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Checklist for Family Scarabaeidae Latreille, 1802 (Dung beetles)
and region Southeast Europe (Europe)


Class
 
Insecta
Linnaeus, 1758
Insects, True insects
638
10421
42784
Families
Genera
Species
 
Taxom
 

Subclass
 
Pterygota
Lang, 1888
Winged insects
634
10403
42747
Families
Genera
Species
 
Taxom
 

Infraclass
 
Neoptera
van der Wulp, 1890
Modern Wing-folding insects
601
10265
42432
Families
Genera
Species
 
Taxom
 

Superorder
 
Holometabola
Heider, 1889
Holometabolous Insects
390
8244
36333
Families
Genera
Species
 
Taxom
 

Order
 
Coleoptera
Linnaeus, 1758
Beetles, Coleopterans
108
2925
15620
Families
Genera
Species
 
Taxom
 

Suborder
 
Polyphaga
Emery, 1886
Polyphags
99
2614
13015
Families
Genera
Species
 
Taxom
 

Infraorder
 
Scarabaeiformia
Crowson, 1960
9
231
799
Families
Genera
Species
 
Taxom
 

Superfamily
 
Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles
9
231
799
Families
Genera
Species
 
Taxom
 
FamilyScarabaeidae
Latreille, 1802
Dung beetles, Flower beetles, Fruit chafers, Rain beetles
189
677
Genera
Species
 
Taxom
 


Genera for Southeast Europe
189 Genera with 677 Species
#ABCDEFGHKLMNOPRSTV
Acanthobodilus Dellacasa, 17991SpeciesTaxom
Acrossus Mulsant, 17583SpeciesTaxom
Adoretus Laporte, 19141SpeciesTaxom
Aegialia Latreille, 17972SpeciesTaxom
Aethiessa Burmeister, 17871SpeciesTaxom
Agoliinus Schmidt, 18924SpeciesTaxom
Agolius Mulsant & Rey, 20053SpeciesTaxom
Agrilinus Mulsant & Rey, 19466SpeciesTaxom
Ahermodontus Baguena, 19391SpeciesTaxom
Alocoderus Schmidt, 17981SpeciesTaxom
Amadotrogus Amadotrogus, 18861SpeciesTaxom
Amerisaprus1SpeciesTaxom
Amidorus Mulsant & Rey, 18005SpeciesTaxom
Ammoecius Mulsant, 18481SpeciesTaxom
Amphimallon Lepeletier &..., 1823European Chafers40SpeciesTaxom
Anisoplia Schönherr, 178336SpeciesTaxom
Anomala Samouelle, 1845Shining Leaf Chafer16SpeciesTaxom
Anoxia Laporte, 189022SpeciesTaxom
Anthoplia Medvedev, 18472SpeciesTaxom
Aphodius Hellwig, 190142SpeciesTaxom
Aplidia Hope, 18012SpeciesTaxom
Ataenius Harold, 18237SpeciesTaxom
Ateuchetus Bedel, 17876SpeciesTaxom
Biralus Mulsant & Rey, 17891SpeciesTaxom
Blitopertha Reitter, 18884SpeciesTaxom
Bodiloides3SpeciesTaxom
Bodilopsis Adám, 03SpeciesTaxom
Bodilus Mulsant & Rey, 18485SpeciesTaxom
Brancoplia1SpeciesTaxom
Brindalus Landin, 18031SpeciesTaxom
Bubas Mulsant, 19382SpeciesTaxom
Butozania Miksic, 19552SpeciesTaxom
Caccobius Thomson, 17675SpeciesTaxom
Calamosternus Motschulsky, 18633SpeciesTaxom
Callistethus Blanchard, 17921SpeciesTaxom
Canthon Hoffmannsegg, 1805Tumblebug3SpeciesTaxom
Cetonia Fabricius, 18337SpeciesTaxom
Cetonischema Reitter, 17861SpeciesTaxom
Chaetonyx Schaum, 19464SpeciesTaxom
Chaetopteroplia Medvedev, 18448SpeciesTaxom
Cheironitis van Lansberge, 17895SpeciesTaxom
Chilothorax Motschulsky, 18058SpeciesTaxom
Colobopterus Mulsant, 17581SpeciesTaxom
Coprimorphus Mulsant, 17891SpeciesTaxom
Copris Geoffroy, 1901Mexican dung beetle4SpeciesTaxom
Cotinis Burmeister, 1758Green June Beetles2SpeciesTaxom
CotinusSmoketree1SpeciesTaxom
Cremastocheilus Knoch, 1885Anteater Scarab Beetles1SpeciesTaxom
Cyclocephala Dejean, 1847Masked Chafers3SpeciesTaxom
Cyphonotus Bell, 01SpeciesTaxom
Cyptochirus Lesne, 18281SpeciesTaxom
Diastictus Mulsant, 18051SpeciesTaxom
Dichelus Lepeletier &..., 18441SpeciesTaxom
Dichotomius Hope, 17671SpeciesTaxom
Dicronorrhina Burmeister, 18411SpeciesTaxom
Digitonthophagus Balthasar, 20174SpeciesTaxom
Diplotaxis Kirby, 1837Wall-Rockets7SpeciesTaxom
Drepanocerus Kirby, 18513SpeciesTaxom
Dynastes MacLeay, 1766Hercules Beetles1SpeciesTaxom
Dyscinetus Harold, 1847Rice Beetles1SpeciesTaxom
Elaphocera Géné, 187416SpeciesTaxom
Epicometis Burmeister, 17611SpeciesTaxom
Eriesthis Dejean, 18441SpeciesTaxom
Erytus Mulsant & Rey, 19071SpeciesTaxom
Esymus Mulsant & Rey, 18924SpeciesTaxom
Eudolus Mulsant & Rey, 18922SpeciesTaxom
Euetheola Bates, 18471SpeciesTaxom
Euheptaulacus Dellacasa, 18065SpeciesTaxom
Euoniticellus Janssens, 18734SpeciesTaxom
Euonthophagus Balthasar, 17907SpeciesTaxom
Euorodalus Dellacasa, 18483SpeciesTaxom
Euphoria Burmeister, 18522SpeciesTaxom
Eupleurus Mulsant, 17582SpeciesTaxom
Eupotosia Miksic, 18421SpeciesTaxom
Exomala Reitter, 18404SpeciesTaxom
Firminus1SpeciesTaxom
Gnorimus Le Peletier ..., 1758Chafer4SpeciesTaxom
Gymnopleurus Illiger, 18219SpeciesTaxom
Haplidia Hope, 18018SpeciesTaxom
Hellaserica Baraud & Nic..., 18871SpeciesTaxom
Hemichaetoplia Baraud, 18171SpeciesTaxom
Heptaulacus Mulsant, 17751SpeciesTaxom
