Checklist for Family Scarabaeidae Latreille, 1802 (Dung beetles)
and region Middle Miocene (Miocene)
Class
| Insecta
Linnaeus, 1758
Insects, True insects | 185
683
1400 | Families
Genera
Species |
Taxom
|
Subclass
| Pterygota
Lang, 1888
Winged insects | 185
683
1400 | Families
Genera
Species |
Taxom
|
Infraclass
| Neoptera
van der Wulp, 1890
Modern Wing-folding insects | 173
646
1347 | Families
Genera
Species |
Taxom
|
Superorder
| Holometabola
Heider, 1889
Holometabolous Insects | 108
459
992 | Families
Genera
Species |
Taxom
|
Order
| Coleoptera
Linnaeus, 1758
Beetles, Coleopterans | 41
238
455 | Families
Genera
Species |
Taxom
|
Suborder
| Polyphaga
Emery, 1886
Polyphags | 37
201
370 | Families
Genera
Species |
Taxom
|
Infraorder
| Scarabaeiformia
Crowson, 1960
| 5
32
64 | Families
Genera
Species |
Taxom
|
Superfamily
| Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles | 5
32
64 | Families
Genera
Species |
Taxom
| | Family | Scarabaeidae
Latreille, 1802
Dung beetles, Flower beetles, Fruit chafers, Rain beetles | 25
56 | Genera
Species |
Taxom
|
Genera for Middle Miocene
25 Genera with 56 Species #ACGHLMOPRSTVZ
| Adoretus Laporte, 1994 | | 2 | Species | Taxom | | Anomala Samouelle, 1994 | Shining Leaf Chafer | 13 | Species | Taxom | | Aphodius Hellwig, 1847 | | 3 | Species | Taxom | | Cheirotonus Hope, 1989 | | 1 | Species | Taxom | | Copris Geoffroy, 1862 | Mexican dung beetle | 2 | Species | Taxom | | Genonota Zhang, 1989 | | 2 | Species | Taxom | | Gnorimus Le Peletier ..., 1862 | Chafer | 1 | Species | Taxom | | Gymnopleurus Illiger, 1847 | | 3 | Species | Taxom | | Holotrichia Hope, 1989 | | 2 | Species | Taxom | | Lepitrix Lepeletier &..., 1862 | | 1 | Species | Taxom | | Macronota Hoffmansegg, 1989 | | 1 | Species | Taxom | | Melolonthites Heer, 1847 | | 6 | Species | Taxom | | Mioserica Zhang, 1989 | | 1 | Species | Taxom | | Oniticellus Serville, 1862 | | 1 | Species | Taxom | | Onthophagus Latreille, 1867 | | 5 | Species | Taxom | | Petraeianus Zhang, 1989 | | 1 | Species | Taxom | | Petulantis Zhang & al., 1994 | | 1 | Species | Taxom | Phalops Erichson, 1847 | | 0 | Species | Taxom | | Rhizotrogus Lepeletier &..., 1847 | | 1 | Species | Taxom | | Scarabaeus Linnaeus, 1862 | | 1 | Species | Taxom | | Scelocopris Zhang, 1989 | | 1 | Species | Taxom | | Serica MacLeay, 1862 | | 1 | Species | Taxom | | Trichius Fabricius, 1862 | Bee beetles | 4 | Species | Taxom | | Valgus Scriba, 1862 | | 1 | Species | Taxom | | Zhangsunia Krell, 1994 | | 1 | Species | Taxom |
Species list for Middle Miocene
56 Species | Adoretus recticlypeus ♰ J. F. Zhang, 1989 | | | Adoretus rhinus ♰ J. Zhang, 1994 | | | Anomala amblobelia ♰ J. F. Zhang, 1989 | | | Anomala brachytarsia ♰ J. Zhang, 1994 | | | Anomala endoxa ♰ J. Zhang, 1994 | | | Anomala eversa ♰ J. F. Zhang, 1989 | | | Anomala fugax ♰ O. Heer, 1862 | | | Anomala furva ♰ Y. C. Hong, 1983 | | | Anomala lochmocola ♰ J. F. Zhang, 1989 | | | Anomala orcina ♰ J. Zhang, 1994 | | | Anomala palaeobrunnea ♰ Hong, 1985 | | | Anomala punctulata ♰ J. F. Zhang, 