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Checklist for Subfamily Harpalinae Bonelli, 1810
and region Caucasus (Eurasia)


Class
 		Insecta
Linnaeus, 1758
Insects, True insects		523
6140
18845		Families
Genera
Species		 
Taxom
 

Subclass
 		Pterygota
Lang, 1888
Winged insects		521
6137
18842		Families
Genera
Species		 
Taxom
 

Infraclass
 		Neoptera
van der Wulp, 1890
Modern Wing-folding insects		497
6044
18637		Families
Genera
Species		 
Taxom
 

Superorder
 		Holometabola
Heider, 1889
Holometabolous Insects		331
4886
16042		Families
Genera
Species		 
Taxom
 

Order
 		Coleoptera
Linnaeus, 1758
Beetles, Coleopterans		98
1952
8628		Families
Genera
Species		 
Taxom
 

Suborder
 		Adephaga
Schellenberg, 1806
Adephagans, Ground Beetles and Water Beetles		5
251
1505		Families
Genera
Species		 
Taxom
 

Superfamily
 		Caraboidea
Latreille, 1802
1
215
1447		Family
Genera
Species		 
Taxom
 
FamilyCarabidae
Latreille, 1802
Carabid beetles, Carabids, Ground Beetles, Tiger Beetles		215
1447		Genera
Species		 
Taxom
 
SubfamilyHarpalinae
Bonelli, 1810
120
682		Genera
Species		 
Taxom
 


Genera for Caucasus
120 Genera with 682 Species
#ABCDEGHILMNOPSTXZ
Abacetus Dejean, 19834SpeciesTaxom
Abacidus LeConte, 18522SpeciesTaxom
Abax Bonelli, 18121SpeciesTaxom
Acinopus Dejean, 18296SpeciesTaxom
Acupalpus Latreille, 188413SpeciesTaxom
Agonum Bonelli, 182921SpeciesTaxom
Amara Bonelli, 0Seed-Eating Ground Beetles77SpeciesTaxom
Amblystomus Erichson, 18836SpeciesTaxom
Anaulacus MacLeay, 18501SpeciesTaxom
Anchomenus Bonelli, 05SpeciesTaxom
Andrewesius Andrewes, 19981SpeciesTaxom
Anisodactylus Dejean, 19339SpeciesTaxom
Anthracus Motschulsky, 18572SpeciesTaxom
Apristus Chaudoir, 18462SpeciesTaxom
Atranus LeConte, 18281SpeciesTaxom
Auchmerus Andrewes, 19291SpeciesTaxom
Badister Clairville, 013SpeciesTaxom
Bradycellus Erichson, 18128SpeciesTaxom
Calathus Bonelli, 196142SpeciesTaxom
Calleida Latreille & ..., 18451SpeciesTaxom
Callistus Bonelli, 17752SpeciesTaxom
Calodromius Reitter, 17982SpeciesTaxom
Carterus Dana, 18435SpeciesTaxom
Cedrorum Borges & Ser..., 19933SpeciesTaxom
Cephalophonus Ganglbauer, 18541SpeciesTaxom
Chilotomus Chaudoir, 19622SpeciesTaxom
Chlaenius Bonelli, 1860Vivid Metallic Ground Beetle36SpeciesTaxom
Colliuris De Geer, 1774
Long-necked Ground Beetles0SpeciesTaxom
Coptodera Dejean, 18251SpeciesTaxom
Cosnania Dejean, 17671SpeciesTaxom
Craspedophorus Hope, 18501SpeciesTaxom
Cryptophonus Brandmayr & ..., 18293SpeciesTaxom
Curtonotus Stephens, 17961SpeciesTaxom
Cyclotrachelus Chaudoir, 18232SpeciesTaxom
Cymindis Latreille, 182320SpeciesTaxom
Daptus Fischer von ..., 18231SpeciesTaxom
Demetrias Bonelli, 18244SpeciesTaxom
Derostichus Motschulsky, 18591SpeciesTaxom
Diachromus Erichson, 17581SpeciesTaxom
Dicaelus Bonelli, 1826Notched-mouth Ground Beetles4SpeciesTaxom
Dicheirotrichus Jacquelin du..., 18298SpeciesTaxom
Dinodes Bonet, 18324SpeciesTaxom
Diplocheila Brullé, 18441SpeciesTaxom
Ditomus Dana, 19932SpeciesTaxom
Dixus Billberg, 18724SpeciesTaxom
Dolichoctis Schmidt-Goebel, 19301SpeciesTaxom
Dromius Bonelli, 18874SpeciesTaxom
Drypta Latreille, 17901SpeciesTaxom
Ega Laporte de C..., 19601SpeciesTaxom
Eucarterus Reitter, 18981SpeciesTaxom
Euphorticus Horn, 18311SpeciesTaxom
Galerita Fabricius, 1792False Bombardier Beetle2SpeciesTaxom
Glycia Chaudoir, 18381SpeciesTaxom
Graniger Motschulsky, 18211SpeciesTaxom
Gynandromorphus Dejean, 19053SpeciesTaxom
Harpalus Latreille, 181291SpeciesTaxom
Helluomorphoides Ball, 1831Flat-horned Ground Beetles1SpeciesTaxom
Idiomelas Tschitschérine, 18321SpeciesTaxom
Laemostenus Bonelli, 184523SpeciesTaxom
Lamprias Bonelli, 18031SpeciesTaxom
Lebia Latreille, 1890colorful foliage ground beetle25SpeciesTaxom
Leptotrachelus Latreille, 18011SpeciesTaxom
Licinus Latreille, 06SpeciesTaxom
Lindrothius Kurnakov, 196116SpeciesTaxom
Lionychus Wissmann, 18122SpeciesTaxom
Loxandrus LeConte, 18521SpeciesTaxom
Loxoncus Schmidt-Gobel, 18581SpeciesTaxom
Metablus Jedlicka, 18391SpeciesTaxom
Metadromius Bedel, 18841SpeciesTaxom
Microderes Faldermann, 18091SpeciesTaxom
Microlestes Schmidt-Goebel, 186312SpeciesTaxom
Mimodromius Chaudoir, 18341SpeciesTaxom
Mnuphorus Chaudoir, 18891SpeciesTaxom
Morion Latreille, 18431SpeciesTaxom
Notiobia Perty, 1830
0SpeciesTaxom
Notonomus Chaudoir, 18401SpeciesTaxom
Odacantha Paykull, 17661SpeciesTaxom
Oedematicus Bedel, 17941SpeciesTaxom
Oedesis Motschulsky, 18311SpeciesTaxom
Olisthopus Dejean, 19831SpeciesTaxom
Ophonus Dejean, 025SpeciesTaxom
Panagaeus Latreille, 17752SpeciesTaxom
Pangus Dejean, 18181SpeciesTaxom
Paradromius Fowler, 18263SpeciesTaxom
Paranchus Lindroth, 17961SpeciesTaxom
Parazuphium Jeannel, 18983SpeciesTaxom
Parophonus Ganglbauer, 18296SpeciesTaxom
Penthus Chaudoir, 18381SpeciesTaxom
Perigona Laporte, 18311SpeciesTaxom
Philorhizus Hope, 18986SpeciesTaxom
Pinacodera Schaum, 18312SpeciesTaxom
Platyderus Stephens, 18323SpeciesTaxom
Platynus Bonelli, 18284SpeciesTaxom
Platytarus Fairmaire, 18262SpeciesTaxom
Poecilus Bonelli, 198324SpeciesTaxom
Polistichus Bonelli, 17902SpeciesTaxom
Psammodromius Peyerimhoff, 19242SpeciesTaxom
Pseudanchomenus Tarnier, 18601SpeciesTaxom
Pseudoophonus Motschulsky, 17743SpeciesTaxom
Pseudotaphoxenus Schaufuss, 18433SpeciesTaxom
Pterostichus Bonelli, 2015Woodland Ground Beetle125SpeciesTaxom
Selenophorus Dejean, 17982SpeciesTaxom
Singilis Rambur, 19631SpeciesTaxom
Sphelodon Townes, 18881SpeciesTaxom
Sphodrus Clairville, 17581SpeciesTaxom
Stenocrepis Chaudoir, 18351SpeciesTaxom
Stenolophus Dejean, 1781Seedcorn Beetle15SpeciesTaxom
Stomis Clairville, 19044SpeciesTaxom
Strigia Brullé, 18011SpeciesTaxom
Syntomus Hope, 17618SpeciesTaxom
Synuchus Gyllenhal, 17982SpeciesTaxom
Taphoxenus Motschulsky, 19523SpeciesTaxom
Tetragonoderus Dejean, 18242SpeciesTaxom
Thermoscelis Putzeys, 18883SpeciesTaxom
Trichocellus Ganglbauer, 18271SpeciesTaxom
Trichotichnus A. Morawitz, 18122SpeciesTaxom
Tschitscherinellus Csiki, 18502SpeciesTaxom
Xanthomelina Iablokoff-Kh..., 19241SpeciesTaxom
Zabrus Clairville, 177727SpeciesTaxom
Zuphium Latreille, 18584SpeciesTaxom