Heterocnemis Albers, 18321SpeciesTaxom
Heterogomphus Burmeister, 18471SpeciesTaxom
Holochelus Reitter, 183514SpeciesTaxom
Homaloplia Stephens, 19652SpeciesTaxom
Hoplia Illiger, 1885Monkey Beetles18SpeciesTaxom
Hybalus Brullé, 18432SpeciesTaxom
Hypotrichia LeConte, 1862
Hypotrichs0SpeciesTaxom
Kubousa Péringuey, 1902
0SpeciesTaxom
Labarrus Mulsant and Rey, 02SpeciesTaxom
Leiopsammodius Rakovic, 19833SpeciesTaxom
Liatongus Reitter, 1893
0SpeciesTaxom
Limarus Mulsant & Rey, 18132SpeciesTaxom
Liocola Thomson, 17921SpeciesTaxom
Liothorax Motschulsky, 17674SpeciesTaxom
Litocopris Waterhouse, 18691SpeciesTaxom
Loraphodius Reitter, 18351SpeciesTaxom
Loraspis Mulsant & Rey, 18701SpeciesTaxom
Maladera Mulsant, 18509SpeciesTaxom
Mecynodes Mulsant & Rey, 18351SpeciesTaxom
Megasoma Kirby, 1785Elephant beetles1SpeciesTaxom
Megatelus Reitter, 18451SpeciesTaxom
Melinopterus Mulsant, 189211SpeciesTaxom
Melolontha Fabricius, 1999Billy witch9SpeciesTaxom
Mendidaphodius Reitter, 18711SpeciesTaxom
Mesontoplatys Motschulsky, 18751SpeciesTaxom
Metacatharsius Paulian, 18601SpeciesTaxom
Mimela Kirby, 18054SpeciesTaxom
Monotropus Erichson, 18511SpeciesTaxom
Mothon Semenov & Me..., 19271SpeciesTaxom
Neagolius Kozhantschikov, 18482SpeciesTaxom
Neosisyphus Müller, 18181SpeciesTaxom
Netocia A. Costa, 17811SpeciesTaxom
Nialus Mulsant & Rey, 18051SpeciesTaxom
Nimbus Mulsant & Rey, 19946SpeciesTaxom
Nobius Mulsant & Rey, 17992SpeciesTaxom
Omaloplia Schönherr, 026SpeciesTaxom
Oniticellus Serville, 18401SpeciesTaxom
Onitis Fabricius, 18754SpeciesTaxom
Onthophagus Latreille, 197754SpeciesTaxom
Orizabus Fairmaire, 18561SpeciesTaxom
Orodaliscus Reitter, 19001SpeciesTaxom
Oromus Mulsant & Rey, 17631SpeciesTaxom
Oryctes Hellwig, 1758Coconut rhinoceros beetles6SpeciesTaxom
Osmanius Branco & Baraud, 19631SpeciesTaxom
Osmoderma Le Peletier ..., 19914SpeciesTaxom
Otophorus Mulsant, 17581SpeciesTaxom
Oxyomus Dejean, 17631SpeciesTaxom
Oxythyrea Mulsant, 17616SpeciesTaxom
Pachnoda Burmeister, 02SpeciesTaxom
Pachycnema Lepeletier &..., 18441SpeciesTaxom
Pachydema Laporte, 1832
0SpeciesTaxom
Pachydemocera Reitter, 18821SpeciesTaxom
Paracoptochirus Balthasar, 19903SpeciesTaxom
Paragymnopleurus Shipp, 18751SpeciesTaxom
Parammoecius Seidlitz, 18172SpeciesTaxom
Pelidnota MacLeay, 1758Precious Metal Scarabs1SpeciesTaxom
Pentodon Hope, 189915SpeciesTaxom
Phalacronothus Motschulsky, 19175SpeciesTaxom
Phanaeus MacLeay, 1819Burrowing Dung Beetles3SpeciesTaxom
Phyllognathus Eschscholtz, 17711SpeciesTaxom
Phyllopertha Stephens, 19132SpeciesTaxom
Phyllophaga Harris, 1888June beetles10SpeciesTaxom
Plagiogonus Mulsant, 17852SpeciesTaxom
Planolinellus Dellacasa an..., 18251SpeciesTaxom
Planolinoides Dellacasa an..., 18271SpeciesTaxom
Planolinus Mulsant and Rey, 17891SpeciesTaxom
Platytomus Mulsant, 17981SpeciesTaxom
Pleurophorus Mulsant, 19615SpeciesTaxom
Polyphylla Harris, 2014Lined June Beetles9SpeciesTaxom
Popillia Leach, 18381SpeciesTaxom
Potosia Mulsant & Rey, 17874SpeciesTaxom
Propomacrus Newman, 17811SpeciesTaxom
Protaetia Burmeister, 183250SpeciesTaxom
Psammodius Fallen, 180210SpeciesTaxom
Psammoporus Thomson, 17972SpeciesTaxom
Pseudacrossus Reitter, 02SpeciesTaxom
Pseudocanthon Bates, 18471SpeciesTaxom
Pseudotrematodes Jacquelin du..., 18361SpeciesTaxom
Pseudotrochalus Quedenfeldt, 18011SpeciesTaxom
Pubinus Mulsant & Rey, 17761SpeciesTaxom
Rhizotrogus Lepeletier &..., 18505SpeciesTaxom
Rhodaphodius1SpeciesTaxom
Rhysothorax Bedel, 17921SpeciesTaxom
Rhyssemus Mulsant, 196310SpeciesTaxom
Scabrostomus Gordon and S..., 19911SpeciesTaxom
Scarabaeus Linnaeus, 17586SpeciesTaxom
Serica MacLeay, 18175SpeciesTaxom
Sigorus Mulsant & Rey, 17921SpeciesTaxom
Sisyphus Latreille, 17584SpeciesTaxom
Smaragdesthes Kraatz, 18361SpeciesTaxom
Subrinus Mulsant & Rey, 18701SpeciesTaxom
Tanyproctus Ménétriés, 19962SpeciesTaxom
Temnorhynchus Hope, 18562SpeciesTaxom
Temnorrhynchus Alluaud, 18561SpeciesTaxom
Teuchestes Mulsant, 17581SpeciesTaxom
Tibiodrepanus Krikken, 18401SpeciesTaxom
Tomarus Erichson, 1847Carrot Beetles2SpeciesTaxom
Tosevskiana Pavicevic, 19983SpeciesTaxom
Trichesthes Erichson, 17871SpeciesTaxom
Trichiorhyssemus Clouët, 19671SpeciesTaxom
Trichiotinus Casey, 1775Bee-Like Flower Scarabs2SpeciesTaxom
Trichius Fabricius, 1906Bee beetles6SpeciesTaxom
Trichonotulus Bedel, 17871SpeciesTaxom
Triodontella Reitter, 19623SpeciesTaxom
Tropinota Mulsant, 17836SpeciesTaxom
Valgus Scriba, 18054SpeciesTaxom
Volinus Mulsant & Rey, 17981SpeciesTaxom


