Checklist for Subfamily Harpalinae Bonelli, 1810
and region Pennsylvania (USA)
Class
| Insecta
Linnaeus, 1758
Insects, True insects | 554
5452
12530 | Families
Genera
Species |
Taxom
|
Subclass
| Pterygota
Lang, 1888
Winged insects | 553
5449
12526 | Families
Genera
Species |
Taxom
|
Infraclass
| Neoptera
van der Wulp, 1890
Modern Wing-folding insects | 525
5347
12273 | Families
Genera
Species |
Taxom
|
Superorder
| Holometabola
Heider, 1889
Holometabolous Insects | 355
4526
10429 | Families
Genera
Species |
Taxom
|
Order
| Coleoptera
Linnaeus, 1758
Beetles, Coleopterans | 102
1569
3485 | Families
Genera
Species |
Taxom
|
Suborder
| Adephaga
Schellenberg, 1806
Adephagans, Ground Beetles and Water Beetles | 5
178
688 | Families
Genera
Species |
Taxom
|
Superfamily
| Caraboidea
Latreille, 1802
| 1
143
597 | Family
Genera
Species |
Taxom
| | Family | Carabidae
Latreille, 1802
Carabid beetles, Carabids, Ground Beetles, Tiger Beetles | 143
597 | Genera
Species |
Taxom
| | Subfamily | Harpalinae
Bonelli, 1810
| 88
366 | Genera
Species |
Taxom
|
Genera for Pennsylvania
88 Genera with 366 Species #ABCDEGHILMNOPRSTXZ
| Abacidus LeConte, 1852 | | 4 | Species | Taxom | | Acupalpus Latreille, 1829 | | 8 | Species | Taxom | | Agonoleptus Casey, 1843 | | 2 | Species | Taxom | | Agonum Bonelli, 1954 | | 37 | Species | Taxom | | Agra Fabricius, 1835 | Elegant canopy beetles | 1 | Species | Taxom | | Amara Bonelli, 1837 | Seed-Eating Ground Beetles | 24 | Species | Taxom | | Amblygnathus Dejean, 1823 | | 1 | Species | Taxom | Amblystomus Erichson, 1837 | | 0 | Species | Taxom | | Amerinus Casey, 1863 | | 1 | Species | Taxom | | Amphasia Newman, 1828 | | 2 | Species | Taxom | | Anisodactylus Dejean, 1848 | | 17 | Species | Taxom | | Apenes LeConte, 1823 | | 3 | Species | Taxom | | Apristus Chaudoir, 1826 | | 1 | Species | Taxom | | Atranus LeConte, 1828 | | 1 | Species | Taxom | | Badister Clairville, 1920 | | 8 | Species | Taxom | | Bradycellus Erichson, 1848 | | 13 | Species | Taxom | | Calathus Bonelli, 1854 | | 2 | Species | Taxom | | Calleida Latreille & ..., 1845 | | 5 | Species | Taxom | | Chlaenius Bonelli, 1876 | Vivid Metallic Ground Beetle | 23 | Species | Taxom | | Colliuris De Geer, 1758 | Long-necked Ground Beetles | 1 | Species | Taxom | | Colpodes W.S.Macleay, 1947 | | 1 | Species | Taxom | | Coptodera Dejean, 1825 | | 1 | Species | Taxom | | Cosnania Dejean, 1767 | | 1 | Species | Taxom | | Cratacanthus Dejean, 1811 | | 1 | Species | Taxom | | Cratoferonia Tschitschérine, 1900 | | 1 | Species | Taxom | | Cyclotrachelus Chaudoir, 1848 | | 11 | Species | Taxom | | Cylindronotum Putzeys, 1845 | | 1 | Species | Taxom | | Cymindis Latreille, 1876 | | 8 | Species | Taxom | | Dicaelus Bonelli, 1876 | Notched-mouth Ground Beetles | 11 | Species | Taxom | | Diplocheila Brullé, 1848 | | 2 | Species | Taxom | | Dromius Bonelli, 1831 | | 1 | Species | Taxom | | Euproctinus Leng & Mutchler, 1883 | | 1 | Species | Taxom | | Galerita Fabricius, 1792 | False Bombardier Beetle | 3 | Species | Taxom | | Gastrellarius Casey, 1823 | | 1 | Species | Taxom | | Geopinus LeConte, 1829 | | 1 | Species | Taxom | | Glypta Gravenhorst, 1919 | | 1 | Species | Taxom | | Harpalus Latreille, 1865 | | 22 | Species | Taxom | | Helluomorphoides Ball, 1825 | Flat-horned Ground Beetles | 2 | Species | Taxom | | Holcaspis de Chaudoir, 1908 | | 1 | Species | Taxom | | Incagonum Liebherr, 1868 | | 1 | Species | Taxom | | Lachnocrepis LeConte, 1830 | | 1 | Species | Taxom | | Lebia