Species list for Caucasus
682 Species and 167 Subspecies
Abacetus (Astigis) inexpectatus Kryzhanovski..., 1983
Abacetus (Astigis) quadripustulatus Peyron, 1858
Abacetus (Triaenabacetus) transcaucasicus Chaudoir, 1876
Abacetus inexspectatus Abdurakhmano..., 1983
Abacidus atratus (Newman, 1838)⇒ Pterostichus atratus Gistel, 1857
Abacidus sculptus (LeConte, 1852)⇒ Pterostichus sculptus LeConte, 1853
Abax (Abacopercus) carinatus Duftschmid, 1812
Acinopus (Acinopus) laevigatus Ménétriés, 1832
Acinopus laevigatus laevigatus Ménétriés..., 1832
Acinopus (Acinopus) picipes Olivier, 1795
Acinopus (Haplacinopus) striolatus Zoubkoff, 1833
Acinopus (Oedematicus) megacephalus (P. Rossi, 1794)⇒ Acinopus picipes Olivier, 1795
Acinopus (Osimus) ammophilus Dejean, 1829
Acupalpus (Acupalpus) brunnipes Sturm, 1825
Acupalpus (Acupalpus) dubius Schilsky, 1888
Acupalpus (Acupalpus) elegans Dejean, 1829
Acupalpus (Acupalpus) exiguus Dejean, 1829
Acupalpus (Acupalpus) flavicollis Sturm, 1825
Acupalpus (Acupalpus) luteatus (Duftschmid, 1812)⇒ Acupalpus dubius Schilsky, 1888
Acupalpus (Acupalpus) maculatus Schaum, 1860
Acupalpus (Acupalpus) meridianus Linnaeus, 1761Midday harp ground beetle
Acupalpus (Acupalpus) parvulus Sturm, 1825
Acupalpus (Acupalpus) suturalis (Dejean, 1829)⇒ Acupalpus meridianus Linnaeus, 1761
Acupalpus suturalis suturalis Dejean, 1829
Acupalpus (Ancylostria) interstitialis Reitter, 1884
Acupalpus (Tachistodes) paludicola Reitter, 1884
Agonum (Agonum) carbonarium Dejean, 1828
Agonum carbonarium hexacoelum Chaudoir, 1850
Agonum (Agonum) chalconotum Ménétriés, 1832
Agonum (Agonum) marginatum Linnaeus, 1758
Agonum (Agonum) mesostictum Bates, 1889
Agonum (Agonum) monachum Duftschmid, 1812
Agonum monachum monachum Duftschmid, 1812
Agonum (Agonum) muelleri Herbst, 1784Müller'S Harp Ground Beetle
Agonum (Agonum) rugicolle (Chaudoir, 1846)⇒ Agonum rugaticolle Gemminger & ..., 1868
Agonum (Melanagonum) gisellae Csiki, 1931
Agonum (Olisares) corvus (LeConte, 1860)⇒ Agonum viridicupreum fulgidicolle Erichson, 1841
Agonum (Olisares) extensicolle Say, 1823
Agonum (Olisares) hypocrita Apfelbeck, 1904
Agonum (Olisares) longicorne Chaudoir, 1846
Agonum (Olisares) lugens Duftschmid, 1812
Agonum (Olisares) pallipes Fabricius, 1787Spotted Warner Valley Dunes June beetle
Agonum (Olisares) viridicupreum Goeze, 1777
Agonum gracillipes gracilippes
Agonum rugaticolle Gemminger & ..., 1868
Agonum sexipunctatum
Agonum thoracicum (Fischer von ..., 1829)⇒ Agonum viridicupreum viridicupreum Goeze, 1777
Amara (Allobradytus) armeniaca Motschulsky, 1839
Amara (Amara) aenea De Geer, 1774Common sun beetle
Amara (Amara) anthobia A. Villa & G..., 1833Italy harp ground beetle
Amara (Amara) communis Panzer, 1797Common harp ground beetle
Amara (Amara) convexior Stephens, 1828
Amara (Amara) curta Dejean, 1828
Amara (Amara) eurynota Panzer, 1796
Amara (Amara) famelica C. Zimmermann, 1832Early sunshiner
Amara (Amara) familiaris Duftschmid, 1812Bareyed goby
Amara (Amara) littorea C.C. Thomson, 1857
Amara (Amara) lucida Duftschmid, 1812
Amara (Amara) lunicollis Schiodte, 1837Golden-banded goby
Amara (Amara) montivaga Sturm, 1825
Amara (Amara) morio Ménétriés, 1832
Amara morio morio Ménétriés..., 1832
Amara (Amara) ovata (Fabricius, 1792)⇒ Amara tricuspidata Dejean, 1831
Amara (Amara) proxima Putzeys, 1866
Amara (Amara) similata Gyllenhal, 1810
Amara (Amara) tibialis (Paykull, 1798)⇒ Amara tumida tumida Morawitz, 1862
Amara (Amarocelia) erratica Duftschmid, 1812
Amara (Amathitis) abdominalis Motschulsky, 1844
Amara (Amathitis) microdera Chaudoir, 1844
Amara (Amathitis) parvicollis Gebler, 1833
Amara (Amathitis) rufescens (Dejean, 1829)⇒ Amara aulica Panzer, 1796
Amara rufescens rufescens Dejean, 1829
Amara (Bradytus) apricaria Paykull, 1790Exposed sun beetle
Amara (Bradytus) consularis Duftschmid, 1812
Amara (Bradytus) crenata Dejean, 1828
Amara (Bradytus) fulva (O.F. Muller, 1776)⇒ Amara apricaria Paykull, 1790
Amara (Celia) bifrons Gyllenhal, 1810Two-faced harp ground beetle
Amara (Celia) cursitans Zimmermann, 1832
Amara (Celia) erratica Duftschmid, 1812Bareyed goby
Amara municipalis municipalis Duftschmid, 1812
Amara (Celia) musculis Say, 1823
Amara (Celia) praetermissa C.R. Sahlberg, 1827
Amara quenseli silvicola Zimm., 1832
Amara (Celia) sabulosa Audinet-Serv..., 1821
Amara (Celia) sollicita Pantel, 1888
Amara (Celia) viridescens Reitter, 1883
Amara (Curtonotus) aulica Panzer, 1796Princely harp ground beetle
Amara (Curtonotus) carinata LeConte, 1848
Amara (Curtonotus) convexiuscula Marsham, 1802
Amara (Curtonotus) cribricollis Chaudoir, 1846
Amara (Curtonotus) gebleri Dejean, 1831
Amara (Curtonotus) propinqua Ménétriés, 1832
Amara (Curtonotus) torrida Panzer, 1796
Amara tumida tumida Morawitz, 1862
Amara (Cyrtonotus) torrida Panzer, 1796
Amara (Harpalodema) bradytoides Reitter, 1889
Amara (Leiocnemis) calathoides Putzeys, 1866
Amara (Leiocnemis) cordicollis (Ménétriés, 1832)⇒ Amara microdera Chaudoir, 1844
Amara (Paracelia) pulchra Baliani, 1943
Amara pulchra alagoensis Hieke, 2006
Amara pulchra pulchra Baliani, 1943
Amara (Paracelia) quenseli Schoenher, 1806Bearded goby
Amara (Paracelia) saxicola C. Zimmermann, 1832
Amara saxicola saxicola Zimmermann, 1832
Amara (Parapercosia) taurica Motschulsky, 1844
Amara (Percosia) equestris Duftschmid, 1812
Amara equestris equestris Duftschmid, 1812
Amara equestris pastica Dejean, 1831
Amara (Phanerodonta) punctipennis Reitter, 1889
Amara (Xenocelia) ambulans C. Zimmermann, 1832
Amara (Xenocelia) bischoffi Jedlička, 1946
Amara (Xenocelia) cursitans C. Zimmermann, 1832
Amara (Xenocelia) elevata (Motschulsky, 1844)⇒ Amara infima Duftschmid, 1812
Amara (Xenocelia) fusca (Dejean, 1828)⇒ Amara quenseli silvicola Zimm., 1832
Amara (Xenocelia) ingenua Duftschmid, 1812
Amara (Xenocelia) municipalis Duftschmid, 1812
Amara (Zezea) chaudoiri Schaum, 1858
Amara chaudoiri chaudoiri Schaum, 1858
Amara chaudoiri transcaucasiens Hieke, 1970