Species list for Southeast Europe
677 Species and 152 Subspecies
Acanthobodilus immundus Creutzer, 1799
Acrossus depressus Kugelann, 1792
Acrossus luridus Fabricius, 1775
Acrossus rufipes Linnaeus, 1758Night-Flying Dung Beetle
Adoretus (Adoretus) lithobius Ohaus, 1914
Aegialia mimicus Pittino, 2006
Aegialia sabuleti Panzer, 1797
Aethiessa floralis Fabricius, 1787
Agoliinus nemoralis Erichson, 1848
Agoliinus piceus Gyllenhal, 1808
Agoliinus pittinoi Carpaneto, 1986
Agoliinus satyrus Reitter, 1892
Agolius abdominalis Bonelli, 1812
Agolius abdominalis abdominalis Bonelli, 1812
Agolius abdominalis tatrensis Dellacasa & ..., 2005
Agrilinus ater De Geer, 1774
Agrilinus constans Duftschmid, 1805
Agrilinus convexus Erichson, 1848
Agrilinus ibericus Harold, 1874
Agrilinus ibericus ibericus
Agrilinus pirinensis Balthasar, 1946
Ahermodontus bischoffi Vsetecka, 1939
Alocoderus hydrochaeris Fabricius, 1798
Amadotrogus oertzeni Brenske, 1886
Amerisaprus valdivia Stebnicka & ..., 2003
Amidorus cribrarius Brullé, 1832
Amidorus koshantschikovi Jacobson, 1911
Amidorus obscurus Fabricius, 1792
Amidorus obscurus obscurus Fabricius, 1792
Amidorus thermicola Sturm, 1800
Ammoecius brevis Erichson, 1848
Amphimallon (Amphimallon) assimile Herbst, 1790
Amphimallon (Amphimallon) atrum Herbst, 1790
Amphimallon atrum atrum Herbst, 1790
Amphimallon (Amphimallon) burmeisteri (Brenske, 1886)⇒ Amphimallon majale bonadonai Paulian, 1959
Amphimallon (Amphimallon) fuscum Scopoli, 1786
Amphimallon (Amphimallon) majale Razoumowsky, 1789European Chafer
Amphimallon majale bonadonai Paulian, 1959
Amphimallon (Amphimallon) ochraceum (Knoch, 1801)⇒ Amphimallon fallenii Gyllenhall, 1817
Amphimallon (Amphimallon) solstitiale Linnaeus, 1758Cliff Beetle
Amphimallon solstitiale dalmatinum Brenske, 1894
Amphimallon solstitiale grossatum Reitter, 1902
Amphimallon solstitiale orientale Reitter, 1902
Amphimallon solstitiale pictum Kraatz, 1902
Amphimallon solstitiale silvestrii Schatzmayr, 1936
Amphimallon solstitiale simlicissimum J. Müller, 1902
Amphimallon solstitiale simplicissimus Müller, 1902
Amphimallon solstitiale solstitiale Linnaeus, 1758
Amphimallon (Amphimallon) solstitialis (Linnaeus, 1758)⇒ Amphimallon volgense Fischer, 1823
Amphimallon alexandri Uliana & Mon..., 2022
Amphimallon altaicum Mannerheim, 1825
Amphimallon arianae Fairmaire, 1879
Amphimallon arnaudi Montreuil & ..., 2022
Amphimallon brucki Fairmaire, 1879
Amphimallon bruckii Fairmaire, 1879
Amphimallon caucasicum Gyllenhall, 1817
Amphimallon dalmatinum Brenske, 1894
Amphimallon furvum Germar, 1817
Amphimallon furvum furvum
Amphimallon furvum graecum Keith, 2010
Amphimallon jeannei Baraud, 1971
Amphimallon krali Montreuil, 2002
Amphimallon maevae Montreuil, 1999
Amphimallon majalis (Razoumowsky, 1789)⇒ Amphimallon majale Razoumowsky, 1789
Amphimallon maniense Montreuil, 2000
Amphimallon ruficorne Fabricius, 1775
Amphimallon spartanum Brenske, 1884
Amphimallon vernale Brullé, 1832
Amphimallon vernale vernale
Amphimallon verticale Burmeister, 1855
Amphimallon volgense Fischer, 1823
Anisoplia (Anisoplia) agricola (Poda von Neu..., 1761)⇒ Anisoplia tempestiva Erichson, 1847
Anisoplia (Anisoplia) andreae Baraud, 1991
Anisoplia (Anisoplia) aprica Erichson, 1847
Anisoplia (Anisoplia) brenskei Reitter, 1889
Anisoplia (Anisoplia) bromicola Germar, 1817
Anisoplia (Anisoplia) bulgarica Apfelbeck, 1909
Anisoplia (Anisoplia) bureschi Sacharieva-S..., 1958
Anisoplia (Anisoplia) deserticola Fischer von ..., 1824
Anisoplia (Anisoplia) dispar Erichson, 1847
Anisoplia (Anisoplia) erichsoni Reitter, 1889
Anisoplia (Anisoplia) faldermanni Reitter, 1883
Anisoplia (Anisoplia) flavipennis Brullé, 1832
Anisoplia (Anisoplia) hebes Reitter, 1903
Anisoplia (Anisoplia) hebrothracica Kral, 1996
Anisoplia (Anisoplia) imitatrix Apfelbeck, 1909
Anisoplia (Anisoplia) insolita Baraud, 1991
Anisoplia (Anisoplia) lanuginosa Erichson, 1847
Anisoplia (Anisoplia) lata Erichson, 1847
Anisoplia lata lamiensis Apfelbeck, 1908
Anisoplia lata lata Erichson, 1847
Anisoplia (Anisoplia) minor J. Müller, 1902
Anisoplia (Anisoplia) monticola Erichson, 1847
Anisoplia monticola adriatica Müller, 1957
Anisoplia monticola minor Müller, 1902
Anisoplia monticola monticola
Anisoplia monticola muelleri Miksic, 1952
Anisoplia (Anisoplia) pubipennis Blanchard, 1851
Anisoplia (Anisoplia) tempestiva Erichson, 1847
Anisoplia (Anisoplia) tenebralis Burmeister, 1844
Anisoplia (Anisoplia) thessalica Reitter, 1889
Anisoplia (Anisoplia) villosa (Goeze, 1777)⇒ Anisoplia faldermanni Reitter, 1883
Anisoplia (Autanisoplia) austriaca (Herbst, 1783)⇒ Anisoplia tempestiva Erichson, 1847
Anisoplia austriaca austriaca Herbst, 1783
Anisoplia austriaca hordearia Petrovitz, 1959
Anisoplia (Autanisoplia) tritici Kiesenwetter, 1858
Anisoplia segetum Herbst, 1783Cereal chafer
Anomala (Anomala) ausonia Erichson, 1847
Anomala (Anomala) dubia (Scopoli, 1763)⇒ Anomala undulata Melsheimer, 1845
Anomala dubia albanica Reitter, 1918
Anomala dubia dubia
Anomala (Anomala) vitis Fabricius, 1775Vine Chafer
Anomala adscita Robinson, 1941
Anomala affinis Ganglbauer, 1882
Anomala cavifrons LeConte, 1868Redeye triplefin
Anomala errans (Fabricius, 1775)⇒ Anomala quadripunctata Olivier, 1789
Anomala errans errans
Anomala immatura Boheman, 1860
Anomala matzenaueri Reitter, 1918
Anomala osmanlis Blanchard, 1851
Anomala quadripunctata Olivier, 1789
Anomala solida (Erichson, 1847)⇒ Anomala immatura Boheman, 1860
Anomala undulata Melsheimer, 1845Blossom anomala
Anoxia (Anoxia) asiatica Desbrochers, 1871
Anoxia asiatica montandoni Dewailly, 1945
Anoxia (Anoxia) caphtor Petrovitz, 1971
Anoxia (Anoxia) cretica Kiesenwetter, 1858
Anoxia (Anoxia) pasiphae Reitter, 1890
Anoxia pasiphae pasiphae
Anoxia (Anoxia) pilosa (Fabricius, 1792)⇒ Anoxia villosa Fabricius, 1781
Anoxia (Anoxia) scutellaris Mulsant, 1842
