Checklist for Family Scarabaeidae Latreille, 1802 (Dung beetles)
and region North Macedonia (Yugoslavia)
Class
| Insecta
Linnaeus, 1758
Insects, True insects | 323
2669
7147 | Families
Genera
Species |
Taxom
|
Subclass
| Pterygota
Lang, 1888
Winged insects | 323
2669
7147 | Families
Genera
Species |
Taxom
|
Infraclass
| Neoptera
van der Wulp, 1890
Modern Wing-folding insects | 314
2646
7108 | Families
Genera
Species |
Taxom
|
Superorder
| Holometabola
Heider, 1889
Holometabolous Insects | 212
2187
6237 | Families
Genera
Species |
Taxom
|
Order
| Coleoptera
Linnaeus, 1758
Beetles, Coleopterans | 50
611
1847 | Families
Genera
Species |
Taxom
|
Suborder
| Polyphaga
Emery, 1886
Polyphags | 48
489
1156 | Families
Genera
Species |
Taxom
|
Infraorder
| Scarabaeiformia
Crowson, 1960
| 7
97
199 | Families
Genera
Species |
Taxom
|
Superfamily
| Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles | 7
97
199 | Families
Genera
Species |
Taxom
| | Family | Scarabaeidae
Latreille, 1802
Dung beetles, Flower beetles, Fruit chafers, Rain beetles | 81
171 | Genera
Species |
Taxom
|
Genera for North Macedonia
81 Genera with 171 Species #ABCEGHLMNOPRSTV
| Acanthobodilus Dellacasa, 1799 | | 1 | Species | Taxom | | Acrossus Mulsant, 1758 | | 3 | Species | Taxom | | Agolius Mulsant & Rey, 1812 | | 1 | Species | Taxom | | Agrilinus Mulsant & Rey, 1805 | | 2 | Species | Taxom | | Amidorus Mulsant & Rey, 1800 | | 3 | Species | Taxom | | Amphimallon Lepeletier &..., 0 | European Chafers | 6 | Species | Taxom | | Anisoplia Schönherr, 1783 | | 7 | Species | Taxom | | Anomala Samouelle, 1847 | Shining Leaf Chafer | 4 | Species | Taxom | | Anoxia Laporte, 1832 | | 2 | Species | Taxom | | Anthoplia Medvedev, 1847 | | 1 | Species | Taxom | | Aphodius Hellwig, 1783 | | 1 | Species | Taxom | | Ataenius Harold, 1869 | | 1 | Species | Taxom | | Ateuchetus Bedel, 1819 | | 2 | Species | Taxom | | Blitopertha Reitter, 0 | | 2 | Species | Taxom | | Bodilopsis Adám, 1782 | | 1 | Species | Taxom | | Bodilus Mulsant & Rey, 1799 | | 2 | Species | Taxom | | Butozania Miksic, 1955 | | 2 | Species | Taxom | | Caccobius Thomson, 1767 | | 2 | Species | Taxom | | Cetonia Fabricius, 1761 | | 2 | Species | Taxom | | Chaetonyx Schaum, 1946 | | 1 | Species | Taxom | | Chaetopteroplia Medvedev, 1832 | | 3 | Species | Taxom | | Cheironitis van Lansberge, 1789 | | 2 | Species | Taxom | | Chilothorax Motschulsky, 1840 | | 2 | Species | Taxom | | Colobopterus Mulsant, 1758 | | 1 | Species | Taxom | | Coprimorphus Mulsant, 1789 | | 1 | Species | Taxom | | Copris Geoffroy, 1758 | Mexican dung beetle | 2 | Species | Taxom | | Elaphocera Géné, 1902 | | 1 | Species | Taxom | | Esymus Mulsant & Rey, 1789 | | 2 | Species | Taxom | | Euheptaulacus Dellacasa, 1783 | | 1 | Species | Taxom | | Euoniticellus Janssens, 1781 | | 2 | Species | Taxom | | Euonthophagus Balthasar, 1790 | | 3 | Species | Taxom | | Euorodalus Dellacasa, 1960 | | 1 | Species | Taxom | | Exomala Reitter, 1832 | | 1 | Species | Taxom | | Gnorimus Le Peletier ..., 1758 | Chafer | 3 | Species | Taxom | | Gymnopleurus Illiger, 1821 | | 4 | Species | Taxom | | Haplidia Hope, 1801 | | 1 | Species | Taxom | | Holochelus Reitter, 1823 | | 6 | Species | Taxom | | Hoplia Illiger, 1805 | Monkey Beetles | 3 | Species | Taxom | | Limarus Mulsant & Rey, 1800 | | 1 | Species | Taxom | | Liothorax Motschulsky, 1767 | | 1 | Species | Taxom | | Loraphodius Reitter, 1835 | | 1 | Species | Taxom | | Maladera Mulsant, 1969 | | 1 | Species | Taxom | | Melinopterus Mulsant, 1798 | | 3 | Species | Taxom | | Melolontha Fabricius, 0 | Billy witch | 4 | Species | Taxom | | Mimela Kirby, 1801 | | 1 | Species | Taxom | | Neagolius Kozhantschikov, 1848 | | 2 | Species | Taxom | | Nialus Mulsant & Rey, 1805 | | 1 | Species | Taxom | | Nimbus Mulsant & Rey, 1823 | | 2 | Species | Taxom | | Omaloplia Schönherr, 1994 | | 9 | Species | Taxom | | Onitis Fabricius, 1806 | | 1 | Species | Taxom | | Onthophagus Latreille, 1781 | | 24 | Species | Taxom | | Oryctes Hellwig, 1758 | Coconut rhinoceros beetles | 3 | Species | Taxom | | Osmoderma Le Peletier ..., 1845 | | 1 | Species | Taxom | | Oxyomus Dejean, 1763 | | 1 | Species | Taxom | | Oxythyrea Mulsant, 1761 | | 4 | Species | Taxom | | Pentodon Hope, 1789 | | 3 | Species | Taxom | | Phalacronothus Motschulsky, 1892 | | 1 | Species | Taxom | | Phyllognathus Eschscholtz, 1771 | | 1 | Species | Taxom | | Phyllopertha Stephens, 1758 | | 1 | Species | Taxom | | Planolinoides Dellacasa an..., 1827 | | 1 | Species | Taxom | | Pleurophorus Mulsant, 1961 | | 2 | Species | Taxom | | Polyphylla Harris, 1758 | Lined June Beetles | 1 | Species | Taxom | | Propomacrus Newman, 1781 | | 1 | Species | Taxom | | Protaetia Burmeister, 1833 | | 8 | Species | Taxom | | Psammodius Fallen, 1844 | | 2 | Species | Taxom | | Pseudotrematodes Jacquelin du..., 1836 | | 1 | Species | Taxom | | Rhizotrogus Lepeletier &..., 1789 | | 1 | Species | Taxom | | Rhodaphodius | | 1 | Species | Taxom | | Rhyssemus Mulsant, 1923 | | 2 | Species | Taxom | | Scarabaeus Linnaeus, 1823 | | 2 | Species | Taxom | | Serica MacLeay, 1758 | | 1 | Species | Taxom | | Sisyphus Latreille, 1758 | | 2 | Species | Taxom | | Subrinus Mulsant & Rey, 1870 | | 1 | Species | Taxom | | Teuchestes Mulsant, 1758 | | 1 | Species | Taxom | | Tosevskiana Pavicevic, 1985 | | 1 | Species | Taxom | | Trichius Fabricius, 1906 | Bee beetles | 2 | Species | Taxom | | Trichonotulus Bedel, 1787 | | 1 | Species | Taxom | | Triodontella Reitter, 1962 | | 1 | Species | Taxom | | Tropinota Mulsant, 1783 | | 4 | Species | Taxom | | Valgus Scriba, 0 | | 2 | Species | Taxom | | Volinus Mulsant & Rey, 1798 | | 1 | Species | Taxom |