Amara (Zezea) reflexicollis Motschulsky, 1844
Amara reflexicollis reflexicollis Motschulsky, 1845
Amara (Zezea) strandi Lutshnik, 1933
Amara (Zezea) tricuspidata Dejean, 1831
Amara caucasicus
Amblystomus levantinus Reitter, 1883
Amblystomus levantinus levantinus Reitter, 1883
Amblystomus metallescens Dejean, 1829
Amblystomus niger (Heer, 1841)⇒ Amblystomus nigrinus Csiki, 1931
Amblystomus nigrinus Csiki, 1931
Amblystomus rectangulus Reitter, 1883
Anaulacus (Aephnidius) ruficornis Chaudoir, 1850
Anchomenus (Anchodemus) kurnakovi Kryzhanovskij, 1983
Anchomenus (Anchomenus) dorsalis Pontoppidan, 1763
Anchomenus dorsalis dorsalis
Anchomenus (Anchomenus) turkestanicus Ballion, 1871
Anchomenus subdepressa
Andrewesius arlettae Morvan, 1998
Anisodactylus (Anadaptus) discoideus Dejean, 1831
Anisodactylus (Anadaptus) sanctaecrucis Fabricius, 1798
Anisodactylus (Anisodactylus) binotatus Fabricius, 1787Two-marked harp ground beetle
Anisodactylus (Anisodactylus) nemorivagus Duftschmid, 1812Heath Short-spur
Anisodactylus (Hexatrichus) poeciloides Stephens, 1828Saltmarsh Short-spur
Anisodactylus poeciloides pseudoaeneus Dejean, 1829
Anisodactylus (Pseudanisodactylus) signatus Panzer, 1796
Anisodactylus (Pseudodichirus) intermedius Dejean, 1829
Anisodactylus pueli pueli Schauberger, 1933
Anthracus consputus Duftschmid, 1812
Anthracus longicornis Schaum, 1857
Apristus reticulatus Schaum, 1857
Apristus subaeneus Chaudoir, 1846
Atranus pubescens Dejean, 1828Bynoe's goby
Auchmerus trachys Andrewes, 1929
Badister (Badister) bullatus Schrank, 1798
Badister (Badister) denticulatus Wrase, 1995
Badister (Badister) flavipes LeConte, 1853
Badister (Badister) meridionalis Puel, 1925
Badister (Badister) unipustulatus Bonelli, 1813
Badister (Bandia) dilatatus Chaudoir, 1837
Badister peltatus peltatus Panzer, 1796
Badister (Baudia) collaris (Motschulsky, 1844)⇒ Badister peltatus peltatus Panzer, 1796
Badister (Baudia) dilatatus Chaudoir, 1837
Badister (Baudia) grandiceps Casey, 1920
Badister (Trimorphus) sodalis Duftschmid, 1812
Badister billatus
Badister bipunctulatus
Bradycellus (Bradycellus) heinzi Jaeger, 1990Engelhardt´s cave beetle
Bradycellus (Stenocellus) nebulosus LeConte, 1853
Bradycellus (Stenocellus) rupestris Say, 1823Rich Mountain cave beetle
Bradycellus (Triliarthrus) caucasicus Chaudoir, 1846
Bradycellus (Triliarthrus) chavesi Alluaud, 1919Ground-beetle
Bradycellus (Triliarthrus) csikii Laczó, 1912
Bradycellus (Triliarthrus) distinctus Dejean, 1829
Bradycellus (Triliarthrus) verbasci Duftschmid, 1812
Calathus (Calathus) alternans Faldermann, 1836
Calathus (Calathus) distinguendus (Chaudoir, 1846)⇒ Calathus gregarius Say, 1823
Calathus (Calathus) fracassii Heyden, 1908
Calathus fracassii fracassii Heyden, 1908
Calathus (Calathus) fuscipes Goeze, 1777Dark-footed harp ground beetle
Calathus fuscipes fuscipes Goeze, 1777
Calathus fuscipes punctipennis Germar, 1824
Calathus (Calathus) libanensis Putzeys, 1873
Calathus libanensis pluriseriatus Putzeys, 1873
Calathus (Calathus) longicollis Motschulsky, 1865
Calathus (Calathus) reflexicollis Faldermann, 1839
Calathus (Lauricalathus) carvalhoi Serrano & Bo..., 1986Ground Beetle
Calathus (Neocalathus) ambiguus Paykull, 1790
Calathus ambiguus ambiguus Paykull, 1790
Calathus (Neocalathus) erratus C.R.Sahlberg, 1827
Calathus erratus marginicollis Chaudoir, 1846
Calathus (Neocalathus) extensicollis Putzeys, 1873Ground beetle
Calathus (Neocalathus) gregarius Say, 1823
Calathus (Neocalathus) lundbladi Colas, 1938Ground beetle
Calathus (Neocalathus) melanocephalus Linnaeus, 1758
Calathus melanocephalus antoinei Puel, 1939
Calathus melanocephalus furvus Tschitschérine, 1895
Calathus melanocephalus melanocephalus Linné, 1758
Calathus melanocephalus paphlagonicus Maran, 1935
Calathus melanocephalus purkynei Maran, 1935
Calathus (Neocalathus) mollis Marsham, 1802
Calathus mollis mollis Marsham, 1802
Calathus (Neocalathus) peltatus Kolenati, 1845
Calathus (Neocalathus) vicenteorum Schatzmayr, 1937Ground beetle
Calathus (Trichocalathus) syriacus Chaudoir, 1863
Calathus aequistriatus Kurnakov, 1961
Calathus caucasicus Chaudoir, 1846
Calathus caucasicus caucasicus Chaudoir, 1846
Calathus caucasicus orbicollis Motschulsky, 1865
Calathus grandiceps Kurnakov, 1961
Calathus grandiceps grandiceps Kurnakov, 1961
Calathus gregaria Say, 1823
Calathus horsti Reitter, 1888
Calathus mandibularis Kurnakov, 1961
Calathus robustus Kurnakov, 1961
Calathus stricticaudis Kurnakov, 1961
Calathus subpraestans Kurnakov, 1961
Calleida cordicollis (Putzeys, 1845)⇒ Calleida marginicollis Chaudoir, 1844
Callistus lunatus Fabricius, 1775
Callistus lunatus lunatus Fabricius, 1775
Calodromius sphex Andrewes, 1933
Calodromius spilotus Illiger, 1798
Carterus (Pristocarterus) angustipennis Chaudoir, 1852
Carterus angustipennis lutshniki Zamotailov, 1988
Carterus (Pristocarterus) angustus Ménétriés, 1832
Carterus (Pristocarterus) gilvipes Piochard de ..., 1873
Carterus (Pristocarterus) rufipes Chaudoir, 1843
Cedrorum azoricus Borges & Ser..., 1993
Cedrorum azoricus azoricus Borges & Ser..., 1993
Cedrorum azoricus caveirensis Borges & Ser..., 1993
Cephalophonus cephalotes Fairmaire & ..., 1854
Chilotomus alexandri Kalashyan, 1999
Chilotomus arnoldii Kryzhanovskij, 1962
Chlaenius (Agostenus) alutaceus Gebler, 1829
Chlaenius (Amblygenius) dimidiatus (Chaudoir, 1842)⇒ Chlaenius ruficauda Chaudoir, 1856
Chlaenius (Anomoglossus) amoenus (Dejean, 1831)⇒ Chlaenius brevilabris Le Conte, 1848
Chlaenius (Callistometus) ruficauda Chaudoir, 1856Mudskipper
Chlaenius (Chlaeniellus) flavipes Ménétriés, 1832
Chlaenius (Chlaeniellus) kindermanni Chaudoir, 1856
Chlaenius kindermanni chrysothorax Krynitzky, 1832
Chlaenius (Chlaeniellus) melampus Ménétriés, 1849
Chlaenius (Chlaeniellus) nemoralis Say, 1823
Chlaenius (Chlaeniellus) nigricornis Fabricius, 1787
Chlaenius (Chlaeniellus) terminatus Dejean, 1826
Chlaenius (Chlaeniellus) tricolor Dejean, 1826Prickly scorpion's-tail