Anoxia scutellaris naxiana Reitter, 1890
Anoxia scutellaris rumelica Apfelbeck, 1899
Anoxia scutellaris scutellaris Mulsant, 1842
Anoxia (Anoxia) villosa Fabricius, 1781
Anoxia villosa tethys Reitter, 1890
Anoxia villosa villosa Fabricius, 1781
Anoxia (Mesanoxia) hirta Reitter, 1890
Anoxia (Mesanoxia) matutinalis Laporte, 1832
Anoxia matutinalis matutinalis Laporte de C..., 1823
Anoxia matutinalis suturalis Reitter, 1890
Anoxia (Mesanoxia) reisseri Petrovitz, 1964
Anoxia (Protanoxia) orientalis Krynizky, 1832
Anoxia candiae Miessen & Sa..., 2022
Anoxia meridionalis (Reitter, 1890)⇒ Anoxia orientalis Krynizky, 1832
Anthoplia (Anthoplia) floricola Fabricius, 1787
Anthoplia floricola floricola Gistl, 1847
Aphodius (Acanthobodilus) immundus (Creutzer, 1799)⇒ Pubinus tomentosus O.F.Muller, 1776
Aphodius (Acrossus) luridus (Fabricius, 1775)⇒ Acrossus luridus Fabricius, 1775
Aphodius (Acrossus) rufipes (Linnaeus, 1758)⇒ Acrossus rufipes Linnaeus, 1758
Aphodius (Amidorus) cribrarius Brullé, 1832
Aphodius (Amidorus) obscurus (Fabricius, 1792)⇒ Amidorus obscurus Fabricius, 1792
Aphodius (Aphodius) coniugatus Panzer, 1795
Aphodius (Aphodius) fimetarius Linnaeus, 1758Common dung beetle
Aphodius (Aphodius) foetidus Herbst, 1783
Aphodius (Aphodius) pedellus (DeGeer, 1774)⇒ Aphodius fimetarius Linnaeus, 1758
Aphodius (Biralus) satellitius (Herbst, 1789)⇒ Biralus satellitius Herbst, 1789
Aphodius (Bodilus) ictericus (Laicharting, 1781)⇒ Bodiloides ictericus Laicharting, 1781
Aphodius (Bodilus) lugens (Creutzer, 1799)⇒ Bodilus lugens Creutzer, 1799
Aphodius (Bodilus) rufus (Moll, 1782)⇒ Bodilopsis rufa Moll, 1782
Aphodius (Calamosternus) granarius (Linnaeus, 1767)⇒ Calamosternus granarius Linnaeus, 1767
Aphodius (Chilothorax) conspurcatus (Linnaeus, 1758)⇒ Chilothorax conspurcatus Linnaeus, 1758
Aphodius (Chilothorax) distinctus O.F. Müller, 1776Maculated Dung Beetle
Aphodius (Chilothorax) lineolatus (Illiger, 1803)⇒ Chilothorax lineolatus Illiger, 1803
Aphodius (Chilothorax) melanostictus (W.L.E. Schmidt, 1840)⇒ Chilothorax melanostictus Schmidt, 1840
Aphodius (Chilothorax) sticticus (Panzer, 1798)⇒ Volinus sticticus Panzer, 1798
Aphodius (Colobopterus) erraticus (Linnaeus, 1758)⇒ Colobopterus erraticus Linnaeus, 1758
Aphodius (Coprimorphus) scrutator (Herbst, 1789)⇒ Coprimorphus scrutator Herbst, 1789
Aphodius (Esymus) merdarius (Fabricius, 1775)⇒ Esymus merdarius Fabricius, 1775
Aphodius (Eudolus) quadriguttatus (Herbst, 1783)⇒ Eudolus quadriguttatus Herbst, 1783
Aphodius (Euorodalus) paracoenosus (Balthasar & ..., 1960)⇒ Euorodalus paracoenosus Balthasar & ..., 1960
Aphodius (Euorodalus) pusillus (Herbst, 1789)⇒ Esymus pusillus Herbst, 1789
Aphodius (Loraphodius) suarius (Faldermann, 1835)⇒ Loraphodius suarius Faldermann, 1835
Aphodius (Megatelus) contractus Klug, 1845
Aphodius (Melinopterus) consputus (Creutzer, 1799)⇒ Melinopterus consputus Creutzer, 1799
Aphodius (Melinopterus) sphacelatus (Panzer, 1798)⇒ Melinopterus sphacelatus Panzer, 1798
Aphodius (Nialus) lividus (Olivier, 1789)⇒ Acrossus luridus Fabricius, 1775
Aphodius (Nialus) plagiatus (Linnaeus, 1758)⇒ Liothorax plagiatus Linnaeus, 1767
Aphodius (Nialus) sturmi (Harold, 1870)⇒ Subrinus sturmi Harold, 1870
Aphodius (Nialus) varians (Duftschmid, 1805)⇒ Nialus varians Duftschmid, 1805
Aphodius (Nimbus) contaminatus (Herbst, 1783)⇒ Nimbus contaminatus Herbst, 1783
Aphodius (Nimbus) obliteratus (Panzer, 1823)⇒ Nimbus obliteratus Panzer, 1823
Aphodius (Otophorus) haemorrhoidalis (Linnaeus, 1758)⇒ Otophorus haemorrhoidalis Linnaeus, 1758
Aphodius (Plagiogonus) arenarius Olivier, 1789
Aphodius (Planolinellus) vittatus Mulsant, 1842
Aphodius (Pseudacrossus) thermicola (Sturm, 1800)⇒ Amidorus thermicola Sturm, 1800
Aphodius (Trichonotulus) scrofa (Fabricius, 1787)⇒ Trichonotulus scrofa Fabricius, 1787
Aphodius cardinalis (Reitter, 1892)⇒ Aphodius fimetarius Linnaeus, 1758
Aphodius intercalaris Péringuey, 1901
Aplidia graeca Kraatz, 1882
Aplidia transversa Fabricius, 1801
Ataenius cognatus (LeConte, 1858)⇒ Ataenius liogaster Bates, 1887
Ataenius horticola Harold, 1869
Ataenius opatroides Blanchard, 1846
Ataenius platensis Blanchard, 1846
Ataenius puncticollis (LeConte, 1868)⇒ Parapsammodius puncticollis LeConte, 1858
Ataenius spretulus Haldeman, 1848Black turfgrass ataenius
Ataenius strigatus Say, 1823Black turfgrass ataenius
Ateuchetus armeniacus Ménétriés..., 1832
Ateuchetus cicatricosus Lucas, 1846
Ateuchetus laticollis Linnaeus, 1767
Ateuchetus puncticollis Latreille, 1819
Ateuchetus semipunctatus Fabricius, 1792
Ateuchetus variolosus Fabricius, 1787
Biralus satellitius Herbst, 1789
Blitopertha (Blitopertha) lineata (Fabricius, 1798)⇒ Blitopertha nigripennis Reitter, 1888
Blitopertha lineolata Fischer von ..., 1824
Blitopertha lineolata lineolata
Blitopertha nigripennis Reitter, 1888
Bodiloides ictericus Laicharting, 1781
Bodiloides ictericus ghardimaouensis Balthasar, 1929
Bodiloides ictericus ictericus Laicharting, 1781
Bodilopsis rufa Moll, 1782
Bodilopsis rufus Moll, 1782
Bodilopsis sordidus sordidus
Bodilus caspius Menestries, 1832
Bodilus circumcinctus Schmidt, 1840
Bodilus ictericus (Laicharting, 1781)⇒ Bodiloides ictericus Laicharting, 1781
Bodilus lugens Creutzer, 1799
Bodilus punctipennis Erichson, 1848
Brancoplia leucaspis Castelnau, 1840
Brindalus porcicollis Illiger, 1803
Bubas bison Linnaeus, 1767
Bubas bubaloides Janssens, 1938
Butozania bulgarica Kantardzhiev..., 1953
Butozania macedonica Miksic, 1955
Caccobius denticollis Harold, 1867
Caccobius histeroides Ménétriés, 1832
Caccobius mundus Ménétriés, 1838
Caccobius obtusus Fahraeus, 1857
Caccobius schreberi Linnaeus, 1767
Calamosternus granarius Linnaeus, 1767Granular Small Dung Beetle
Calamosternus mayeri Pilleri, 1953
Calamosternus trucidatus Harold, 1863
Callistethus marginatus Fabricius, 1792