1989 | | | Anomala scia ♰ J. F. Zhang, 1989 | | | Anomala synemosyna ♰ J. F. Zhang, 1989 | | | Anomala ursa ♰ J. Zhang, 1994 | | | Aphodius anteactus ♰ Krell, 2000 | | | Aphodius brevipennis ♰ (O. Heer, 1862) | ⇒ Aphodius foetidus Herbst, 1783 | | Aphodius meyeri ♰ Heer, 1847 | | | Cheirotonus otai ♰ K. Ueda, 1989 | | | Copris druidum ♰ O. Heer, 1862 | | | Copris subterranea ♰ O. Heer, 1862 | | | Genonota mochthera ♰ J. F. Zhang, 1989 | | | Genonota scola ♰ J. F. Zhang, 1989 | | | Gnorimus aedilis ♰ Heer, 1862 | | | Gymnopleurus deperditus ♰ O. Heer, 1862 | | | Gymnopleurus rotundatus ♰ O. Heer, 1862 | | | Gymnopleurus sisyphus ♰ O. Heer, 1847 | | | Holotrichia cressona ♰ J. Zhang, 1994 | | | Holotrichia spatha ♰ J. F. Zhang, 1989 | | | Lepitrix germanica ♰ O. Heer, 1862 | | | Macronota shandongiana ♰ J. F. Zhang, 1989 | | | Melolonthites aciculata ♰ O. Heer, 1847 | | | Melolonthites deperdita ♰ Heer, 1847 | | | Melolonthites kollari ♰ Heer, 1847 | | | Melolonthites lavateri ♰ Heer, 1847 | | | Melolonthites obsoleta ♰ Heer, 1847 | | | Melolonthites parschlugiana ♰ Heer, 1847 | | | Mioserica margelis ♰ J. F. Zhang, 1989 | | | Oniticellus amplicollis ♰ (O. Heer, 1862) | ⇒ Helictopleurus neoamplicollis Krell, 2000 | | Onthophagus bisontinus ♰ Heer, 1862 | | | Onthophagus crassus ♰ (Heer, 1862) | ⇒ Parascatonomus tricornis Wiedemann, 1823 | | Onthophagus ovatulus ♰ Heer, 1847 | | | Onthophagus prodromus ♰ Heer, 1862 | | | Onthophagus urus (Harold, 1867) | ⇒ Onthophagus urusheeri Krell, 2000 | | Petraeianus ruderatus ♰ J. F. Zhang, 1989 | | | Petulantis yimengensis ♰ J. Zhang, 1994 | | | Rhizotrogus longimanus ♰ Heer, 1847 | | | Scarabaeus proserpinae O. Heer, 1862 | | | Scelocopris enertheus ♰ J. F. Zhang, 1989 | | | Serica minutula ♰ (O. Heer, 1862) | ⇒ Gynaecoserica pellecta Brenske, 1896 | | Trichius amoenus ♰ Heer, 1847 | | | Trichius lugubris ♰ O. Heer, 1862 | | | Trichius rotundatus ♰ O. Heer, 1862 | | | Trichius unifasciatus ♰ O. Heer, 1862 | | | Valgus oeningensis ♰ O. Heer, 1862 | | | Zhangsunia extumida ♰ J. Zhang, 1994 | |
Subnodes of Scarabaeidae with occurrences in Middle Miocene | Subfamily | Aphodiinae
Leach, 1815
Aphodiine Dung Beetles, Small dung beetles | 1
3 | Genus
Species |
Taxom
| | Subfamily | Cetoniinae
Leach, 1815
Flower beetles, Flower chafers, Flower scarabs, Fruit and flower chafers | 5
9 | Genera
Species |
Taxom
| | Subfamily | Euchirinae
Hope, 1840
| 1
1 | Genus
Species |
Taxom
| | Subfamily | Melolonthinae
Samouelle, 1819
May Beetles and Junebugs, White grubs | 4
5 | Genera
Species |
Taxom
| | Subfamily | Rutelinae
MacLeay, 1819
Shining Flower Chafers, Shining Leaf Chafers | 2
15 | Genera
Species |
Taxom
| | Subfamily | Scarabaeinae
Latreille, 1802
Dung Beetles, True dung beetles | 7
13 | Genera
Species |
Taxom
|
 | | Scarabaeidae (Latreille 1802) |
Links, sources and further information Scarabaeidae at artfakta.se
Scarabaeidae at dyntaxa.se
Scarabaeidae at gbif.org
Scarabaeidae at irmng.org
Scarabaeidae at itis.gov
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