Chlaenius (Chlaeniellus) tristis (Schaller, 1783)⇒ Chlaenius moestus Csiki, 1931
Chlaenius tristis tristis Schaller, 1783
Chlaenius (Chlaeniellus) vestitus Paykull, 1790
Chlaenius (Chlaenites) spoliatus P. Rossi, 1792
Chlaenius spoliatus spoliatus P. Rossi, 1792
Chlaenius (Chlaenius) festivus Panzer, 1796
Chlaenius festivus festivus Panzer, 1796
Chlaenius (Chlaenius) reitteri Jakobson, 1906
Chlaenius (Epomis) circumscriptus Duftschmid, 1812
Chlaenius (Eurydactylus) aurolimbatus LaFerté-Séne..., 1851
Chlaenius (Eurydactylus) tomentosus Say, 1823
Chlaenius (Homalolachnus) moestus Csiki, 1931
Chlaenius (Stenochlaenius) coeruleus (Steven, 1809)⇒ Chlaenius aurolimbatus LaFerté-Séne..., 1851
Chlaenius (Stenochlaenius) lederi Reitter, 1888
Chlaenius (Trichochlaenius) aeneocephalus Dejean, 1826
Chlaenius aeneocephalus aeneocephalus Dejean, 1826
Chlaenius (Trichochlaenius) gotschii Chaudoir, 1846
Chlaenius (Trichochlaenius) steveni Quensel, 1806
Chlaenius (Trichochlaenius) stevenii Quensel In S..., 1806
Chlaenius (Wilmernstus) ophonoides Fairmaire, 1843
Chlaenius armenus Jedlicka, 1950
Chlaenius cruralis Fischer von ..., 1829
Chlaenius decipiens L.Dufour, 1820
Chlaenius viridis Montrouzier, 1860
Coptodera (Coptodera) aerata Dejean, 1825
Cosnania pensylvanica Linnaeus, 1767
Craspedophorus australasiae Chaudoir, 1850
Cryptophonus litigiosus Dejean, 1829
Cryptophonus melancholicus Dejean, 1829
Cryptophonus tenebrosus Dejean, 1829
Curtonotus aulicus (Panzer, 1796)⇒ Amara aulica Panzer, 1796
Cyclotrachelus (Evarthrus) incisus LeConte, 1848
Cyclotrachelus (Evarthrus) sigillatus Say, 1823
Cymindis (Cymindis) axillaris (Fabricius, 1794)⇒ Cymindis scapularis Schaum, 1857
Cymindis axillaris axillaris Fabricius, 1794
Cymindis axillaris omiades Faldermann, 1836
Cymindis (Cymindis) ovipennnis Motschulsky, 1844
Cymindis (Cymindis) scapularis Schaum, 1857
Cymindis scapularis scapularis Schaum, 1857
Cymindis (Eremocymindis) pallidula Chaudoir, 1846
Cymindis hierichontica hierichontica Reiche & Saulcy, 1855
Cymindis (Menas) miliaris Fabricius, 1801
Cymindis (Neomenas) antonowi Semenov, 1891
Cymindis (Tarsostinus) equestris Gebler, 1825
Cymindis (Tarulus) andreae Ménétriés, 1832
Cymindis (Tarulus) decora Fischer von ..., 1829
Cymindis (Tarulus) intermedia Chaudoir, 1873
Cymindis (Tarulus) lineata Quensel In S..., 1806
Cymindis (Tarulus) ovipennis Motschulsky, 1844
Cymindis (Tarulus) pallidula Chaudoir, 1846
Cymindis (Tarulus) picta (Pallas, 1771)⇒ Mimodromius lepidus Brullé, 1834
Cymindis picta picta Pallas, 1771
Cymindis (Tarulus) pilosa (Say, 1823)⇒ Cymindis faldermanni Gistel, 1838
Daptus vittatus Fischer von ..., 1823
Demetrias (Aetophorus) imperialis Germar, 1824
Demetrias (Demetrias) longicornis Chaudoir, 1846
Demetrias (Demetrias) monostigma Samouelle, 1819
Demetrias (Risophilus) imperialis Germar, 1824
Derostichus caucasicus Motschulsky, 1859
Diachromus germanus Linnaeus, 1758
Dicaelus (Dicaelus) purpuratus Bonelli, 1813
Dicaelus (Paradicaelus) dilatatus Say, 1823
Dicaelus (Paradicaelus) elongatus Bonelli, 1813
Dicaelus (Paradicaelus) furvus Dejean, 1826Sponge goby
Dicheirotrichus (Cardiostenus) microderus Solsky, 1874
Dicheirotrichus (Trichocellus) discolor Faldermann, 1836
Dicheirotrichus discolor discolor Faldermann, 1836
Dicheirotrichus (Trichocellus) lacustris L.Redtenbacher, 1858
Dicheirotrichus (Trichocellus) obscuricornis Reitter In T..., 1899
Dicheirotrichus (Trichocellus) pallidus Dejean, 1829
Dicheirotrichus (Trichocellus) placidus (Gyllenhal, 1827)⇒ Trichocellus placidus Gyllenhal, 1827
Dicheirotrichus (Trichocellus) ustulatus Dejean, 1829
Dinodes (Dinodes) armenus Jedlička, 1950
Dinodes (Dinodes) cruralis Fischer von ..., 1829
Dinodes (Dinodes) decipiens (L. Dufour, 1820)⇒ 
Dinodes (Dinodes) viridis (Ménétriés, 1832)⇒ Chlaenius nigricornis Fabricius, 1787
Diplocheila (Isorembus) striatopunctata LeConte, 1844Aboma de mar
Ditomus calydonius P. Rossi, 1790
Ditomus calydonius oriens Dvorak, 1993
Dixus eremita Dejean, 1825
Dixus obscuratus
Dixus obscurus Dejean, 1825
Dixus semicylindricus Piochard de ..., 1872
Dolichoctis (Dolichoctis) microdera Andrewes, 1930
Dromius (Dromius) meridionalis Dejean, 1825
Dromius (Dromius) piceus Dejean, 1831
Dromius (Dromius) quadrimaculatus Linnaeus, 1758
Dromius (Dromius) semiplagiatus Reitter, 1887
Drypta (Drypta) dentata P.Rossi, 1790
Ega sallei (Park, 1960)⇒ Calybe sallei Chevrolat, 1839
Eucarterus sparsutus Reitter, 1898
Euphorticus pubescens Dejean, 1831
Galerita (Progaleritina) bicolor Drury, 1773False bombardier beetle
Galerita (Progaleritina) janus Fabricius, 1792
Glycia spencei Gistel, 1838
Graniger cordicollis Audinet-Serv..., 1821
Gynandromorphus etruscus Quensel, 1806
Gynandromorphus etruscus etruscus Quensel, 1806
Gynandromorphus peyroni Carret, 1905
Harpalus (Acardystus) atratus Latreille, 1804
Harpalus (Acardystus) attenuatus Stephens, 1828
Harpalus (Acardystus) pygmaeus Dejean, 1829
Harpalus (Actephilus) albanicus Reitter, 1900
Harpalus (Actephilus) pumilus Sturm, 1818
Harpalus (Amblystus) latus Linnaeus, 1758
Harpalus (Amblystus) rubripes Dufischmid, 1812Bareyed goby
Harpalus (Artabas) dispar Dejean, 1829
Harpalus dispar elegantulus Ménétries, 1832
Harpalus (Cephalophonus) cephalotes (Fairmaire & ..., 1854)⇒ Harpalus capito A.Morawitz, 1862
Harpalus cephalotes cephalotes
Harpalus cephalotes somcheticus Schauberger, 1933
Harpalus (Cryptophonus) litigiosus (Dejean, 1829)⇒ Cryptophonus litigiosus Dejean, 1829
Harpalus litigiosus litigiosus Dejean, 1829
Harpalus (Cryptophonus) melancholicus (Dejean, 1829)⇒ Cryptophonus melancholicus Dejean, 1829
Harpalus melancholicus melancholicus
Harpalus (Cryptophonus) tenebrosus (Dejean, 1829)⇒ 
Harpalus tenebrosus tenebrosus
Harpalus (Harpalobius) froelichii Sturm, 1818Brush-thighed Seed-eater
Harpalus (Harpalobius) fuscipalpis Sturm, 1818
Harpalus (Harpalobius) hirtipes Panzer, 1796
Harpalus (Harpaloderus) honestus Dufischmid, 1812St. Bees Seed-eater