Latreille, 1896 | colorful foliage ground beetle | 22 | Species | Taxom | | Leptotrachelus Latreille, 1801 | | 1 | Species | Taxom | | Lophoglossus LeConte, 1823 | | 2 | Species | Taxom | | Loxandrus LeConte, 1918 | | 4 | Species | Taxom | | Microlestes Schmidt-Goebel, 1863 | | 1 | Species | Taxom | | Myas Sturm, 1828 | | 3 | Species | Taxom | | Nemotarsus LeConte, 1853 | | 1 | Species | Taxom | | Neocolpodes Jeannel, 1935 | | 1 | Species | Taxom | | Notiobia Perty, 1823 | | 3 | Species | Taxom | | Notonomus Chaudoir, 1916 | | 3 | Species | Taxom | | Olisthopus Dejean, 1823 | | 1 | Species | Taxom | | Oodes Bonelli, 1863 | | 3 | Species | Taxom | | Ophonus Dejean, 1798 | | 2 | Species | Taxom | Oxygonium Basilewsky, 1951 | | 0 | Species | Taxom | | Oxypselaphus Chaudoir, 1854 | | 1 | Species | Taxom | | Panagaeus Latreille, 1823 | | 1 | Species | Taxom | | Penthus Chaudoir, 1838 | | 1 | Species | Taxom | | Perigona Laporte, 1853 | | 2 | Species | Taxom | | Philodes LeConte, 1853 | | 2 | Species | Taxom | | Pinacodera Schaum, 1823 | | 2 | Species | Taxom | | Piosoma LeConte, 1848 | | 2 | Species | Taxom | | Platynus Bonelli, 1854 | | 10 | Species | Taxom | | Plochionus Dejean, 1775 | | 2 | Species | Taxom | | Poecilus Bonelli, 1823 | | 2 | Species | Taxom | | Pogonodaptus Horn, 1878 | | 1 | Species | Taxom | | Prosopogmus Chaudoir, 1843 | | 1 | Species | Taxom | | Pseudamara Lindroth, 1848 | | 1 | Species | Taxom | Pseudamphasia Casey, 1914 | | 0 | Species | Taxom | | Pseudaptinus Castelnau de..., 1872 | | 1 | Species | Taxom | | Pseudomorpha Kirby, 1925 | Western False-form Beetles | 1 | Species | Taxom | | Pterostichus Bonelli, 1890 | Woodland Ground Beetle | 34 | Species | Taxom | | Rhadine LeConte, 1863 | | 1 | Species | Taxom | | Selenophorus Dejean, 1863 | | 4 | Species | Taxom | | Siopelus Murray, 1926 | | 1 | Species | Taxom | | Sphelodon Townes, 1888 | | 1 | Species | Taxom | | Stenocrepis Chaudoir, 1843 | | 1 | Species | Taxom | | Stenolophus Dejean, 1829 | Seedcorn Beetle | 11 | Species | Taxom | | Stenomorphus Dejean, 1843 | | 1 | Species | Taxom | | Stereocerus Kirby, 1831 | | 1 | Species | Taxom | | Syntomus Hope, 1831 | | 1 | Species | Taxom | | Synuchus Gyllenhal, 1823 | | 1 | Species | Taxom | | Tetragonoderus Dejean, 1843 | | 1 | Species | Taxom | | Tetraleucus Casey, 1844 | | 1 | Species | Taxom | | Trichosternus Chaudoir, 1833 | | 1 | Species | Taxom | | Trichotichnus A. Morawitz, 1823 | | 3 | Species | Taxom | | Xestonotus LeConte, 1829 | | 1 | Species | Taxom | | Zuphioides Ball & Shpeley, 1831 | | 1 | Species | Taxom |
Species list for Pennsylvania
366 Species and 5 Subspecies | Abacidus atratus (Newman, 1838) | ⇒ Pterostichus atratus Gistel, 1857 | | Abacidus hamiltoni (G.Horn, 1880) | ⇒ Pterostichus hamiltoni G. Horn, 1880 | | Abacidus permundus (Say, 1830) | ⇒ Pterostichus permundus Say, 1830 | | Abacidus sculptus (LeConte, 1852) | ⇒ Pterostichus sculptus LeConte, 1853 | | Acupalpus (Acupalpus) carus LeConte, 1863 | | | Acupalpus (Acupalpus) hydropicus LeConte, 1863 | | | Acupalpus (Acupalpus) pumilus Lindroth, 1968 | Leopard-spotted goby | | Acupalpus (Philodes) rectangulus (Chaudoir, 1868) | ⇒ Philodes rectangulus Chaudoir, 1868 | | Acupalpus (Tachistodes) indistinctus Dejean, 1831 | | | Acupalpus (Tachistodes) partiarius Say, 1823 | | | Acupalpus (Tachistodes) pauperculus Dejean, 1829 | Checked goby | | Acupalpus (Tachistodes) testaceus Dejean, 1829 | Checked goby | | Agonoleptus conjunctus Say, 1823 | | | Agonoleptus