Species list for North Macedonia
171 Species and 45 Subspecies | Acanthobodilus immundus Creutzer, 1799 | | | Acrossus depressus Kugelann, 1792 | | | Acrossus luridus Fabricius, 1775 | | | Acrossus rufipes Linnaeus, 1758 | Night-Flying Dung Beetle | | Agolius abdominalis abdominalis Bonelli, 1812 | | | Agrilinus ater De Geer, 1774 | | | Agrilinus constans Duftschmid, 1805 | | | Amidorus cribrarius Brullé, 1832 | | | Amidorus obscurus obscurus Fabricius, 1792 | | | Amidorus thermicola Sturm, 1800 | | | Amphimallon (Amphimallon) assimile Herbst, 1790 | | | Amphimallon atrum atrum Herbst, 1790 | | | Amphimallon (Amphimallon) ochraceum (Knoch, 1801) | ⇒ Amphimallon fallenii Gyllenhall, 1817 | | Amphimallon (Amphimallon) solstitiale Linnaeus, 1758 | Cliff Beetle | | Amphimallon solstitiale solstitiale Linnaeus, 1758 | | | Amphimallon vernale vernale | | | Anisoplia (Anisoplia) agricola (Poda von Neu..., 1761) | ⇒ Anisoplia tempestiva Erichson, 1847 | | Anisoplia (Anisoplia) bureschi Sacharieva-S..., 1958 | | | Anisoplia (Anisoplia) hebes Reitter, 1903 | | | Anisoplia (Anisoplia) imitatrix Apfelbeck, 1909 | | | Anisoplia lata lata Erichson, 1847 | | | Anisoplia (Anisoplia) thessalica Reitter, 1889 | | | Anisoplia austriaca austriaca Herbst, 1783 | | | Anomala dubia dubia | | | Anomala (Anomala) vitis Fabricius, 1775 | Vine Chafer | | Anomala errans errans | | | Anomala solida (Erichson, 1847) | ⇒ Anomala immatura Boheman, 1860 | | Anoxia scutellaris rumelica Apfelbeck, 1899 | | | Anoxia (Protanoxia) orientalis Krynizky, 1832 | | | Anthoplia floricola floricola Gistl, 1847 | | | Aphodius (Aphodius) foetidus Herbst, 1783 | | | Ataenius horticola Harold, 1869 | | | Ateuchetus armeniacus Ménétriés..., 1832 | | | Ateuchetus puncticollis Latreille, 1819 | | | Blitopertha lineolata Fischer von ..., 1824 | | | Blitopertha lineolata lineolata | | | Bodilopsis rufus Moll, 1782 | | | Bodilus circumcinctus Schmidt, 1840 | | | Bodilus lugens Creutzer, 1799 | | | Butozania bulgarica Kantardzhiev..., 1953 | | | Butozania macedonica Miksic, 1955 | | | Caccobius histeroides Ménétriés, 1832 | | | Caccobius schreberi Linnaeus, 1767 | | | Cetonia aurata Linnaeus, 1758 | Flower beetle | | Cetonia aurata aurata Linnaeus, 1761 | | | Chaetonyx schatzmayri Mariani, 1946 | | | Chaetopteroplia segetum Herbst, 1783 | | | Chaetopteroplia segetum segetum Herbst, 1783 | Flatspine triangle crab | | Chaetopteroplia segetum straminea Brullé, 1832 | Rhinoceros crab | | Cheironitis furcifer Rossi, 1792 | | | Cheironitis ungaricus Herbst, 1789 | | | Chilothorax distinctus (Müller, 1776) | ⇒ Aphodius distinctus O.F. Müller, 1776 | | Chilothorax melanostictus melanostictus W. Schmidt, 1840 | | | Colobopterus erraticus Linnaeus, 1758 | Erratic small dung beetle | | Coprimorphus scrutator Herbst, 1789 | | | Copris hispanus