Harpalus neglectus neglectus Audinet-Serv..., 1821
Harpalus (Harpaloderus) rufipalpis Sturm, 1818
Harpalus rufipalpis rufipalpis Sturm, 1818
Harpalus (Harpalophonus) circumpunctatus Chaudoir, 1846
Harpalus (Harpalophonus) hospes Sturm, 1818
Harpalus hospes armenus K.Daniel, 1904
Harpalus (Harpalus) aeneipennis Faldermann, 1836
Harpalus (Harpalus) affinis (Schrank, 1781)⇒ Harpalus serripes ernsti Kataev, 1995
Harpalus (Harpalus) akinini Tschitschérine, 1895
Harpalus (Harpalus) alpivagus Tschitschérine, 1899
Harpalus (Harpalus) araraticus Mlynar, 1979
Harpalus (Harpalus) brevicornis Germar, 1824
Harpalus (Harpalus) breviusculus Chaudoir, 1846
Harpalus (Harpalus) caeruleatus Bates, 1878
Harpalus (Harpalus) caspius Steven, 1806
Harpalus (Harpalus) chrysopus Reitter, 1887
Harpalus chrysopus abasinus Rost, 1892
Harpalus chrysopus contumax Lutshnik, 1933
Harpalus (Harpalus) cisteloides Motschulsky, 1844
Harpalus cisteloides schouberti Tschitschérine, 1898
Harpalus (Harpalus) convexus (Faldermann, 1836)⇒ 
Harpalus (Harpalus) inexspectatus Kataev, 1989
Harpalus (Harpalus) meteorus Basilewsky, 1946
Harpalus oblitus oblitus Dejean, 1829
Harpalus (Harpalus) quadratus Chaudoir, 1846
Harpalus (Harpalus) reflexus Putzeys, 1878
Harpalus reflexus anadoluensis Kataev, 1993
Harpalus reflexus reflexus Putzeys, 1878
Harpalus (Harpalus) saxicola Dejean, 1829
Harpalus (Harpalus) smyrnensis Heyden, 1888
Harpalus smyrnensis medicus Kataev, 1993
Harpalus smyrnensis raddei Tschitschérine, 1897
Harpalus (Harpalus) somnulentus Dejean, 1829Checked goby
Harpalus (Harpalus) subcylindricus Dejean, 1829
Harpalus (Harpalus) subtruncatus (Chaudoir, 1846)⇒ Harpalus rubripes Dufischmid, 1812
Harpalus (Harpalus) xanthopus Gemminger & ..., 1868
Harpalus xanthopus winkleri Schauberger, 1923
Harpalus (Harpalus) zabroides Dejean, 1829
Harpalus (Lasioharpalus) cupreus Dejean, 1829
Harpalus cupreus fastuosus Faldermann, 1835
Harpalus (Lasioharpalus) dimidiatus P. Rossi, 1790
Harpalus (Lasioharpalus) distinguendus Duftschmid, 1812
Harpalus distinguendus distinguendus Duftschmid, 1812
Harpalus (Lasioharpalus) smaragdinus Duftschmid, 1812
Harpalus (Opadius) laevipes Zetterstedt, 1828
Harpalus (Pheuginus) flavicornis Dejean, 1829
Harpalus flavicornis flavicornis Dejean, 1829
Harpalus (Pheuginus) politus Dejean, 1829
Harpalus politus vasilinini Lutshnik, 1916
Harpalus (Pheuginus) serripes Quensel, 1806
Harpalus serripes ernsti Kataev, 1995
Harpalus serripes serripes Quensel, 1806
Harpalus (Pheuginus) tardus Panzer, 1796
Harpalus (Plectralidus) amplicollis Ménétriés, 1848
Harpalus (Plectralidus) angulatus Putzeys, 1878
Harpalus angulatus angulatus Putzeys, 1878
Harpalus (Plectralidus) fuscicornis Ménétriés, 1832
Harpalus (Plectralidus) metallinus Ménétriés, 1836
Harpalus metallinus metallinus Ménétries, 1836
Harpalus (Plectralidus) pulvinatus Ménétriés, 1848
Harpalus pulvinatus lubricus Reitter, 1900
Harpalus (Pseudoophonus) calceatus (Duftschmid, 1812)⇒ Pseudoophonus calceatus Duftschmid, 1812
Harpalus (Pseudoophonus) capito A.Morawitz, 1862
Harpalus (Pseudoophonus) compar (LeConte, 1848)⇒ Harpalus cohni Ball, 1972
Harpalus (Pseudoophonus) erythropus Dejean, 1829Checked goby
Harpalus (Pseudoophonus) griseus (Panzer, 1796)⇒ Pseudoophonus griseus Panzer, 1797
Harpalus (Pseudoophonus) pensylvanicus DeGeer, 1774Pennsylvania Dingy Ground Beetle
Harpalus (Pseudoophonus) rufipes (DeGeer, 1774)⇒ 
Harpalus (Semiophonus) signaticornis Duftschmid, 1812
Helluomorphoides nigripennis Dejean, 1831
Idiomelas (Idiomelas) morio Ménétriés, 1832
Laemostenus (Antisphodroides) levushkini Vareschagina, 1986
Laemostenus (Antisphodroides) suramensis Reitter, 1885
Laemostenus (Antisphodrus) bicolor Reitter, 1890
Laemostenus bicolor bicolor Reitter, 1890
Laemostenus (Laemostenus) caspius Meneiries, 1832
Laemostenus (Laemostenus) complanatus Dejean, 1828Bynoe's goby
Laemostenus (Pristonychus) cimmerius Fischer von ..., 1823
Laemostenus cimmerius cimmerius Fischer-Wald..., 1823
Laemostenus (Pristonychus) gratus Faldermann, 1836
Laemostenus (Pristonychus) lederi Reitter, 1885
Laemostenus lederi arenicus
Laemostenus lederi khnzoriani Kalashyan, 1983
Laemostenus lederi lederi Reitter, 1885
Laemostenus (Pristonychus) mannerheimi Kolenati, 1845
Laemostenus mannerheimi mannerheimi Kolenati, 1845
Laemostenus (Pristonychus) piceus Motschulsky, 1850
Laemostenus (Pristonychus) pretiosus Faldermann, 1837
Laemostenus (Pristonychus) sericeus Fischer von ..., 1824
Laemostenus sericeus hepaticus Faldermann, 1836
Laemostenus sericeus sericeus Fischer von ..., 1824
Laemostenus sericeus tauricus Dejean, 1828
Laemostenus mannerheimii Kolenati, 1845
Laemostenus mannerheimii mannerheimii Kolenati, 1845
Lamprias chlorocephalus (J.J. Hoffmann, 1803)⇒ Lebia chlorocephala Hoffmann, 1803
Lebia (Glyciolebia) arcuata (Reiche & Saulcy, 1855)⇒ Singilis arcuatus Reiche, 1855
Lebia (Lamprias) chlorocephala Hoffmann, 1803
Lebia (Lamprias) cyanocephala Linnaeus, 1758Blue plunderer
Lebia cyanocephala cyanocephala Linnaeus, 1758
Lebia (Lamprias) festiva Faldermann, 1836
Lebia festiva festiva Faldermann, 1836
Lebia (Lebia) analis Dejean, 1825
Lebia (Lebia) cruxminor Linnaeus, 1758
Lebia cruxminor cruxminor Linnaeus, 1758
Lebia (Lebia) fuscata (Dejean, 1825)⇒ Lebia fraterna Péringuey, 1896
Lebia (Lebia) humeralis Dejean, 1825
Lebia (Lebia) marginata Geoffroy, 1785
Lebia (Lebia) marginicollis Dejean, 1825
Lebia (Lebia) ornata Say, 1823
Lebia (Lebia) pulchella Dejean, 1826Beautiful Banded Lebia
Lebia (Lebia) scapularis (Geoffroy, 1785)⇒ Lebia solea Hentz, 1830
Lebia scapularis scapularis Geoffroy, 1785
Lebia (Lebia) solea Hentz, 1830
Lebia (Lebia) trimaculata (Villers, 1789)⇒ 
Lebia (Lebia) viridis Say, 1823Flower Lebia Beetle
Lebia (Loxopeza) atriventris Say, 1823
Lebia (Loxopeza) grandis Hentz, 1830Large Foliage Ground Beetle
Lebia (Stephana) holomera Chaudoir, 1871
Lebia (Stephana) turkestanica Jedlička, 1966
Lebia circumducta Reitter, 1890
Leptotrachelus dorsalis Fabricius, 1801
