philorhizus) quadrisignatus Dejean, 1825Mab's Lantern
Philorhizus (Philorhizus) sigma P.Rossi, 1790
Platyderus (Platyderus) atticus Apfelbeck, 1904
Platyderus (Platyderus) dalmatinus L. Miller, 1881
Platyderus dalmatinus dalmatinus L. Miller, 1881
Platyderus (Platyderus) rufus Duftschmid, 1812
Platyderus rufus rufus Duftschmid, 1812
Platynus (Batenus) scrobiculatus Fabricius, 1801
Platynus scrobiculatus scrobiculatus Fabricius, 1801
Platynus (Platynus) assimilis Paykull, 1790
Platynus (Platynus) krynickii Sperk, 1835
Platynus (Platynus) longiventris Mannerheim, 1825
Platynus (Trapezodera) scrobiculatus Fabricius, 1801
Platynus scrobiculatus scrobiculatus Fabricius, 1801
Poecilus (Ancholeus) puncticollis (Dejean, 1828)⇒ 
Poecilus (Macropoecilus) koyi Germar, 1824
Poecilus koyi dinaricus Apfelbeck, 1904
Poecilus koyi goricianus Müller G., 1921
Poecilus koyi koyi (Germar, 1824)⇒ Poecilus sericeus desbrochersi Jeannel, 1942
Poecilus koyi lossinianus Fairmaire, 1860
Poecilus koyi mosorensis Jedlicka, 1924
Poecilus lepidus lepidus Leske, 1785
Poecilus (Macropoecilus) sericeus Fischer von ..., 1824
Poecilus (Macropoecilus) striatopunctatus Duftschmid, 1812
Poecilus (Metapedius) striatopunctatus Duftschmid, 1812
Poecilus (Poecilus) cupreus (Linnaeus, 1758)⇒ Poecilus lepidus lepidus Leske, 1785
Poecilus cupreus cupreus Linnaeus, 1758
Poecilus (Poecilus) rebeli Apfelbeck, 1904
Poecilus (Poecilus) versicolor Sturm, 1824
Poecilus (Sogines) punctulatus Schaller, 1783
Polistichus connexus Geoffroy, 1785
Polistichus fasciolatus (P.Rossi, 1790)⇒ Polistichus connexus Geoffroy, 1785
Pseudoophonus (Platus) calceatus Duftschmid, 1812
Pseudoophonus (Pseudoophonus) griseus Panzer, 1797
Pseudoophonus (Pseudoophonus) rufipes De Geer, 1774
Pterostichus (Adelosia) macer Marsham, 1802
Pterostichus macer macer Marsham, 1802
Pterostichus (Argus) strenuus Panzer, 1797
Pterostichus aterrimus aterrimus Herbst, 1784
Pterostichus (Argutor) cursor Dejean, 1828
Pterostichus (Argutor) oblongopunctatus Fabricius, 1787
Pterostichus (Argutor) vernalis Panzer, 1796Spring Blackclock
Pterostichus (Bothriopterus) oblongopunctatus Fabricius, 1787
Pterostichus (Bothriopterus) quadrifoveolatus Letzner, 1852
Pterostichus selmanni selmanni Duftschmid, 1812
Pterostichus (Cheporus) burmeisteri Heer, 1838
Pterostichus burmeisteri burmeisteri Heer, 1838
Pterostichus (Cheporus) transversalis Duftschmid, 1812
Pterostichus (Cophosus) cylindricus Herbst, 1784
Pterostichus (Cryobius) unctulatus Duftschmid, 1812
Pterostichus (Eosteropus) aethiops Panzer, 1796
Pterostichus (Eosteropus) faldermanni Schatzmayr, 1929
Pterostichus (Feronidius) melas Creutzer, 1799
Pterostichus melas depressus Dejean, 1828
Pterostichus melas italicus Dejean, 1828
Pterostichus melas melas Creutzer, 1799
Pterostichus (Haptoderus) unctulatus Duftschmid, 1812
Pterostichus (Lagarus) vernalis Panzer, 1796
Pterostichus (Lamenius) caudicalis Say, 1823
Pterostichus (Leptoferonia) idahoae Csiki, 1930Key goby
Pterostichus (Melanius) anthracinus (Illiger, 1798)⇒ Pterostichus quadrifoveolatus Letzner, 1852
Pterostichus anthracinus anthracinus Illiger, 1798
Pterostichus anthracinus biimpressus Küster, 1853
Pterostichus (Melanius) aterrimus Herbst, 1784
Pterostichus aterrimus aterrimus Herbst, 1784
Pterostichus (Melanius) elongatus (Duftschmid, 1812)⇒ Pterostichus idahoae Csiki, 1930
Pterostichus (Melanius) gracilis Dejean, 1828
Pterostichus gracilis gracilis Dejean, 1828
Pterostichus (Melanius) minor Gyllenhal, 1827