rotundicollis Haldeman, 1843 | | | Agonum (Agonum) muelleri Herbst, 1784 | Müller'S Harp Ground Beetle | | Agonum (Agonum) placidum Say, 1823 | | | Agonum (Europhilus) anchomenoides Randall, 1838 | | | Agonum (Europhilus) canadense Goulet, 1969 | Dogtoothed goby | | Agonum (Europhilus) consimile (Gyllenhal, 1810) | ⇒ Agonum scitulum Dejean, 1828 | | Agonum (Europhilus) galvestonicum Casey, 1920 | | | Agonum (Europhilus) gratiosum Mannerheim, 1853 | | | Agonum (Europhilus) lutulentum LeConte, 1854 | | | Agonum (Europhilus) palustre Goulet, 1969 | Dogtoothed goby | | Agonum (Europhilus) retractum LeConte, 1848 | Goldspot goby | | Agonum (Europhilus) simile (Kirby, 1837) | ⇒ | | Agonum (Europhilus) thoreyi Dejean, 1828 | | | Agonum (Olisares) aeruginosum (Dejean, 1828) | ⇒ Agonum muelleri Herbst, 1784 | | Agonum (Olisares) affine (Kirby, 1837) | ⇒ Agonum thoreyi Dejean, 1828 | | Agonum (Olisares) basale (LeConte, 1848) | ⇒ | | Agonum (Olisares) collare (Say, 1830) | ⇒ | | Agonum (Olisares) cupripenne Say, 1823 | | | Agonum (Olisares) decorum Say, 1823 | | | Agonum (Olisares) excavatum Dejean, 1828 | | | Agonum (Olisares) extensicolle Say, 1823 | | | Agonum (Olisares) ferreum Haldeman, 1843 | | | Agonum (Olisares) fidele Casey, 1920 | | | Agonum (Olisares) harrisii LeConte, 1848 | Goldspot goby | | Agonum (Olisares) melanarium Dejean, 1828 | | | Agonum (Olisares) metallescens (LeConte, 1854) | ⇒ | | Agonum (Olisares) moerens Dejean, 1828 | | | Agonum (Olisares) mutatum Gemminger & ..., 1868 | | | Agonum (Olisares) nutans Say, 1823 | | | Agonum (Olisares) octopunctatum Fabricius, 1798 | Saline Valley snow-front June beetle | | Agonum (Olisares) pallipes Fabricius, 1787 | Spotted Warner Valley Dunes June beetle | | Agonum (Olisares) punctiforme Say, 1823 | | | Agonum (Olisares) quinquepunctatum (Motschulsky, 1844) | ⇒ Agonum muelleri Herbst, 1784 | | Agonum (Olisares) rufipes (Dejean, 1828) | ⇒ Agonum fuliginosum Panzer, 1809 | | Agonum (Olisares) striatopunctatum Dejean, 1828 | | | Agonum (Olisares) tenue LeConte, 1854 | | | Agonum (Olisares) texanum LeConte, 1878 | | | Agonum (Olisares) trigeminum Lindroth, 1954 | Painted goby | | Agra oblongopunctata Chevrolat, 1835 | | | Amara (Amara) aenea De Geer, 1774 | Common sun beetle | | Amara (Amara) bamidunyae Bates, 1878 | | | Amara (Amara) basillaris Say, 1823 | | | Amara (Amara) confusa LeConte, 1848 | | | Amara (Amara) convexa LeConte, 1848 | | | Amara (Amara) cupreolata Putzeys, 1866 | Bumblebee gobies | | Amara (Amara) familiaris Duftschmid, 1812 | Bareyed goby | | Amara (Amara) impuncticollis Say, 1823 | | | Amara (Amara) littoralis Mannerheim, 1843 | | | Amara (Amara) lunicollis Schiodte, 1837 | Golden-banded goby | | Amara (Amarocelia) laevipennis Kirby, 1837 | | | Amara (Amarocelia) patruelis Dejean, 1831 | | | Amara (Bradytus) apricaria Paykull, 1790 | Exposed sun beetle | | Amara (Bradytus) avida Say, 1823 | | | Amara (Bradytus) exarata (Dejean, 1828) | ⇒ Amara aenea De Geer, 1774 | | Amara (Bradytus) latior Kirby, 1837 | | | Amara (Celia) interstitialis Dejean, 1828 | | | Amara (Celia) rubrica Haldeman, 1843 | | | Amara (Curtonotus) pennsylvanica Hayward, 1908 | | | Amara (Paracelia) quenseli Schoenher, 1806 | Bearded goby | | Amara (Percosia) obesa (Say, 1823) | ⇒ Amara ignatovitschi Tschitschérine, 1894 | | Amara (Zezea) angustata (Say, 1823) | ⇒ Amara alpina Paykull, 1790 | | Amara (Zezea) angustatoides Hieke, 2000 | Bumblebee fish | | Amara (Zezea) pallipes Kirby, 1837 | | | Amblygnathus