Licinus (Licinus) cassideus Fabricius, 1792
Licinus cassideus cassideus Fabricius, 1792
Licinus (Licinus) convexus Heyden, 1889
Licinus (Licinus) depressus (Paykull, 1790)⇒ Licinus cassideus cassideus Fabricius, 1792
Licinus (Licinus) punctatulus Fabricius, 1792
Licinus punctatulus punctatulus
Lindrothius aequistriatus Kurnakov, 1961
Lindrothius caucasicus Chaudoir, 1846
Lindrothius caucasicus caucasicus
Lindrothius caucasicus orbicollis Motschulsky, 1865
Lindrothius grandiceps Kurnakov, 1961
Lindrothius grandiceps grandiceps Kurnakov, 1961
Lindrothius horsti Reitter, 1888
Lindrothius laticaudis Kurnakov, 1961
Lindrothius mandibularis Kurnakov, 1961
Lindrothius praestans Heyden, 1885
Lindrothius pseudopraestans Kurnakov, 1961
Lindrothius recticaudis Kurnakov, 1961
Lindrothius robustus Kurnakov, 1961
Lindrothius sotshiensis Zamotajlov, 1997
Lindrothius stricticaudis Kurnakov, 1961
Lindrothius subpraestans Kurnakov, 1961
Lionychus (Lionychus) orientalis K.Daniel, 1901
Lionychus (Paralionychus) quadrillum Duftschmid, 1812
Loxandrus pusillus (LeConte, 1852)⇒ Loxandrus piciventris LeConte, 1848
Loxoncus (Loxoncus) procerus Schaum, 1858
Metablus paracenthesis Motschulsky, 1839
Metadromius signifer Reitter, 1884
Microderes (Microderes) brachypus Steven, 1809
Microlestes corticalis L. Dufour, 1820
Microlestes fissuralis Reitter, 1901
Microlestes fulvibasis Reitter, 1901
Microlestes luctuosus Holdhaus, 1904
Microlestes luctuosus luctuosus Holdhaus, 1904
Microlestes maurus Sturm, 1827
Microlestes maurus maurus Sturm, 1827
Microlestes minutulus Goeze, 1777
Microlestes negrita Wollaston, 1854
Microlestes plagiatus (Duftschmid, 1812)⇒ Microlestes corticalis L. Dufour, 1820
Microlestes politulus Reitter, 1901
Microlestes pusio LeConte, 1863
Mimodromius (Mimodromius) lepidus Brullé, 1834
Mnuphorus callistoides Reitter, 1889
Morion (Neomorion) olympicus L.Redtenbacher, 1843
Notonomus (Leiradira) marginatus Castelnau, 1840
Odacantha (Odacantha) melanura Linnaeus, 1766
Oedematicus megacephalus Rossi, 1794
Oedesis caucasicus Dejean, 1831
Olisthopus inclavatus Israelson, 1983Ground Beetle
Ophonus (Hesperophonus) azureus Fabricius, 1775
Ophonus (Hesperophonus) convexicollis Ménétriés, 1832
Ophonus (Hesperophonus) cribricollis Dejean, 1829
Ophonus (Hesperophonus) minimus Motschulsky, 1845
Ophonus (Hesperophonus) similis Dejean, 1829
Ophonus (Hesperophonus) subquadratus (Dejean, 1829)⇒ 
Ophonus (Macrophonus) oblongus Schaum, 1858
Ophonus (Metophonus) austrocaspicus Kataev & Bel..., 2001
Ophonus (Metophonus) austrocaspius Kataev & Bel..., 2001
Ophonus (Metophonus) cordatus Duftschmid, 1812
Ophonus (Metophonus) laticollis (Mannerheim, 1825)⇒ Ophonus puncticollis Paykull, 1798
Ophonus (Metophonus) melletii Heer, 1837Mellet's Downy-back
Ophonus melletii hyrcanus Kataev, 2001
Ophonus melletii melletii Heer, 1837
Ophonus (Metophonus) puncticeps (Stephens, 1828)⇒ Ophonus puncticollis Paykull, 1798
Ophonus (Metophonus) puncticollis Paykull, 1798
Ophonus (Metophonus) rufibarbis Fabricius, 1792
Ophonus (Metophonus) rupicola Sturm, 1818
Ophonus (Metophonus) scharifi Morvan, 1977
Ophonus (Metophonus) sciakyi Wrase, 1990
Ophonus (Ophonus) ardosiacus Lutshnik, 1922
Ophonus (Ophonus) diffinis (Dejean, 1829)⇒ Ophonus ardosiacus Lutshnik, 1922
Ophonus (Ophonus) sabulicola Panzer, 1796
Ophonus (Ophonus) stictus Stephens, 1828Oolite Downy-back
Ophonus punticollis
Panagaeus (Hologaeus) fasciatus Say, 1823
Panagaeus (Panagaeus) bipustulatus Fabricius, 1775
Pangus scaritides Sturm, 1818
Paradromius (Manodromius) linearis Olivier, 1795
Paradromius linearis linearis Olivier, 1795
Paradromius (Paradromius) longiceps Dejean, 1826
Paranchus albipes Fabricius, 1796White-legged harp ground beetle
Parazuphium (Parazuphium) chevrolatii Laporte, 1833
Parazuphium chevrolatii chevrolatii Laporte de C..., 1833
Parazuphium chevrolatii schelkownikovi Carnet, 1898
Parophonus (Ophonomimus) hirsutulus Dejean, 1829
Parophonus (Ophonomimus) interstitialis Reitter, 1889
Parophonus (Parophonus) laeviceps Ménétriés, 1832
Parophonus (Parophonus) maculicornis Duftschmid, 1812
Parophonus (Parophonus) mendax P.Rossi, 1790
Parophonus (Parophonus) planicollis (Dejean, 1829)⇒ Parophonus laeviceps Ménétriés, 1832
Penthus tenebrioides Waltl, 1838
Perigona (Trechicus) nigriceps Dejean, 1831Tawny Harp Ground Beetle
Philorhizus (Philorhizus) crucifer Lucas, 1846
Philorhizus crucifer crucifer Lucas, 1846
Philorhizus (Philorhizus) melanocephalus Dejean, 1825Frillfin gobies
Philorhizus (Philorhizus) notatus Stephens, 1827
Philorhizus (Philorhizus) sigma P.Rossi, 1790
Philorhizus nonfriedi Reitter, 1898
Pinacodera complanata Dejean, 1826
Pinacodera limbata (Dejean, 1831)⇒ Cymindis limbata Dejean, 1831
Platyderus (Platyderus) caucasicus Kryzhanovskij, 1968
Platyderus (Platyderus) talyschensis Reitter, 1887
Platyderus (Platyderus) umbratus Ménétriés, 1832
Platynus (Batenus) cincticollis Say, 1823
Platynus (Batenus) hypolithos Say, 1823
Platynus elongatulus Fischer von ..., 1829
Platynus erythropus Dejean, 1828
Platytarus faminii Dejean, 1826
Platytarus faminii faminii Dejean, 1826
Poecilus (Ancholeus) crenuliger Chaudoir, 1876
Poecilus crenuliger crenuliger Chaudoir, 1876
Poecilus (Ancholeus) iranicus Morvan, 1974
Poecilus (Ancholeus) laevicollis (Chaudoir, 1842)⇒ Poecilus peregrinus Tschitschérine, 1898
Poecilus (Ancholeus) nitens Chaudoir, 1850
Poecilus nitens nitens Chaudoir, 1850
Poecilus (Ancholeus) peregrinus Tschitschérine, 1898
Poecilus (Ancholeus) pertusus Schaum, 1858
Poecilus (Ancholeus) puncticollis (Dejean, 1828)⇒ 
Poecilus (Ancholeus) subsimilis Tschitschérine, 1893
Poecilus (Derus) advena Quensel, 1806
Poecilus lepidus lepidus Leske, 1785
Poecilus (Poecilus) chalcites Say, 1823
Poecilus (Poecilus) crassipes Fischer von ..., 1829
Poecilus (Poecilus) cupreus (Linnaeus, 1758)⇒ Poecilus lepidus lepidus Leske, 1785
Poecilus cupreus cupreus Linnaeus, 1758
Poecilus cupreus erythropus Dejean, 1828