cavolinii Petagna, 1792 | | | Copris lunaris Linnaeus, 1758 | Horned Dung Beetle | | Elaphocera denticornis Reitter, 1902 | | | Esymus merdarius Fabricius, 1775 | | | Esymus pusillus pusillus Herbst, 1789 | | | Euheptaulacus sus Herbst, 1783 | | | Euoniticellus fulvus Goeze, 1777 | | | Euoniticellus pallipes Fabricius, 1781 | Southern sandy dung beetle | | Euonthophagus amyntas alces Fabricius, 1792 | | | Euonthophagus atramentarius Ménétriés, 1832 | | | Euonthophagus gibbosus Scriba, 1790 | | | Euorodalus paracoenosus Balthasar & ..., 1960 | | | Exomala (Trichopertha) hirtella Brullé, 1832 | | | Gnorimus nobilis Linnaeus, 1758 | Noble Chafer | | Gnorimus nobilis macedonicus Baraud, 1992 | | | Gnorimus nobilis nobilis Linnaeus, 1758 | | | Gymnopleurus geoffroyi geoffroyi | | | Gymnopleurus mopsus Pallas, 1781 | | | Gymnopleurus mopsus mopsus | | | Gymnopleurus sturmii MacLeay, 1821 | | | Haplidia transversa transversa (Fabricius, 1801) | ⇒ Aplidia transversa transversa Fabricius, 1801 | | Holochelus (Holochelus) costulatus Frivaldszky, 1835 | | | Holochelus (Miltotrogus) aequinoctialis Herbst, 1790 | | | Holochelus (Miltotrogus) caucasicus Gyllenhal, 1817 | | | Holochelus (Miltotrogus) fallax Marseul, 1879 | | | Holochelus (Miltotrogus) nocturnus Nonveiller, 1958 | | | Holochelus (Miltotrogus) vernus Germar, 1823 | | | Hoplia argentea Poda, 1761 | | | Hoplia hungarica Burmeister, 1844 | | | Hoplia praticola Duftschmidt, 1805 | | | Limarus maculatus Sturm, 1800 | | | Liothorax plagiatus Linnaeus, 1767 | | | Loraphodius suarius Faldermann, 1835 | | | Maladera (Aserica) apfelbecki Petrovitz, 1969 | | | Melinopterus prodromus Brahm, 1790 | Spring small dung beetle | | Melinopterus pubescens Sturm, 1800 | | | Melinopterus sphacelatus Panzer, 1798 | | | Melolontha (Melolontha) melolontha Linnaeus, 1758 | Cockchafer | | Melolontha hippocastani Fabricius, 1801 | Chestnut cockchafer | | Melolontha pectoralis farinosa Kraatz, 1864 | | | Melolontha pectoralis pectoralis | | | Mimela aurata Fabricius, 1801 | | | Neagolius bilimecki Seidlitz, 1891 | | | Neagolius montanus Erichson, 1848 | | | Nialus varians Duftschmid, 1805 | | | Nimbus affinis affinis | | | Nimbus obliteratus Panzer, 1823 | | | Omaloplia (Omaloplia) caeca Baraud, 1965 | | | Omaloplia (Omaloplia) erythroptera Frivaldszky, 1835 | | | Omaloplia (Omaloplia) gibbosa Baraud, 1965 | | | Omaloplia (Omaloplia) illyrica Baraud, 1965 | | | Omaloplia (Omaloplia) iris Reitter, 1887 | | | Omaloplia (Omaloplia) lonae Schatzmayr, 1923 | | | Omaloplia (Omaloplia) ruricola (Fabricius, 1775) | ⇒ Omaloplia alternata Küster, 1849 | | Omaloplia ruricola ruricola Fabricius, 1775 | | | Omaloplia baraudi pontica Adám, 1994 | | | Onitis damoetas Steven, 1806 | | | Onthophagus (Furconthophagus) furcatus (Fabricius, 1781) | ⇒ Onthophagus furcaticeps Masters, 1886 | | Onthophagus (Onthophagus) illyricus Scopoli, 1763 | | | Onthophagus (Onthophagus) taurus Schreber, 1759 | Bull Headed Dung Beetle | | Onthophagus (Palaeonthophagus) angorensis Petrovitz, 1963 | | | Onthophagus (Palaeonthophagus) coenobita Herbst, 1783 | | | Onthophagus (Palaeonthophagus) dellacasai Pittino & Ma..., 1981 | | | Onthophagus (Palaeonthophagus) fissicornis Steven, 1809 | | | Onthophagus (Palaeonthophagus) fracticornis (Preyssler, 1790) | ⇒ Onthophagus similis Scriba, 1790 | | Onthophagus (Palaeonthophagus) grossepunctatus Reitter, 1905 | | | Onthophagus (Palaeonthophagus) kindermanni Harold, 1877 | | | Onthophagus (Palaeonthophagus) lemur Fabricius, 1781 | | | Onthophagus (Palaeonthophagus) lucidus Illiger, 1800 | | | Onthophagus (Palaeonthophagus) macedonicus Miksic, 1959 | | | Onthophagus (Palaeonthophagus) marginalis Gebler, 1817 | | | Onthophagus marginalis marginalis | | | Onthophagus (Palaeonthophagus) medius (Kugelann, 1792) | ⇒ Onthophagus vacca Linnaeus, 1767 | | Onthophagus (Palaeonthophagus) opacicollis Reitter, 1893 | | | Onthophagus ruficapillus ruficapillus Brullé, 1832 | | | Onthophagus (Palaeonthophagus) semicornis Panzer, 1798 | | | Onthophagus (Palaeonthophagus) sericatus Reitter, 1893 | | | Onthophagus (Palaeonthophagus) similis (Scriba, 1790) | ⇒ Onthophagus fracticornis Preyssler, 1790 | | Onthophagus truchmenus truchmenus | | | Onthophagus (Palaeonthophagus) vacca Linnaeus, 1767 | | | Onthophagus (Palaeonthophagus) verticicornis (Laicharting, 1781) | ⇒ Onthophagus nuchicornis Linnaeus, 1758 | | Oryctes (Oryctes) nasicornis Linnaeus, 1758 | European rhinoceros beetle | | Oryctes nasicornis kuntzeni Minck, 1914 | | | Oryctes nasicornis nasicornis Linnaeus, 1758 | European rhinoceros beetle | | Osmoderma barnabita (Motschulsky, 1845) | ⇒ Osmoderma opicum Lewis, 1887 | | Oxyomus sylvestris Scopoli, 1763 | Forest small dung beetle | | Oxythyrea albopicta Motschulsky, 1845 | | | Oxythyrea cinctella Schaum, 1841 | | | Oxythyrea dulcis Reitter, 1898 | | | Oxythyrea funesta Poda, 1761 | | | Pentodon bidens bidens Pallas, 1771 | | | Pentodon bidens punctatus (Villers, 1789) | ⇒ Pentodon bidens punctatum Villers, 1789 | | Pentodon idiota idiota Herbst, 1789 | | | Phalacronothus fumigatulus Reitter, 1892 | | | Phyllognathus excavatus Forster, 1771 | | | Phyllopertha horticola (Linnaeus, 1758) | ⇒ Phyllopertha intermixta Arrow, 1913 | | Planolinoides borealis Gyllenhal, 1827 | | | Pleurophorus caesus Creutzer, 1796 | Cut small dung beetle | | Pleurophorus pannonicus Petrovitz, 1961 | | | Polyphylla fullo fullo Linnaeus, 1758 | | | Propomacrus bimucronatus Pallas, 1781 | | | Protaetia (Cetonischema) speciosissima (Scopoli, 1786) | ⇒ Cetonischema speciossissima Scopoli, 1786 | | Protaetia (Eupotosia) affinis Andersch, 1797 | | | Protaetia marmorata marmorata