iripennis Say, 1823 | | | Amerinus linearis LeConte, 1863 | | | Amphasia (Amphasia) interstitialis Say, 1823 | Red-Headed Ground Beetle | | Amphasia (Pseudamphasia) sericea T.W.Harris, 1828 | Freshwater goby | | Anisodactylus (Anadaptus) discoideus Dejean, 1831 | | | Anisodactylus (Anadaptus) sanctaecrucis Fabricius, 1798 | | | Anisodactylus (Anisodactylus) agricola Say, 1823 | | | Anisodactylus (Anisodactylus) carbonarius Say, 1823 | | | Anisodactylus (Anisodactylus) harrisii LeConte, 1863 | | | Anisodactylus (Anisodactylus) kirbyi Lindroth, 1953 | Durban goby | | Anisodactylus (Anisodactylus) melanopus Haldeman, 1843 | | | Anisodactylus (Anisodactylus) nigerrimus Dejean, 1831 | | | Anisodactylus (Anisodactylus) nigrita Dejean, 1829 | Checked goby | | Anisodactylus (Anisodactylus) pseudagricola Noonan, 1996 | Meander goby | | Anisodactylus (Aplocentrus) caenus Say, 1823 | | | Anisodactylus (Gynandrotarsus) merula Germar, 1824 | | | Anisodactylus (Gynandrotarsus) opaculus LeConte, 1863 | | | Anisodactylus (Gynandrotarsus) ovularis Casey, 1914 | | | Anisodactylus (Gynandrotarsus) rusticus Say, 1823 | | | Anisodactylus (Pseudaplocentrus) laetus Dejean, 1829 | Checked goby | | Anisodactylus (Spongopus) verticalis LeConte, 1848 | | | Apenes (Apenes) sinuatus Say, 1823 | | | Apenes lucidula Dejean, 1831 | | | Apenes sinuata Say, 1823 | | | Apristus subsulcatus Dejean, 1826 | | | Atranus pubescens Dejean, 1828 | Bynoe's goby | | Badister (Badister) elegans LeConte, 1880 | | | Badister (Badister) maculatus LeConte, 1853 | | | Badister (Badister) neopulchellus Lindroth, 1954 | Painted goby | | Badister (Badister) notatus Haldeman, 1843 | | | Badister (Baudia) grandiceps Casey, 1920 | | | Badister (Baudia) micans LeConte, 1844 | | | Badister (Baudia) reflexus LeConte, 1880 | | | Badister (Baudia) transversus Casey, 1920 | | | Bradycellus (Catharellus) lecontei Csiki, 1932 | | | Bradycellus (Lipalocellus) nigrinus Dejean, 1829 | | | Bradycellus (Stenocellus) congener LeConte, 1848 | Limestone Cave beetle | | Bradycellus (Stenocellus) festinans Casey, 1914 | Deceptive cave beetle | | Bradycellus (Stenocellus) neglectus LeConte, 1848 | | | Bradycellus (Stenocellus) nigriceps LeConte, 1868 | | | Bradycellus (Stenocellus) rupestris Say, 1823 | Rich Mountain cave beetle | | Bradycellus (Stenocellus) suavis Casey, 1914 | | | Bradycellus (Stenocellus) tantillus Dejean, 1829 | | | Bradycellus (Triliarthrus) atrimedeus Say, 1823 | | | Bradycellus (Triliarthrus) badipennis Haldeman, 1843 | | | Bradycellus (Triliarthrus) kirbyi G. Horn, 1883 | Goggle goby | | Bradycellus (Triliarthrus) lugubris LeConte, 1848 | | | Calathus (Neocalathus) gregarius Say, 1823 | | | Calathus (Neocalathus) opaculus LeConte, 1854 | | | Calleida (Calleida) circumcincta Bates, 1883 | | | Calleida (Calleida) planulata LeConte, 1858 | Kalia | | Calleida (Calleida) punctata LeConte, 1848 | Goldspot goby | | Calleida (Calleida) viridipennis Say, 1823 | | | Calleida cordicollis (Putzeys, 1845) | ⇒ Calleida marginicollis Chaudoir, 1844 | | Chlaenius (Agostenus) niger (Randall, 1838) | ⇒ Chlaenius nigerrimus JedliÄka, 1958 | | Chlaenius (Anomoglossus) emarginatus Say, 1823 | Emarginate Chlaenius | | Chlaenius (Anomoglossus) pusillus Say, 1823 | | | Chlaenius (Brachylobus) lithophilus Say, 1823 | | | Chlaenius (Chlaeniellus) brevilabris Le Conte, 1848 | | | Chlaenius (Chlaeniellus) flaccidus G. Horn, 1876 | | | Chlaenius (Chlaeniellus) impunctifrons (Say, 1823) | ⇒ Chlaenius pennsylvanicus pennsylvanicus