Poecilus (Poecilus) cursorius (Dejean, 1828)⇒ Poecilus cupreus cupreus Linnaeus, 1758
Poecilus (Poecilus) festivus Chaudoir, 1868
Poecilus (Poecilus) gotschii Chaudoir, 1846
Poecilus stenoderus gressorius
Poecilus (Poecilus) versicolor Sturm, 1824
Poecilus (Sogines) punctulatus Schaller, 1783
Poecilus bogatshevi Kryzhanovskij, 1983
Polistichus connexus Geoffroy, 1785
Polistichus fasciolatus (P.Rossi, 1790)⇒ Polistichus connexus Geoffroy, 1785
Psammodromius pallidicolor Mandl, 1973
Psammodromius zajtzewi Eichler, 1924
Pseudanchomenus aptinoides Tarnier, 1860Ground-beetle
Pseudoophonus (Platus) calceatus Duftschmid, 1812
Pseudoophonus (Pseudoophonus) griseus Panzer, 1797
Pseudoophonus (Pseudoophonus) rufipes De Geer, 1774
Pseudotaphoxenus rufitarsis Fischer von ..., 1823
Pseudotaphoxenus rufitarsis rufitarsis Fischer von ..., 1823
Pseudotaphoxenus thoracicus Gebler, 1843
Pterostichus (Adelosia) macer Marsham, 1802
Pterostichus macer macer Marsham, 1802
Pterostichus (Agastillus) capitatus Chaudoir, 1850
Pterostichus (Agastillus) cratocephalus Tschitschérine, 1897
Pterostichus (Agastillus) cucujinus Reitter, 1892
Pterostichus (Agastillus) eriwanicus Tschitschérine, 1897
Pterostichus (Aphaonus) miroshnikovi Zamotajlov, 1991
Pterostichus (Aphaonus) pseudopercus Reitter, 1889
Pterostichus (Argus) strenuus Panzer, 1797
Pterostichus (Argutor) cursor Dejean, 1828
Pterostichus (Argutor) oblongopunctatus Fabricius, 1787
Pterostichus (Argutor) vernalis Panzer, 1796Spring Blackclock
Pterostichus (Bothriopterus) quadrifoveolatus Letzner, 1852
Pterostichus selmanni selmanni Duftschmid, 1812
Pterostichus (Cryobius) colchicus Chaudoir, 1850
Pterostichus (Cryobius) firmus Lutshnik, 1933
Pterostichus (Cryobius) kadjaranze Jedlička, 1947
Pterostichus (Cryobius) pithyusicus Lutshnik, 1933
Pterostichus (Cryobius) platyderus Chaudoir, 1850
Pterostichus (Cryobius) quadraticollis Chaudoir, 1846
Pterostichus (Cryobius) stomoides Chaudoir, 1868
Pterostichus (Falsargutor) kataevi Kryzhanovskij, 1989
Pterostichus (Falsargutor) ponticus Kirschenhofer, 1987
Pterostichus (Feronidius) fornicatus Kolenati, 1845
Pterostichus (Haplomaseus) andreae Tschitschérine, 1897
Pterostichus (Haplomaseus) arator Faldermann, 1836
Pterostichus arator arator Faldermann, 1836
Pterostichus arator perlidaghensis Reitter, 1902
Pterostichus (Haplomaseus) armenus Faldermann, 1836
Pterostichus armenus armenus Faldermann, 1836
Pterostichus armenus tuberculifer König, 1906
Pterostichus (Haplomaseus) caucasicola Tschitschérine, 1893
Pterostichus (Haplomaseus) chydaeus Tschitschérine, 1897
Pterostichus chydaeus krestovyanus Vysoky, 1984
Pterostichus chydaeus kretovyanus Vysoký, 1984
Pterostichus chydaeus osseticus Poppius, 1908
Pterostichus chydaeus ronchettii Reitter, 1909
Pterostichus (Haplomaseus) cristicaudis Kurnakov, 1962
Pterostichus (Haplomaseus) goriensis Tschitschérine, 1897
Pterostichus goriensis goriensis Tschitschérine, 1897
Pterostichus goriensis telavense
Pterostichus (Haplomaseus) meskheticus Belousov, 1991
Pterostichus (Haplomaseus) novotnyorum Vysoký, 1981
Pterostichus novotnyorum novotnyorum Vysoký, 1981
Pterostichus (Haplomaseus) olegi Belousov, 1991
Pterostichus olegi askhicus Belousov, 1991
Pterostichus olegi olegi Belousov, 1991
Pterostichus (Haplomaseus) porcellus Kurnakov, 1962
Pterostichus (Haplomaseus) rousi Kirschenhofer, 1982
Pterostichus (Haplomaseus) tamsii Dejean, 1831
Pterostichus tamsii tamsii Dejean, 1831
Pterostichus (Haptoderus) latiusculus Chaudoir, 1868
Pterostichus (Hypherpes) adoxus Say, 1823
Pterostichus (Lagarus) vernalis Panzer, 1796
Pterostichus (Lamenius) caudicalis Say, 1823
Pterostichus (Lyrothorax) caspius Ménétriés, 1832
Pterostichus (Melanius) anthracinus (Illiger, 1798)⇒ Pterostichus quadrifoveolatus Letzner, 1852
Pterostichus anthracinus depressiusculus Chaudoir, 1844
Pterostichus (Melanius) gracilis Dejean, 1828
Pterostichus gracilis gracilis Dejean, 1828
Pterostichus melanarius cardioderus Chaudoir, 1846
Pterostichus (Myosodus) aapsorum Belousov, 1991
Pterostichus (Myosodus) aibgensis Starck, 1890
Pterostichus aibgensis adelaidae Reitter, 1899
Pterostichus (Myosodus) avaricus Kryzhanovski..., 1983
Pterostichus (Myosodus) filipjevi Lutshnik, 1928
Pterostichus (Myosodus) lacunosus Chaudoir, 1844
Pterostichus lacunosus intricatus Motschulsky, 1845
Pterostichus lacunosus lacunosus Chaudoir, 1844
Pterostichus lacunosus roubalianus Lutshnik, 1928
Pterostichus (Myosodus) lutshnikianus Bogachev & K..., 1958
Pterostichus lutshnikianus lutshnikianus Bogachev & K..., 1958
Pterostichus (Myosodus) nivicola Ménétriés, 1832
Pterostichus (Myosodus) ordinatus Fischer, 1823
Pterostichus ordinatus ordinatus Fischer von ..., 1823
Pterostichus (Myosodus) rudestriatus Reitter, 1883
Pterostichus (Myosodus) schoenherri Faldermann, 1836
Pterostichus (Myosodus) svanicus Lutshnik, 1922
Pterostichus (Myosodus) swaneticus Reitter, 1883
Pterostichus (Oreoplatysma) araraticus Kirschenhofer, 1987
Pterostichus (Oreoplatysma) belousovi Kryzhanovskij, 1989
Pterostichus (Oreoplatysma) borcka Jedlička, 1963
Pterostichus (Oreoplatysma) casaleianus Kirschenhofer, 1991
Pterostichus (Oreoplatysma) cecchiniae Jakobson, 1907
Pterostichus cecchiniae avadcharicus Kirschenhofer, 1987
Pterostichus (Oreoplatysma) chefsuricus Reitter, 1896
Pterostichus (Oreoplatysma) depressidorsis Reitter, 1896
Pterostichus (Oreoplatysma) faunus Kurnakav, 1962
Pterostichus (Oreoplatysma) fencli Dvorak, 1995
Pterostichus (Oreoplatysma) ghilarovi Kryzhanovskij, 1988
Pterostichus (Oreoplatysma) kirschenblatti Kryzhanovskij, 1988
Pterostichus (Oreoplatysma) koenigi Reitter, 1887
Pterostichus koenigi tscherkessicus Reitter, 1896
Pterostichus (Oreoplatysma) kvirensis Belousov, 1991
Pterostichus (Oreoplatysma) mediocris Dvorak, 1995
Pterostichus (Oreoplatysma) nikodymi Dvorak, 1995
Pterostichus (Oreoplatysma) odvarkai Dvorak, 1995
Pterostichus (Oreoplatysma) pasanauricus Kirschenhofer, 1987
Pterostichus (Oreoplatysma) pulchellus Faldermann, 1836