Canthon (Canthon) humectus Say, 1831Longhorn blenny
Canthon (Canthon) imitator Brown, 1946
Canthon (Glaphyrocanthon) viridis (Palisot de B..., 1805)⇒ Epilissus viridis Klug, 1832
Cetonia asiatica Gory & Perch..., 1833
Cetonia aurata Linnaeus, 1758Flower beetle
Cetonia aurata aurata Linnaeus, 1761
Cetonia aurata naxiana Mitter, 2019
Cetonia aurata pallida Drury, 1770
Cetonia aurata pisana Heer, 1841
Cetonia carthami Gory & Perch..., 1833
Cetonischema speciossissima Scopoli, 1786
Chaetonyx binaghii Mariani, 1946
Chaetonyx robustus Schaum, 1862
Chaetonyx robustus robustus
Chaetonyx schatzmayri Mariani, 1946
Chaetopteroplia segetum Herbst, 1783
Chaetopteroplia segetum balcanicola Machatschke, 1961
Chaetopteroplia segetum cretia Baraud, 1993
Chaetopteroplia segetum croatica Machatschke, 1961
Chaetopteroplia segetum segetum Herbst, 1783Flatspine triangle crab
Chaetopteroplia segetum straminea Brullé, 1832Rhinoceros crab
Chaetopteroplia segetum velutina Erichson, 1847
Chaetopteroplia syriaca Burmeister, 1844
Cheironitis furcifer Rossi, 1792
Cheironitis haroldi Ballion, 1870
Cheironitis hungaricus Herbst, 1789
Cheironitis pamphilus Ménétriés, 1849
Cheironitis ungaricus Herbst, 1789
Chilothorax conspurcatus Linnaeus, 1758
Chilothorax distinctus (Müller, 1776)⇒ Aphodius distinctus O.F. Müller, 1776
Chilothorax hieroglyphicus Klug, 1845
Chilothorax lineolatus Illiger, 1803
Chilothorax melanostictus Schmidt, 1840
Chilothorax melanostictus melanostictus W. Schmidt, 1840
Chilothorax paykulli Bedel, 1907
Chilothorax pictus Sturm, 1805
Colobopterus erraticus Linnaeus, 1758Erratic small dung beetle
Coprimorphus scrutator Herbst, 1789
Copris hispanus Linnaeus, 1764
Copris hispanus cavolinii Petagna, 1792
Copris lunaris Linnaeus, 1758Horned Dung Beetle
Copris umbilicatus Abeille de P..., 1901
Cotinis (Cotinis) mutabilis Gory & Perch..., 1883Fig beetle
Cotinis (Cotinis) nitida Linnaeus, 1758Common Green June Beetle
Cotinus coggygriaSmoke-tree
Cremastocheilus (Trinodia) planipes Horn, 1885
Cyclocephala immaculata Olivier, 1789Southern masked chafer
Cyclocephala lurida (Bland, 1863)⇒ Cyclocephala immaculata Olivier, 1789
Cyclocephala signaticollis Burmeister, 1847Argentinian scarab
Cyphonotus incertus
Cyptochirus ambiguus (Kirby, 1828)⇒ Drepanocerus trogiformis Roth, 1851
Diastictus vulneratus Sturm, 1805
Dichelus minutus Burmeister, 1844
Dichotomius carolinus Linnaeus, 1767Carolina copris
Dicronorrhina cavifrons Westwood, 1841
Digitonthophagus aksumensis Génier, 2017
Digitonthophagus eucatta Génier, 2017
Digitonthophagus gazella Fabricius, 1787Brown dung beetle
Digitonthophagus sahelicus Moretto, 2017
Diplotaxis belfragei Fall, 1909
Diplotaxis liberta Germar, 1824
Diplotaxis muralisAnnual Wall-rocket
Diplotaxis muricata Schaeffer, 1907Prairie grain wireworm
Diplotaxis punctata LeConte, 1856Aquatic oligochaete worm
Diplotaxis thoracica Fall, 1909
Diplotaxis tristis Kirby, 1837
Drepanocerus ambiguus Kirby, 1828
Drepanocerus kirbyi Kirby, 1828
Drepanocerus trogiformis Roth, 1851
Dynastes tityus Linnaeus, 1766Eastern Hercules beetle
Dyscinetus picipes Burmeister, 1847Spotted kelpfish
Elaphocera cretica Reitter, 1902
Elaphocera dalmatina Motschulsky, 1859
Elaphocera denticornis Reitter, 1902
Elaphocera elongata Schaufuss, 1874
Elaphocera erberi Kraatz, 1882
Elaphocera gibbifrons Reitter, 1902
Elaphocera gracilis Waltl, 1838
Elaphocera hiemalis Erichson, 1840
Elaphocera insularis Fairmaire, 1877
Elaphocera kosensis Keith & Trichas, 2007
Elaphocera nigroflabellata Burmeister, 1855
Elaphocera ochsi Baraud, 1987
Elaphocera phungae Baraud, 1987
Elaphocera roessneri Keith, 2005
Elaphocera schmidti Keith, 2005
Elaphocera suturalis Schaufuss, 1874
Epicometis hirta (Poda, 1761)⇒ Tropinota hirta Poda, 1761
Eriesthis guttata Burmeister, 1844
Erytus aequalis Schmidt, 1907
Esymus merdarius Fabricius, 1775
Esymus pusillus Herbst, 1789
Esymus pusillus pusillus Herbst, 1789
Esymus sculpturatus Reitter, 1892
Eudolus quadriguttatus Herbst, 1783
Eudolus quadrinaevulus Reitter, 1892
Euetheola humilis Burmeister, 1847Spotted kelpfish
Euheptaulacus carinatus Germar, 1824
Euheptaulacus carinatus carinatus Germar, 1824
Euheptaulacus porcellus Frivaldszky, 1879
Euheptaulacus sus Herbst, 1783
Euheptaulacus villosus Gyllenhal, 1806
Euoniticellus fulvus Goeze, 1777
Euoniticellus pallens Olivier, 1789
Euoniticellus pallipes Fabricius, 1781Southern sandy dung beetle
Euoniticellus triangulatus Harold, 1873
Euonthophagus amyntas Olivier, 1789
Euonthophagus amyntas alces Fabricius, 1792
Euonthophagus amyntas amyntas Olivier, 1789
Euonthophagus atramentarius Ménétriés, 1832
Euonthophagus dorbignyi Olsoufieff, 1900
Euonthophagus dorbignyi neutto Olsoufieff, 1900
Euonthophagus gibbosus Scriba, 1790
Euorodalus coenosus Panzer, 1798
Euorodalus paracoenosus Balthasar & ..., 1960
Euorodalus tersus Erichson, 1848
Euphoria inda Linnaeus, 1758Brown Fruit Chafer
Euphoria kernii Haldeman, 1852Kern´s Flower Scarab
Eupleurus subterraneus Linnaeus, 1758Subterranean small dung beetle
Eupleurus subterraneus subterraneus Linnaeus, 1758
Eupotosia mirifica Mulsant, 1842
Exomala (Neoblitopertha) adriatica Petrovitz, 1968
Exomala (Neoblitopertha) campestris Latreille, 1804
Exomala (Trichopertha) hirtella Brullé, 1832
Exomala succincta Castelnau, 1840
Firminus lautiusculus Schaufuss, 1864
Gnorimus nobilis Linnaeus, 1758Noble Chafer
Gnorimus nobilis macedonicus Baraud, 1992
Gnorimus nobilis nobilis Linnaeus, 1758
Gnorimus variabilis Linnaeus, 1758Variable Chafer
Gymnopleurus cyaneus (Fabricius, 1798)⇒ Gymnopleurus sericeifrons Fairmaire, 1887
Gymnopleurus flagellatus flagellatus Fabricius, 1787
Gymnopleurus geoffroyi Fuessly, 1775
Gymnopleurus geoffroyi geoffroyi
Gymnopleurus geoffroyi virescens Müller, 1902
Gymnopleurus mopsus Pallas, 1781
Gymnopleurus mopsus mopsus