Say, 1823 | | Chlaenius (Chlaeniellus) nemoralis Say, 1823 | | | Chlaenius (Chlaeniellus) pennsylvanicus Say, 1823 | Say´s Ground Beetle | | Chlaenius pennsylvanicus pennsylvanicus Say, 1823 | | | Chlaenius (Chlaeniellus) tricolor Dejean, 1826 | Prickly scorpion's-tail | | Chlaenius (Chlaeniellus) vafer LeConte, 1852 | | | Chlaenius (Chlaenius) aestivus Say, 1823 | | | Chlaenius (Chlaenius) laticollis (Say, 1823) | ⇒ | | Chlaenius (Chlaenius) orbus G. Horn, 1871 | | | Chlaenius (Chlaenius) platyderus Chaudoir, 1856 | Mudskipper | | Chlaenius (Chlaenius) sericeus Forster, 1771 | Green Pubescent Ground Beetle | | Chlaenius sericeus sericeus Forster, 1771 | | | Chlaenius (Eurydactylus) tomentosus Say, 1823 | | | Chlaenius (Lithochlaenius) cordicollis (Kirby, 1837) | ⇒ Chlaenius kindermanni chrysothorax Krynitzky, 1832 | | Chlaenius (Lithochlaenius) leucoscelis Chevrolat, 1835 | River goby | | Chlaenius (Lithochlaenius) prasinus Dejean, 1826 | Sponge goby | | Chlaenius punctulatus ♰ G. H. Horn, 1876 | | | Colliuris (Cosnania) pensylvanica (Linnaeus, 1758) | ⇒ Cosnania pensylvanica Linnaeus, 1767 | | Colpodes phengodes Andrewes, 1947 | | | Coptodera (Coptodera) aerata Dejean, 1825 | | | Cosnania pensylvanica Linnaeus, 1767 | | | Cratacanthus dubius Palisot de B..., 1811 | Goatee blenny | | Cratoferonia phylarchus Sloane, 1900 | | | Cyclotrachelus (Cyclotrachelus) approximatus LeConte, 1848 | | | Cyclotrachelus (Cyclotrachelus) faber Germar, 1824 | | | Cyclotrachelus (Cyclotrachelus) fucatus Freitag, 1969 | | | Cyclotrachelus (Cyclotrachelus) vinctus LeConte, 1852 | | | Cyclotrachelus (Evarthrus) convivus LeConte, 1852 | | | Cyclotrachelus (Evarthrus) engelmani LeConte, 1852 | | | Cyclotrachelus (Evarthrus) furtivus LeConte, 1852 | | | Cyclotrachelus (Evarthrus) incisus LeConte, 1848 | | | Cyclotrachelus (Evarthrus) seximpressus LeConte, 1848 | | | Cyclotrachelus (Evarthrus) sigillatus Say, 1823 | | | Cyclotrachelus sodalis sodalis LeConte, 1848 | | | Cylindronotum aeneum Putzeys, 1845 | Locusthead | | Cymindis (Cymindis) americanus Dejean, 1826 | | | Cymindis (Cymindis) neglectus Haldeman, 1843 | | | Cymindis (Pinacodera) limbatus Dejean, 1831 | | | Cymindis (Tarulus) americana Dejean, 1826 | Sponge goby | | Cymindis (Tarulus) cribricollis Dejean, 1831 | | | Cymindis (Tarulus) neglecta Haldeman, 1843 | | | Cymindis (Tarulus) pilosa (Say, 1823) | ⇒ Cymindis faldermanni Gistel, 1838 | | Cymindis aurora ♰ G. H. Horn, 1876 | | | Dicaelus (Dicaelus) costatus LeConte, 1848 | | | Dicaelus (Dicaelus) crenatus LeConte, 1853 | | | Dicaelus (Dicaelus) purpuratus Bonelli, 1813 | | | Dicaelus (Paradicaelus) ambiguus LaFerté-Séne..., 1841 | | | Dicaelus (Paradicaelus) dilatatus Say, 1823 | | | Dicaelus (Paradicaelus) elongatus Bonelli, 1813 | | | Dicaelus (Paradicaelus) furvus Dejean, 1826 | Sponge goby | | Dicaelus (Paradicaelus) politus Dejean, 1826 | Sponge goby | | Dicaelus (Paradicaelus) sculptilis Say, 1823 | | | Dicaelus (Paradicaelus) teter Bonelli, 1813 | | | Dicaelus alutaceus ♰ G. H. Horn, 1876 | | | Diplocheila (Isorembus) impressicollis Dejean, 1831 | | | Diplocheila (Isorembus) obtusa LeConte, 1848 | | | Dromius (Dromius) piceus Dejean, 1831 | | | Euproctinus (Neoeuproctus) abjectus Bates, 1883 | | | Galerita (Galerita) esmeraldina Reichardt, 1967 | | | Galerita (Progaleritina) bicolor Drury, 1773 | False bombardier beetle | | Galerita (Progaleritina) janus Fabricius, 1792 | | | Gastrellarius honestus Say, 1823 | | | Geopinus incrassatus Dejean, 