Pterostichus (Oreoplatysma) rubripalpis Csiki, 1930
Pterostichus (Oreoplatysma) satunini Tschitschérine, 1903
Pterostichus (Oreoplatysma) satyrus Kurnakov, 1962
Pterostichus (Oreoplatysma) schodaicus Kirschenhofer, 1987
Pterostichus (Oreoplatysma) sojaki Kirschenhofer, 1987
Pterostichus (Oreoplatysma) tamarae Wrase & Kirs..., 1991
Pterostichus (Oreoplatysma) zolotarewi Reitter, 1911
Pterostichus (Phonias) apfelbecki Csiki, 1908
Pterostichus (Phonias) patruelis Dejean, 1831
Pterostichus (Platysma) niger (Schaller, 1783)⇒ Pterostichus selmanni selmanni Duftschmid, 1812
Pterostichus niger niger Schaller, 1783
Pterostichus niger planipennis Sahlberg, 1844
Pterostichus (Pseudomaseus) anthracinus Illiger, 1798
Pterostichus anthracinus depressiusculus Chaudoir, 1844
Pterostichus (Pseudomaseus) fuscicornis Reiche & Saulcy, 1855
Pterostichus (Pseudomaseus) gracilis Dejean, 1828
Pterostichus (Pseudomaseus) minor Gyllenhal, 1827
Pterostichus (Unitrichus) alexeji Zamotajlov &..., 1992
Pterostichus (Unitrichus) carradei Gautier Des ..., 1866
Pterostichus crenatus (Duftschmid, 1812)⇒ Pterostichus vernalis Panzer, 1796
Pterostichus mariamae Zamotajlov &..., 2015
Pterostichus neilgaimani Chaladze & K..., 2017
Pterostichus nigritus
Pterostichus ordinates kacheticus
Pterostichus setosicus Davies, 2004
Pterostichus svetlanae Zamotajlov &..., 2015
Selenophorus (Selenophorus) fatuus LeConte, 1863
Selenophorus (Selenophorus) palliatus Fabricius, 1798
Singilis (Singilis) turcicus Jedlička, 1963
Sphelodon phoxopteridis Weed, 1888
Sphodrus leucophthalmus Linnaeus, 1758
Stenocrepis (Stenous) mexicana Chevrolat, 1835
Stenolophus (Agonoderus) comma Fabricius, 1775
Stenolophus (Agonoderus) infuscatus Dejean, 1829
Stenolophus (Agonoderus) lecontei Chaudoir, 1868Leconte'S Seedcorn Beetle
Stenolophus (Egadroma) marginatus Dejean, 1829
Stenolophus (Stenolophus) abdominalis Géné, 1836
Stenolophus abdominalis persicus Mannerheim, 1844
Stenolophus (Stenolophus) discophorus Fischer von ..., 1823
Stenolophus (Stenolophus) fuliginosus Dejean, 1829Checked goby
Stenolophus (Stenolophus) mixtus Herbst, 1784
Stenolophus (Stenolophus) ochropezus Say, 1823
Stenolophus (Stenolophus) persicus Mannerheim, 1844
Stenolophus (Stenolophus) proximus Dejean, 1829
Stenolophus (Stenolophus) skrimshiranus Stephens, 1828
Stenolophus (Stenolophus) steveni Krynicki, 1832
Stenolophus (Stenolophus) teutonus Schrank, 1781
Stomis (Stomis) danielanus Semenov, 1904
Stomis (Stomis) pumicatus Panzer, 1796
Stomis pumicatus pumicatus Panzer, 1796
Stomis (Stomis) tschitscherini Semenov, 1904
Strigia stigma Fabricius, 1801
Syntomus foveatus Geoffroy, 1785
Syntomus fuscomaculatus Motschulsky, 1844
Syntomus mongolicus Motschulsky, 1844
Syntomus motschulskyi (Iablokoff-Kh..., 1978)⇒ Syntomus mongolicus Motschulsky, 1844
Syntomus obscuroguttatus Duftschmid, 1812
Syntomus pallipes Dejean, 1825
Syntomus parallelus Ballion, 1871
Syntomus truncatellus Linnaeus, 1761
Synuchus (Synuchus) vivalis Illiger, 1798
Synuchus vivalis vivalis (Illiger, 1798)⇒ Synuchus vivalis Illiger, 1798
Taphoxenus (Lychnifugus) cellarum Adams, 1817
Taphoxenus cellarum cellarum M.Adams, 1817
Taphoxenus cellarum talyschensis Jedlicka, 1952
Tetragonoderus (Crossonychus) fasciatus Haldeman, 1843
Tetragonoderus (Crossonychus) intersectus Germar, 1824
Thermoscelis insignis Chaudoir, 1846
Thermoscelis insignis insignis Chaudoir, 1846
Thermoscelis insignis kamberskyi Reitter, 1888
Trichocellus placidus Gyllenhal, 1827
Trichotichnus (Trichotichnus) dichrous Dejean, 1829
Trichotichnus (Trichotichnus) laevicollis Duftschmid, 1812
Tschitscherinellus oxygonus Chaudoir, 1850
Tschitscherinellus oxygonus oxygonus Chaudoir, 1850
Xanthomelina zajtzewi Eichler, 1924
Zabrus (Aulacozabrus) distinctus Lucas, 1842
Zabrus (Eutroctes) aciculatus Schaum, 1864
Zabrus (Eutroctes) aurichalceus M.Adams, 1817
Zabrus aurichalceus aurichalceus M. F. Adams, 1817
Zabrus aurichalceus punctipennis
Zabrus (Eutroctes) chalceus Faldermann, 1836
Zabrus (Iberozabrus) ambiguus Rambur, 1838
Zabrus (Pelor) anatolicus Ganglbauer, 1915
Zabrus (Pelor) corpulentus Schaum, 1864
Zabrus corpulentus armeniacus Ganglbauer, 1915
Zabrus corpulentus azerbaidjanicus Freude, 2000
Zabrus corpulentus corpulentus Schaum, 1864
Zabrus (Pelor) foveolatus Schaum, 1864
Zabrus (Pelor) melancholicus Schaum, 1864
Zabrus melancholicus melancholicus Schaum, 1864
Zabrus (Pelor) rotundicollis (Ménétriés, 1836)⇒ Zabrus ambiguus Rambur, 1838
Zabrus (Pelor) seriatus Ganglbauer, 1915
Zabrus (Pelor) spectabilis Hampe, 1852
Zabrus (Pelor) spinipes Fabricius, 1798
Zabrus spinipes rugosus Ménétries, 1832
Zabrus (Pelor) trinii Fischer von ..., 1817
Zabrus trinii anatolicus Ganglbauer, 1915
Zabrus trinii araxidis Reitter, 1889
Zabrus trinii trinii Fischer von ..., 1817
Zabrus (Zabrus) morio Mandrias, 1832
Zabrus morio morio Ménétries, 1832
Zabrus (Zabrus) tenebrioides Goeze, 1777Corn ground beetle
Zuphium (Zuphium) araxidis Iablokoff-Kh..., 1972
Zuphium (Zuphium) olens P.Rossi, 1790
Zuphium ciliatum Vauloger De ..., 1897
Zuphium cilicium Peyron, 1858

Subnodes of Harpalinae with occurrences in Caucasus
SupertribeAnthiitae
Bonelli, 1813
1
1		Genus
Species
Taxom

SupertribeChlaeniitae
Bonelli, 1810
11
63		Genera
Species
Taxom

SupertribeCtenodactylitae
Castelnau, 1834
1
1		Genus
Species
Taxom

SupertribeDryptitae
Bonelli, 1810
5
10		Genera
Species
Taxom

SupertribeHarpalitae
Erwin, 1985
34
192		Genera
Species
Taxom

SupertribeLebiitae
Bonelli, 1810
34
101		Genera
Species
Taxom

SupertribePlatynitae
Bonelli, 1810
18
95		Genera
Species
Taxom

SupertribePterostichitae
Bonelli, 1810
13
216		Genera
Species
Taxom

TribeGlyptini
Cushman & Rohwer, 1920
1
1		Genus
Species
Taxom

TribeIdiomorphini
Bates, 1891
1
1		Genus
Species
Taxom

TribePseudomorphini
Newman, 1842
0
0		Genera
Species
Taxom



Links, sources and further information
Harpalinae at artfakta.se
Harpalinae at dyntaxa.se
Harpalinae at itis.gov