Fabricius, 1792 | | | Protaetia (Netocia) angustata Germar, 1817 | | | Protaetia (Netocia) cuprea (Fabricius, 1775) | ⇒ Protaetia distincta Antoine & Pa..., 1994 | | Protaetia cuprea obscura Andersch, 1797 | | | Protaetia ungarica ungarica Herbst, 1790 | | | Protaetia (Netocia) vidua Gory & Perch..., 1833 | | | Psammodius asper (Fabricius, 1775) | ⇒ Psammodius sulcicollis Illiger, 1802 | | Psammodius laevipennis Costa, 1844 | Awkward-winged small dung beetle | | Pseudotrematodes frivaldszkyi Ménétriés, 1836 | | | Rhizotrogus aestivus A. G. Olivier, 1789 | | | Rhodaphodius foetens Fabricius, 1787 | | | Rhyssemus annaedicatus Pierotti, 1980 | | | Rhyssemus macedonicus Benard, 1923 | | | Scarabaeus (Scarabaeus) pius Illiger, 1803 | | | Scarabaeus (Scarabaeus) typhon (Fischer von ..., 1823) | ⇒ Megasoma gyas Herbst, 1785 | | Serica (Serica) brunnea (Linnaeus, 1758) | ⇒ Serica similis Lewis, 1895 | | Sisyphus schaefferi Linnaeus, 1758 | | | Sisyphus schaefferi schaefferi Linnaeus, 1758 | | | Subrinus sturmi Harold, 1870 | | | Teuchestes fossor Linnaeus, 1758 | Digger small dung beetle | | Tosevskiana inexpectata Pavicevic, 1985 | | | Trichius fasciatus Linnaeus, 1758 | Bee Beetle | | Trichius sexualis Bedel, 1906 | | | Trichonotulus scrofa Fabricius, 1787 | Sow small dung beetle | | Triodontella dalmatica Baraud, 1962 | | | Tropinota (Epicometis) hirta Poda, 1761 | Blossom feeder | | Tropinota hirta suturalis Reitter, 1913 | | | Tropinota (Tropinota) squalida Scopoli, 1783 | | | Tropinota squalida squalida Scopoli, 1783 | | | Valgus hemipterus Linnaeus, 1758 | Bug flower chafer | | Valgus hemipterus hemipterus | | | Volinus sticticus Panzer, 1798 | |
Subnodes of Scarabaeidae with occurrences in North Macedonia | Subfamily | Aphodiinae
Leach, 1815
Aphodiine Dung Beetles, Small dung beetles | 33
48 | Genera
Species |
Taxom
| | Subfamily | Cetoniinae
Leach, 1815
Flower beetles, Flower chafers, Flower scarabs, Fruit and flower chafers | 8
19 | Genera
Species |
Taxom
| | Subfamily | Dynastinae
MacLeay, 1819
Horn beetles, Rhinoceros beetles | 3
4 | Genera
Species |
Taxom
| | Subfamily | Euchirinae
Hope, 1840
| 1
1 | Genus
Species |
Taxom
| | Subfamily | Melolonthinae
Samouelle, 1819
May Beetles and Junebugs, White grubs | 16
38 | Genera
Species |
Taxom
| | Subfamily | Orphninae
Erichson, 1847
| 1
1 | Genus
Species |
Taxom
| | Subfamily | Rutelinae
MacLeay, 1819
Shining Flower Chafers, Shining Leaf Chafers | 8
17 | Genera
Species |
Taxom
| | Subfamily | Scarabaeinae
Latreille, 1802
Dung Beetles, True dung beetles | 10
41 | Genera
Species |
Taxom
|
 | | Scarabaeidae (Latreille 1802) |
Links, sources and further information Scarabaeidae at artfakta.se
Scarabaeidae at dyntaxa.se
Scarabaeidae at gbif.org
Scarabaeidae at irmng.org
Scarabaeidae at itis.gov
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