Gymnopleurus sturmi (MacLeay, 1821)⇒ Gymnopleurus sturmii MacLeay, 1821
Gymnopleurus sturmii MacLeay, 1821
Haplidia baudii Kraatz, 1882
Haplidia chloes Keith & Lo C..., 2005
Haplidia graeca Kraatz, 1882
Haplidia hirticollis Burmeister, 1855
Haplidia transversa (Fabricius, 1801)⇒ Aplidia transversa Fabricius, 1801
Haplidia transversa cretica Petrovitz, 1971
Haplidia transversa peloponnisica Petrovitz, 1971
Haplidia transversa transversa (Fabricius, 1801)⇒ Aplidia transversa transversa Fabricius, 1801
Hellaserica elongata Reitter, 1887
Hemichaetoplia pallidipennis Gyllenhal, 1817
Heptaulacus testudinarius Fabricius, 1775
Heterocnemis (Heterocnemis) graeca Brullé, 1832
Heterogomphus schoenherri Burmeister, 1847
Holochelus (Holochelus) costulatus Frivaldszky, 1835
Holochelus (Miltotrogus) aequinoctialis Herbst, 1790
Holochelus (Miltotrogus) caucasicus Gyllenhal, 1817
Holochelus (Miltotrogus) fallax Marseul, 1879
Holochelus (Miltotrogus) fraxinicola Sturm & Hage..., 1825
Holochelus (Miltotrogus) gradojevici Nonveiller, 1965
Holochelus (Miltotrogus) nocturnus Nonveiller, 1958
Holochelus (Miltotrogus) pilicollis Gyllenhall, 1817
Holochelus pilicollis pilicollis Gyllenhal, 1817
Holochelus (Miltotrogus) serrifunis Marseul, 1879
Holochelus (Miltotrogus) tauricus Blanchard, 1851
Holochelus (Miltotrogus) vernus Germar, 1823
Holochelus jarkae Sehnal & Nik..., 2011
Holochelus rusticus Faldermann, 1835
Homaloplia (Acarina) spireae Pallas, 1776
Homaloplia spireae longiclava Baraud, 1965
Hoplia argentea Poda, 1761
Hoplia auriventris Apfelbeck, 1912
Hoplia brunnipes Bonelli, 1807
Hoplia coerulea Drury, 1773
Hoplia corniculata Reitter, 1890
Hoplia dilutipes Reitter, 1890
Hoplia fissa Reitter, 1890
Hoplia flavipes Germar, 1824
Hoplia graminicola Fabricius, 1792
Hoplia herminiana Apfelbeck, 1912
Hoplia hungarica Burmeister, 1844
Hoplia laconiae Petrovitz, 1958
Hoplia nigrina Reitter, 1885
Hoplia philanthus philanthus Fuesslin, 1775
Hoplia praticola Duftschmidt, 1805
Hoplia pseudophilanthus Petrovitz, 1967
Hoplia stenolepis Apfelbeck, 1912
Hoplia uniformis Reitter, 1885
Hybalus cornifrons (Brullé, 1832)⇒ Hybalus graecus Sturm, 1843
Hybalus graecus Sturm, 1843
Labarrus lividus Olivier, 1789Scarab beetle
Labarrus lividus lividus
Leiopsammodius belloi Pierotti, 1980
Leiopsammodius haruspex Adám, 1980
Leiopsammodius strumae Chromy, 1983
Limarus maculatus Sturm, 1800
Limarus zenkeri Germar, 1813
Liocola marmorata Fabricius, 1792
Liothorax isikdagensis Balthasar, 1952
Liothorax kraatzi Harold, 1868
Liothorax niger Illiger, 1798Beaulieu dung beetle
Liothorax plagiatus Linnaeus, 1767
Litocopris simplex Harold, 1869
Loraphodius suarius Faldermann, 1835
Loraspis frater Mulsant & Rey, 1870
Maladera (Aserica) apfelbecki Petrovitz, 1969
Maladera (Aserica) graeca Petrovitz, 1969
Maladera (Aserica) japonica Motschulsky, 1857Velvety chafer
Maladera (Aserica) krueperi Petrovitz, 1969
Maladera (Aserica) punctatissima Falderman, 1835
Maladera (Maladera) holosericea Scopoli, 1772
Maladera elata Küster, 1849
Maladera fusconitens Fairmaire, 1892
Maladera rugosa (C. É. Blanch..., 1850)⇒ Maladera punctatissima Falderman, 1835
Mecynodes striatulus Waltl, 1835
Megasoma gyas Herbst, 1785
Megatelus contractus Klug, 1845
Melinopterus consputus Creutzer, 1799
Melinopterus prodromus Brahm, 1790Spring small dung beetle
Melinopterus pubescens Sturm, 1800
Melinopterus punctatosulcatus Sturm, 1805
Melinopterus punctatosulcatus hirtipes Fischer von ..., 1844
Melinopterus punctatosulcatus ponticus Petrovitz, 1967
Melinopterus punctatosulcatus punctatosulcatus Sturm, 1805
Melinopterus reyi Reitter, 1892
Melinopterus sphacelatus Panzer, 1798
Melinopterus stolzi Reitter, 1906
Melinopterus tingens Reitter, 1892
Melolontha (Melolontha) melolontha Linnaeus, 1758Cockchafer
Melolontha albida (Frivaldszky, 1835)⇒ Melolontha melolontha Linnaeus, 1758
Melolontha farinosa (Kraatz, 1864)⇒ Hoplia graminicola Fabricius, 1792
Melolontha hippocastani Fabricius, 1801Chestnut cockchafer
Melolontha hippocastani hippocastani Fabricius, 1801
Melolontha pectoralis (Megerle von ..., 1812)⇒ Melolontha hippocastani Fabricius, 1801
Melolontha pectoralis farinosa Kraatz, 1864
Melolontha pectoralis pectoralis
Melolontha taygetana Rey, 1999
Mendidaphodius armiger Harold, 1871
Mesontoplatys arabicus Harold, 1875
Metacatharsius exiguus Boheman, 1860
Mimela aurata Fabricius, 1801
Mimela aurata meridionalis Miksic, 1954
Mimela junii Duftschmid, 1805
Mimela junii junii Duftschmid, 1805
Monotropus nordmanni Blanchard, 1851
Mothon sarmaticus Semenov & Me..., 1927
Neagolius bilimecki Seidlitz, 1891
Neagolius montanus Erichson, 1848
Neosisyphus spinipes Thunberg, 1818Grey dungball roller
Netocia morio morio Fabricius, 1781
Nialus varians Duftschmid, 1805
Nimbus affinis Panzer, 1823
Nimbus affinis affinis
Nimbus contaminatus Herbst, 1783
Nimbus johnsoni Baraud, 1976
Nimbus obliteratus Panzer, 1823
Nimbus proximus Adám, 1994
Nobius rhodiensis Baraud, 1976
Nobius serotinus Panzer, 1799
Omaloplia (Omaloplia) alternata Küster, 1849
Omaloplia (Omaloplia) caeca Baraud, 1965
Omaloplia (Omaloplia) cerrutii Sabatinelli, 1977
Omaloplia (Omaloplia) depilis Müller, 1910
Omaloplia (Omaloplia) erythroptera Frivaldszky, 1835
Omaloplia (Omaloplia) gibbosa Baraud, 1965
Omaloplia (Omaloplia) hericius Chobaut, 1907
Omaloplia (Omaloplia) illyrica Baraud, 1965
Omaloplia (Omaloplia) iris Reitter, 1887
Omaloplia (Omaloplia) lonae Schatzmayr, 1923
Omaloplia (Omaloplia) marginata Fuessly, 1775
Omaloplia (Omaloplia) minuta Brenske, 1887
Omaloplia (Omaloplia) polita Baraud, 1965
Omaloplia (Omaloplia) ruricola (Fabricius, 1775)⇒ Omaloplia alternata Küster, 1849
Omaloplia ruricola nicolasi Baraud, 1965
Omaloplia ruricola ruricola Fabricius, 1775
Omaloplia baraudi pontica Adám, 1994
Omaloplia corcyrae Baraud, 1965
Omaloplia gobbii Piattella & ..., 1993