Pterostichus (Melanius) nigrita (Paykull, 1790)⇒ Pterostichus anthracinus anthracinus Illiger, 1798
Pterostichus (Morphnosoma) melanarius Illiger, 1798European Dark Harp Ground Beetle
Pterostichus melanarius melanarius Illiger, 1798
Pterostichus (Omaseus) aterrimus (Herbst, 1784)⇒ Pterostichus transversalis Duftschmid, 1812
Pterostichus (Oreophilus) variolatus Dejean, 1828
Pterostichus variolatus carniolicus Ganglbauer, 1891
Pterostichus (Parahaptoderus) brevis (Duftschmid, 1812)⇒ Pterostichus faldermanni Schatzmayr, 1929
Pterostichus (Phonias) apfelbecki Csiki, 1908
Pterostichus (Phonias) diligens Sturm, 1824
Pterostichus (Phonias) taksonyis Csiki, 1930
Pterostichus (Platysma) niger (Schaller, 1783)⇒ Pterostichus selmanni selmanni Duftschmid, 1812
Pterostichus niger niger Schaller, 1783
Pterostichus (Pseudomaseus) anthracinus Illiger, 1798
Pterostichus anthracinus anthracinus Illiger, 1798
Pterostichus anthracinus biimpressus Küster, 1853
Pterostichus (Pseudomaseus) gracilis Dejean, 1828
Pterostichus (Pseudomaseus) minor Gyllenhal, 1827
Pterostichus (Pseudomaseus) nigrita Paykull, 1790
Pterostichus nigrita nigrita
Pterostichus (Pseudosteropus) cognatus (Dejean, 1831)⇒ Pterostichus aethiops Panzer, 1796
Pterostichus (Pseudosteropus) schmidtii Chaudoir, 1838
Pterostichus (Pterostichus) bruckii Schaum, 1859
Pterostichus (Pterostichus) fasciatopunctatus Creutzer, 1799
Pterostichus fasciatopunctatus fasciatopunctatus Creutzer, 1799
Pterostichus (Rhagadus) thorectes Bates, 1873
Pterostichus crenatus (Duftschmid, 1812)⇒ Pterostichus vernalis Panzer, 1796
Scybalicus oblongiusculus Dejean, 1829
Speluncarius (Speluncarius) anophthalmus Reitter, 1886
Speluncarius biokovensis Hlavac & Lak..., 2016
Sphodrus leucophthalmus Linnaeus, 1758
Stenolophus (Egadroma) marginatus Dejean, 1829
Stenolophus (Stenolophus) abdominalis Géné, 1836
Stenolophus abdominalis persicus Mannerheim, 1844
Stenolophus (Stenolophus) discophorus Fischer von ..., 1823
Stenolophus (Stenolophus) mixtus Herbst, 1784
Stenolophus (Stenolophus) narentinus Drovenik & Peks, 1999
Stenolophus (Stenolophus) proximus Dejean, 1829
Stenolophus (Stenolophus) skrimshiranus Stephens, 1828
Stenolophus (Stenolophus) steveni Krynicki, 1832
Stenolophus (Stenolophus) teutonus Schrank, 1781
Stomis (Stomis) rostratus Duftschmid, 1812
Stomis rostratus rostratus Sturm, 1812
Syntomus foveatus Geoffroy, 1785
Syntomus impressus Dejean, 1825
Syntomus impressus impressus Dejean, 1825
Syntomus obscuroguttatus Duftschmid, 1812
Syntomus pallipes Dejean, 1825
Syntomus truncatellus Linnaeus, 1761
Synuchus (Synuchus) vivalis Illiger, 1798
Synuchus vivalis vivalis (Illiger, 1798)⇒ Synuchus vivalis Illiger, 1798
Trechicus nigriceps (Dejean, 1831)⇒ Perigona nigriceps Dejean, 1831
Trichotichnus (Trichotichnus) laevicollis Duftschmid, 1812
Zabrus (Pelor) bischoffi J.Muller, 1936
Zabrus (Pelor) incrassatus Ahrens, 1814
Zabrus (Zabrus) tenebrioides Goeze, 1777Corn ground beetle
Zuphium (Zuphium) olens P.Rossi, 1790

Subnodes of Harpalinae with occurrences in Croatia
SupertribeChlaeniitae
Bonelli, 1810
8
34		Genera
Species
Taxom

SupertribeDryptitae
Bonelli, 1810
4
8		Genera
Species
Taxom

SupertribeHarpalitae
Erwin, 1985
22
145		Genera
Species
Taxom

SupertribeLebiitae
Bonelli, 1810
15
44		Genera
Species
Taxom

SupertribePlatynitae
Bonelli, 1810
14
70		Genera
Species
Taxom

SupertribePterostichitae
Bonelli, 1810
10
140		Genera
Species
Taxom



Links, sources and further information
Harpalinae at artfakta.se
Harpalinae at dyntaxa.se
Harpalinae at itis.gov