1829 | Homely Geopinus | | Glypta mutica Cushman, 1919 | | | Harpalus (Glanodes) cohni Ball, 1972 | | | Harpalus (Glanodes) puncticeps Casey, 1914 | Ground beetle | | Harpalus (Harpalobius) autumnalis Duftschmid, 1812 | | | Harpalus (Harpalus) affinis (Schrank, 1781) | ⇒ Harpalus serripes ernsti Kataev, 1995 | | Harpalus (Harpalus) fulgens Csiki, 1932 | | | Harpalus (Harpalus) herbivagus Say, 1823 | | | Harpalus (Harpalus) meteorus Basilewsky, 1946 | | | Harpalus (Harpalus) somnulentus Dejean, 1829 | Checked goby | | Harpalus (Megapangus) caliginosus Fabricius, 1798 | Dusky ground beetle | | Harpalus (Megapangus) katiae F.Battoni, 1985 | | | Harpalus (Opadius) desertus LeConte, 1859 | Bullwood | | Harpalus (Opadius) indigens Casey, 1924 | Goose barnacles | | Harpalus (Opadius) providens Casey, 1914 | | | Harpalus (Opadius) reversus Casey, 1924 | Goose barnacles | | Harpalus (Opadius) spadiceus Dejean, 1829 | Checked goby | | Harpalus (Plectralidus) erraticus (Say, 1823) | ⇒ Harpalus eraticus Say, 1823 | | Harpalus (Pseudoophonus) compar (LeConte, 1848) | ⇒ Harpalus cohni Ball, 1972 | | Harpalus (Pseudoophonus) erythropus Dejean, 1829 | Checked goby | | Harpalus (Pseudoophonus) faunus Say, 1823 | | | Harpalus (Pseudoophonus) pensylvanicus DeGeer, 1774 | Pennsylvania Dingy Ground Beetle | | Harpalus (Pseudoophonus) rufipes (DeGeer, 1774) | ⇒ | | Harpalus (Pseudoophonus) vagans LeConte, 1865 | | | Helluomorphoides ferrugineus LeConte, 1853 | | | Helluomorphoides praeustus Dejean, 1825 | Bluespot goby | | Holcaspis odontella Broun, 1908 | | | Incagonum fuscoaeneum Gemminger & ..., 1868 | | | Lachnocrepis parallela Say, 1830 | | | Lebia (Lebia) abdominalis Chaudoir, 1843 | | | Lebia (Lebia) analis Dejean, 1825 | | | Lebia (Lebia) bitaeniata Chevrolat, 1834 | | | Lebia (Lebia) bivittata Fabricius, 1798 | | | Lebia (Lebia) calliope Bates, 1883 | | | Lebia (Lebia) fuscata (Dejean, 1825) | ⇒ Lebia fraterna Péringuey, 1896 | | Lebia (Lebia) marginicollis Dejean, 1825 | | | Lebia (Lebia) ornata Say, 1823 | | | Lebia (Lebia) pleuritica LeConte, 1848 | Goldspot goby | | Lebia (Lebia) pulchella Dejean, 1826 | Beautiful Banded Lebia | | Lebia (Lebia) pumila Dejean, 1831 | | | Lebia (Lebia) rhodope Bates, 1883 | | | Lebia (Lebia) rufopleura Schaeffer, 1910 | | | Lebia (Lebia) scapularis (Geoffroy, 1785) | ⇒ Lebia solea Hentz, 1830 | | Lebia (Lebia) solea Hentz, 1830 | | | Lebia (Lebia) viridipennis Dejean, 1826 | Green-winged Lebia | | Lebia (Lebia) viridis Say, 1823 | Flower Lebia Beetle | | Lebia (Lebia) vittata (Fabricius, 1777) | ⇒ Lebia cruxminor cruxminor Linnaeus, 1758 | | Lebia (Loxopeza) atriventris Say, 1823 | | | Lebia (Loxopeza) grandis Hentz, 1830 | Large Foliage Ground Beetle | | Lebia (Loxopeza) tricolor Say, 1823 | | | Lebia (Nematopeza) fraterna Péringuey, 1896 | | | Leptotrachelus dorsalis Fabricius, 1801 | | | Lophoglossus substrenuus Csiki, 1930 | | | Lophoglossus tartaricus Say, 1823 | | | Loxandrus agilis Dejean, 1828 | Bynoe's goby | | Loxandrus brevicollis LeConte, 1848 | | | Loxandrus pravitubus Allen, 1972 | | | Loxandrus vulneratus Casey, 1918 | | | Microlestes pusio LeConte, 1863 | | | Myas (Neomyas) cyanescens Dejean, 1828 | | | Myas (Trigonognatha) coracinus Say, 1823 | | | Myas (Trigonognatha) cyanescens Dejean, 1828 | | | Nemotarsus elegans LeConte, 1853 | | | Neocolpodes hylobius Alluaud, 1935 | | | Notiobia (Anisotarsus) nitidipennis LeConte, 1848 | | | Notiobia (Anisotarsus) sayi Blatchley, 1910 | | | Notiobia (Anisotarsus) terminata