Omaloplia graeca Reitter, 1887
Omaloplia hericia Chobaut, 1907
Omaloplia labrata Burmeister, 1855
Omaloplia nigromarginata Herbst, 1786
Omaloplia pubipennis Rossner & Ah..., 2004
Omaloplia spireae longiclava Baraud, 1965
Omaloplia spireae spireae
Oniticellus planatus Castelnau, 1840
Onitis alexis septentrionalis Balthasar, 1942
Onitis damoetas Steven, 1806
Onitis humerosus Pallas, 1771
Onitis pecuarius Lansberge, 1875
Onthophagus (Furconthophagus) furcatus (Fabricius, 1781)⇒ Onthophagus furcaticeps Masters, 1886
Onthophagus (Onthophagus) illyricus Scopoli, 1763
Onthophagus (Onthophagus) taurus Schreber, 1759Bull Headed Dung Beetle
Onthophagus (Onthophagus) velutinus Horn, 1875
Onthophagus (Palaeonthophagus) angorensis Petrovitz, 1963
Onthophagus (Palaeonthophagus) coenobita Herbst, 1783
Onthophagus (Palaeonthophagus) cruciatus (Ménétriés, 1832)⇒ Onthophagus lemur Fabricius, 1781
Onthophagus (Palaeonthophagus) dellacasai Pittino & Ma..., 1981
Onthophagus (Palaeonthophagus) eulaminicornis Pittino, 2006
Onthophagus (Palaeonthophagus) excisus Reiche & Saulcy, 1856
Onthophagus (Palaeonthophagus) fissicornis Steven, 1809
Onthophagus (Palaeonthophagus) fracticornis (Preyssler, 1790)⇒ Onthophagus similis Scriba, 1790
Onthophagus (Palaeonthophagus) gibbulus Pallas, 1781
Onthophagus gibbulus gibbulus
Onthophagus (Palaeonthophagus) grossepunctatus Reitter, 1905
Onthophagus (Palaeonthophagus) joannae Goljan, 1953
Onthophagus (Palaeonthophagus) kindermanni Harold, 1877
Onthophagus (Palaeonthophagus) lemur Fabricius, 1781
Onthophagus (Palaeonthophagus) lucidus Illiger, 1800
Onthophagus (Palaeonthophagus) macedonicus Miksic, 1959
Onthophagus (Palaeonthophagus) marginalis Gebler, 1817
Onthophagus marginalis marginalis
Onthophagus (Palaeonthophagus) medius (Kugelann, 1792)⇒ Onthophagus vacca Linnaeus, 1767
Onthophagus (Palaeonthophagus) nuchicornis Linnaeus, 1758Mottled dung beetle
Onthophagus (Palaeonthophagus) opacicollis Reitter, 1893
Onthophagus (Palaeonthophagus) ovatus Linnaeus, 1767
Onthophagus (Palaeonthophagus) panici Petrovitz, 1964
Onthophagus (Palaeonthophagus) parmatus Reitter, 1892
Onthophagus (Palaeonthophagus) ponticus Harold, 1883
Onthophagus (Palaeonthophagus) rodentium Pittino, 2004
Onthophagus (Palaeonthophagus) ruficapillus Brullé, 1832
Onthophagus ruficapillus ruficapillus Brullé, 1832
Onthophagus (Palaeonthophagus) semicornis Panzer, 1798
Onthophagus (Palaeonthophagus) sericatus Reitter, 1893
Onthophagus (Palaeonthophagus) similis (Scriba, 1790)⇒ Onthophagus fracticornis Preyssler, 1790
Onthophagus (Palaeonthophagus) suturellus Brullé, 1832
Onthophagus (Palaeonthophagus) tesquorum Semenov & Me..., 1927
Onthophagus truchmenus truchmenus
Onthophagus (Palaeonthophagus) vacca Linnaeus, 1767
Onthophagus (Palaeonthophagus) verticicornis (Laicharting, 1781)⇒ Onthophagus nuchicornis Linnaeus, 1758
Onthophagus (Palaeonthophagus) vitulus (Fabricius, 1776)⇒ Onthophagus immundus Boheman, 1858
Onthophagus andalusicus andalusicus
Onthophagus (Paleonthophagus) fracticornis Preyssler, 1790
Onthophagus (Paleonthophagus) nuchicornis Linnaeus, 1758
Onthophagus (Paleonthophagus) similis Scriba, 1790
Onthophagus (Paleonthophagus) vacca Linnaeus, 1767
Onthophagus amphicoma Boucomont, 1914
Onthophagus binodis Thunberg, 1818Humpbacked Dung Beetle
Onthophagus fissinasus Fairmaire, 1895
Onthophagus furcaticeps Masters, 1886
Onthophagus immundus Boheman, 1858
Onthophagus malthinus Gillet, 1930
Onthophagus mexicanus Bates, 1887
Onthophagus xanthochlorus Walter & Cam..., 1977
Orizabus clunalis LeConte, 1856
Orodaliscus rotundangulus Reitter, 1900
Oromus alpinus Scopoli, 1763
Oryctes (Oryctes) nasicornis Linnaeus, 1758European rhinoceros beetle
Oryctes nasicornis corniculatus Villa & Villa, 1833
Oryctes nasicornis holdhausi Minck, 1914
Oryctes nasicornis kuntzeni Minck, 1914
Oryctes nasicornis laevigatus Heer, 1841
Oryctes nasicornis nasicornis Linnaeus, 1758European rhinoceros beetle
Osmanius balthasari Petrovitz, 1963
Osmoderma barnabita (Motschulsky, 1845)⇒ Osmoderma opicum Lewis, 1887
Osmoderma eremicola Knoch, 1801Hermit Flower Beetle
Osmoderma eremita Scopoli, 1763Hermit Beetle
Osmoderma lassallei Baraud & Tauzin, 1991
Otophorus haemorrhoidalis Linnaeus, 1758Bloody small dung beetle
Oxyomus sylvestris Scopoli, 1763Forest small dung beetle
Oxythyrea abigailoides (Miksic, 1977)⇒ Oxythyrea dulcis Reitter, 1898
Oxythyrea albopicta Motschulsky, 1845
Oxythyrea cinctella Schaum, 1841
Oxythyrea cinctelloides Reitter, 1898
Oxythyrea dulcis Reitter, 1898
Oxythyrea funesta Poda, 1761
Pachnoda savignyi Gory & Perch..., 1833
Pachnoda savignyi savignyi
Pachycnema flavolineata Burmeister, 1844
Pachydemocera lucidicollis Kraatz, 1882
Paracoptochirus kozanensis Pittino, 2001
Paracoptochirus singularis Harold, 1868
Paracoptochirus vignai Carpaneto & ..., 1990
Paragymnopleurus sinuatus abax Sharp, 1875
Parammoecius corvinus Erichson, 1848
Parammoecius gibbus Germar, 1817
Pelidnota (Pelidnota) punctata Linnaeus, 1758Grapevine beetle
Pentodon algerinum Herbst, 1789
Pentodon algerinus (Fuessly, 1778)⇒ Pentodon algerinum Herbst, 1789
Pentodon algerinus algerinus (Fuessly, 1778)⇒ Pentodon algerinum algerinum Herbst, 1789
Pentodon algerinus dispar Baudi, 1870
Pentodon bidens Pallas, 1771
Pentodon bidens bidens Pallas, 1771
Pentodon bidens punctatum Villers, 1789
Pentodon bidens punctatus (Villers, 1789)⇒ Pentodon bidens punctatum Villers, 1789
Pentodon bidens sulcifrons Kuster, 1848
Pentodon caminarius (Faldermann, 1835)⇒ Pentodon idiota Herbst, 1789
Pentodon caminarius caminarius
Pentodon idiota Herbst, 1789
Pentodon idiota idiota Herbst, 1789
Pentodon millini Miksic, 1962
Pentodon variolopunctatus deserti Heyden, 1899
Phalacronothus biguttatus Germar, 1824
Phalacronothus citellorum Semenov & Me..., 1929
Phalacronothus fumigatulus Reitter, 1892
Phalacronothus quadrimaculatus Linnaeus, 1761