Say, 1823 | | | Notonomus (Acanthoferonia) angustibasis Sloane, 1902 | | | Notonomus (Leiradira) amabilis Castelnau, 1867 | | | Notonomus (Leiradira) truncatus Sloane, 1916 | | | Olisthopus parmatus Say, 1823 | | | Oodes amaroides Dejean, 1831 | | | Oodes americanus Dejean, 1826 | Sponge goby | | Oodes fluvialis (LeConte, 1863) | ⇒ Oodes americanus Dejean, 1826 | | Ophonus (Metophonus) puncticeps (Stephens, 1828) | ⇒ Ophonus puncticollis Paykull, 1798 | | Ophonus (Metophonus) puncticollis Paykull, 1798 | | | Oxypselaphus pusillus LeConte, 1854 | | | Panagaeus (Hologaeus) fasciatus Say, 1823 | | | Penthus tenebrioides Waltl, 1838 | | | Perigona (Trechicus) nigriceps Dejean, 1831 | Tawny Harp Ground Beetle | | Perigona (Trechicus) pallipennis LeConte, 1853 | | | Philodes (Goniolophus) rectangulus Chaudoir, 1868 | Quagga goby | | Philodes (Philodes) alternans LeConte, 1853 | | | Pinacodera limbata (Dejean, 1831) | ⇒ Cymindis limbata Dejean, 1831 | | Pinacodera platicollis Say, 1823 | | | Piosoma brevipennis | | | Piosoma setosum LeConte, 1848 | | | Platynus (Batenus) angustatus Dejean, 1828 | | | Platynus (Batenus) cincticollis Say, 1823 | | | Platynus (Batenus) hypolithos Say, 1823 | | | Platynus (Platynus) daviesi Bousquet, 2012 | Bigmouth sleepers | | Platynus (Platynus) decentis Say, 1823 | | | Platynus (Platynus) indecentis Liebherr & Will, 1996 | Manson's goby | | Platynus (Platynus) opaculus LeConte, 1863 | | | Platynus (Platynus) parmarginatus Hamilton, 1893 | | | Platynus (Platynus) tenuicollis LeConte, 1848 | | | Platynus laevis LeConte, 1854 | | | Plochionus (Menidius) timidus Haldeman, 1843 | Timid harp ground beetle | | Plochionus (Plochionus) pallens Fabricius, 1775 | Ground beetle | | Poecilus (Poecilus) chalcites Say, 1823 | | | Poecilus (Poecilus) lucublandus Say, 1823 | Woodland Ground Beetle | | Pogonodaptus mexicanus H.W.Bates, 1878 | | | Prosopogmus chalybeipennis Chaudoir, 1843 | | | Pseudamara arenaria LeConte, 1848 | | | Pseudaptinus (Thalpius) horni Chaudoir, 1872 | Railway glider | | Pseudomorpha (Pseudomorpha) tenebroides Notman, 1925 | | | Pterostichus (Abacidus) hamiltoni G. Horn, 1880 | Candystick goby | | Pterostichus (Argutor) commutabilis Motschulsky, 1866 | | | Pterostichus (Bothriopterus) lustrans LeConte, 1851 | | | Pterostichus (Bothriopterus) mutus Say, 1823 | | | Pterostichus (Bothriopterus) pensylvanicus LeConte, 1873 | | | Pterostichus (Bothriopterus) trinarius Casey, 1918 | | | Pterostichus amoenus amoenus Dejean, 1828 | | | Pterostichus (Cylindrocharis) rostratus Newman, 1838 | | | Pterostichus (Eosteropus) moestus Say, 1823 | | | Pterostichus (Euferonia) coracinus (Newman, 1838) | ⇒ Pterostichus multipunctatus Dejean, 1828 | | Pterostichus (Euferonia) lachrymosus Newman, 1838 | | | Pterostichus (Euferonia) novus Straneo, 1944 | | | Pterostichus (Euferonia) relictus (Newman, 1838) | ⇒ Pterostichus amoenus amoenus Dejean, 1828 | | Pterostichus (Euferonia) stygicus Say, 1823 | | | Pterostichus (Gastrosticta) sayanus Csiki, 1930 | Key goby | | Pterostichus (Hypherpes) adoxus Say, 1823 | | | Pterostichus (Hypherpes) californicus Dejean, 1828 | Bynoe's goby | | Pterostichus (Hypherpes) tristis Dejean, 1828 | Bynoe's goby | | Pterostichus (Lamenius) caudicalis Say, 1823 | | | Pterostichus (Lenapterus) punctatissimus Randall, 1838 | | | Pterostichus (Melanius) corvinus Dejean, 1828 | Bynoe's goby | | Pterostichus (Melanius) ebeninus Dejean, 1828 | Bynoe's goby | | Pterostichus (Monoferonia) diligendus Chaudoir, 1868 | Quagga goby | | Pterostichus (Monoferonia) mancus LeConte, 1852 | | | Pterostichus (Morphnosoma) melanarius Illiger, 1798 | European Dark Harp Ground Beetle | | Pterostichus (Oreophilus) multipunctatus Dejean, 1828 | | | Pterostichus (Phonias) femoralis Kirby, 1837 | | | Pterostichus (Phonias) patruelis Dejean, 1831 | | | Pterostichus (Pseudomaseus) luctuosus Dejean, 1828 | Bynoe's goby | | Pterostichus (Pterostichus) funestes Csiki, 1930 | | | Pterostichus funestes funestes Csiki, 1930 | | | Pterostichus (Unitrichus) moestus (Say, 1823) | ⇒ Pterostichus funestes funestes Csiki, 1930 | | Pterostichus (Unitrichus) superciliosus Say, 1823 | | | Pterostichus laevigatus ♰ (S. H. Scudder, 1890) | ⇒ Pterostichus minor Gyllenhal, 1827 | | Rhadine caudata LeConte, 1863 | | | Selenophorus (Selenophorus) fatuus LeConte, 1863 | | | Selenophorus (Selenophorus) gagatinus Dejean, 1829 | Checked goby | | Selenophorus (Selenophorus) hylacis Say, 1823 | | | Selenophorus (Selenophorus) opalinus LeConte, 1863 | | | Siopelus (Neosiopelus) pediobius Alluaud, 1926 | | | Sphelodon phoxopteridis Weed, 1888 | | | Stenocrepis (Stenous) cuprea Chaudoir, 1843 | Horny goby | | Stenolophus (Agonoderus) comma Fabricius, 1775 | | | Stenolophus (Agonoderus) lecontei Chaudoir, 1868 | Leconte'S Seedcorn Beetle | | Stenolophus (Agonoderus) lineola Fabricius, 1775 | Lined Stenolophus | | Stenolophus (Agonoleptus) thoracicus Casey, 1914 | | | Stenolophus (Stenolophus) carbo Bousquet, 1993 | Scaleless goby | | Stenolophus (Stenolophus) dissimilis Dejean, 1829 | Checked goby | | Stenolophus (Stenolophus) fuliginosus Dejean, 1829 | Checked goby | | Stenolophus (Stenolophus) fuscatus Dejean, 1829 | Checked goby | | Stenolophus (Stenolophus) humidus Hamilton, 1893 | | | Stenolophus (Stenolophus) ochropezus Say, 1823 | | | Stenolophus (Stenolophus) plebejus Dejean, 1829 | Checked goby | | Stenomorphus californicus Ménétriés, 1843 | Kalia | | Stereocerus haematopus Dejean, 1831 | | | Syntomus americanus Dejean, 1831 | | | Synuchus (Pristodactyla) impunctatus Say, 1823 | | | Tetragonoderus (Crossonychus) fasciatus Haldeman, 1843 | | | Tetraleucus picticornis Newman, 1844 | | | Trichosternus vigorsi Gory, 1833 | | | Trichotichnus (Iridessus) autumnalis Say, 1823 | | | Trichotichnus (Trichotichnus) dichrous Dejean, 1829 | | | Trichotichnus (Trichotichnus) vulpeculus Say, 1823 | | | Xestonotus lugubris Dejean, 1829 | | | Zuphioides americanum Dejean, 1831 | |
Subnodes of Harpalinae with occurrences in Pennsylvania | Supertribe | Anthiitae
Bonelli, 1813
| 1
2 | Genus
Species |
Taxom
| | Supertribe | Chlaeniitae
Bonelli, 1810
| 8
48 | Genera
Species |
Taxom
| | Supertribe | Ctenodactylitae
Castelnau, 1834
| 1
1 | Genus
Species |
Taxom
| | Supertribe | Dryptitae
Bonelli, 1810
| 3
5 | Genera
Species |
Taxom
| | Supertribe | Harpalitae
Erwin, 1985
| 23
100 | Genera
Species |
Taxom
| | Supertribe | Lebiitae
Bonelli, 1810
| 19
56 | Genera
Species |
Taxom
| | Supertribe | Platynitae
Bonelli, 1810
| 13
58 | Genera
Species |
Taxom
| | Supertribe | Pterostichitae
Bonelli, 1810
| 16
93 | Genera
Species |
Taxom
| | Tribe | Anisodactylini
Lacordaire, 1854
| 1
0 | Genus
Species |
Taxom
| | Tribe | Glyptini
Cushman & Rohwer, 1920
| 2
2 | Genera
Species |
Taxom
| | Tribe | Pseudomorphini
Newman, 1842
| 1
1 | Genus
Species |
Taxom
|
Links, sources and further information Harpalinae at artfakta.se
Harpalinae at dyntaxa.se
Harpalinae at itis.gov
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