Phalacronothus suturinigra Schmidt, 1917
Phanaeus (Phanaeus) amithaon Harold, 1875
Phanaeus (Phanaeus) palliatus Sturm, 1843
Phanaeus (Phanaeus) vindex MacLeay, 1819Laguna Madre fiddler
Phyllognathus excavatus Forster, 1771
Phyllopertha horticola (Linnaeus, 1758)⇒ Phyllopertha intermixta Arrow, 1913
Phyllopertha intermixta Arrow, 1913
Phyllophaga (Phyllophaga) anxia LeConte, 1850Cranberry White Grub
Phyllophaga (Phyllophaga) epigaea Wickham, 1903
Phyllophaga (Phyllophaga) fusca Frölich, 1792Northern June beetle
Phyllophaga (Phyllophaga) hirsuta Knoch, 1801
Phyllophaga (Phyllophaga) hirticula Knoch, 1801
Phyllophaga (Phyllophaga) ilicis Knoch, 1801
Phyllophaga (Phyllophaga) longispina Smith, 1889
Phyllophaga (Phyllophaga) panorpa Sanderson, 1950
Phyllophaga (Phyllophaga) quercus Knoch, 1801
Phyllophaga polyphylla Bates, 1888
Plagiogonus arenarius Olivier, 1789
Plagiogonus putridus (Geoffroy, 1785)⇒ Plagiogonus arenarius Olivier, 1789
Planolinellus vittatus Say, 1825
Planolinoides borealis Gyllenhal, 1827
Planolinus fasciatus Olivier, 1789
Platytomus tibialis Fabricius, 1798
Pleurophorus anatolicus Petrovitz, 1961
Pleurophorus caesus Creutzer, 1796Cut small dung beetle
Pleurophorus maghrebinicus Pittino & Ma..., 1986
Pleurophorus mediterranicus Pittino & Ma..., 1986
Pleurophorus pannonicus Petrovitz, 1961
Polyphylla (Polyphylla) albertischulzi Kuntzen, 1933
Polyphylla (Polyphylla) boryi (Brullé, 1832)⇒ Polyphylla olivieri Castelnau, 1840
Polyphylla boryi albertischultzi Kuntzen, 1933
Polyphylla boryi boryi
Polyphylla (Polyphylla) fullo (Linnaeus, 1758)⇒ Polyphylla olivieri Castelnau, 1840
Polyphylla fullo fullo Linnaeus, 1758
Polyphylla (Polyphylla) naxiana Reitter, 1902
Polyphylla (Polyphylla) olivieri Castelnau, 1840
Polyphylla hassi Hass & Reich..., 2014
Popillia japonica Newman, 1838Japanese beetle
Potosia cuprea (Mulsant, 1842)⇒ Protaetia brevitarsis Lewis, 1879
Potosia cuprea metallica Herbst, 1786
Potosia fieberi Kraatz, 1880
Potosia opaca Fabricius, 1787
Propomacrus bimucronatus Pallas, 1781
Protaetia (Calopotosia) inquinata Arrow, 1913
Protaetia (Cetonischema) aeruginosa (Drury, 1770)⇒ Cetonischema speciossissima Scopoli, 1786
Protaetia (Cetonischema) speciosa Adams, 1817
Protaetia (Cetonischema) speciosissima (Scopoli, 1786)⇒ Cetonischema speciossissima Scopoli, 1786
Protaetia (Eupotosia) affinis Andersch, 1797
Protaetia affinis affinis (Andersch, 1797)⇒ Eupotosia affinis affinis Andersch, 1797
Protaetia (Eupotosia) mirifica (Mulsant, 1842)⇒ Eupotosia mirifica Mulsant, 1842
Protaetia mirifica koenigi Reitter, 1894
Protaetia mirifica mirifica (Mulsant, 1842)⇒ Eupotosia mirifica mirifica Mulsant, 1842
Protaetia (Foveopotosia) judith Reiche, 1871
Protaetia (Liocola) brevitarsis Lewis, 1879
Protaetia (Liocola) lugubris (Herbst, 1786)⇒ Liocola marmorata Fabricius, 1792
Protaetia marmorata marmorata Fabricius, 1792
Protaetia (Macroprotaetia) distincta Antoine & Pa..., 1994
Protaetia (Netocia) afflicta Gory & Perch..., 1833
Protaetia afflicta afflicta Gory & Perch..., 1833
Protaetia (Netocia) angustata Germar, 1817
Protaetia angustata angustata Germar, 1817
Protaetia (Netocia) cretica Kraatz, 1880
Protaetia (Netocia) cuprea (Fabricius, 1775)⇒ Protaetia distincta Antoine & Pa..., 1994
Protaetia cuprea cuprea (Fabricius, 1775)⇒ Potosia cuprea cuprea Fabricius, 1775
Protaetia cuprea ignicollis Gory & Perch..., 1833
Protaetia cuprea levantina Schatzmayr, 1936
Protaetia cuprea metallica (Herbst, 1782)⇒ Potosia cuprea metallica Herbst, 1786
Protaetia cuprea obscura Andersch, 1797
Protaetia (Netocia) cuprina Motschulsky, 1849
Protaetia (Netocia) fieberi (Kraatz, 1880)⇒ Potosia fieberi Kraatz, 1880
Protaetia fieberi boldyrevi Jacobson, 1909
Protaetia fieberi fabriciana Reitter, 1898
Protaetia fieberi fieberi (Kraatz, 1880)⇒ Potosia fieberi fieberi Kraatz, 1880
Protaetia (Netocia) funesta Ménétriés, 1836
Protaetia morio morio (Fabricius, 1781)⇒ Netocia morio morio Fabricius, 1781
Protaetia (Netocia) opaca (Fabricius, 1787)⇒ Potosia opaca Fabricius, 1787
Protaetia opaca cretica Kraatz, 1880
Protaetia (Netocia) rhodensis Rataj, 1998
Protaetia (Netocia) subpilosa Desbrochers ..., 1869
Protaetia subpilosa subpilosa
Protaetia (Netocia) trojana Gory & Perch..., 1833
Protaetia trojana ariadne Mitter, 2019
Protaetia trojana ephesia Tauzin, 1993
Protaetia trojana trojana Gory & Perch..., 1833
Protaetia (Netocia) ungarica Herbst, 1790
Protaetia ungarica ungarica Herbst, 1790
Protaetia ungarica viridana Brullé, 1832
Protaetia (Netocia) vidua Gory & Perch..., 1833
Protaetia hieroglyphica cuprina Motschulsky, 1849
Protaetia (Potosia) metallica Herbst, 1782
Protaetia hungarica hungarica
Protaetia hungarica viridana Brullé, 1832
Psammodius asper (Fabricius, 1775)⇒ Psammodius sulcicollis Illiger, 1802
Psammodius asperoides Pittino, 2007
Psammodius basalis Mulsant & Rey, 1870Crested blenny
Psammodius bulgaricus Mencl, 1982
Psammodius danubialis Adám, 1989
Psammodius laevipennis Costa, 1844Awkward-winged small dung beetle
Psammodius nocturnus Reitter, 1892
Psammodius pierottii Pittino, 1979Mud blenny
Psammodius plicicollis Erichson, 1848
Psammodius sulcicollis Illiger, 1802
Psammoporus mimicus Pittino, 2006
Psammoporus sabuleti (Panzer, 1797)⇒ Psammoporus mimicus Pittino, 2006
Pseudacrossus suffertus Schmidt, 1922
Pseudacrossus suffertus suffertus
Pseudocanthon perplexus LeConte, 1847
Pseudotrematodes frivaldszkyi Ménétriés, 1836
Pseudotrochalus quadrilineatus Fabricius, 1801
Pubinus tomentosus O.F.Muller, 1776
Rhizotrogus (Amphimallon) solstitialis (Linnaeus, 1758)⇒ Amphimallon volgense Fischer, 1823
Rhizotrogus aestivus A. G. Olivier, 1789
Rhizotrogus creticus (Brenske, 1891)⇒ Amphimallon atrum Herbst, 1790
Rhizotrogus marginipes Mulsant, 1842
Rhizotrogus pallidipennis Blanchard, 1850
Rhodaphodius foetens Fabricius, 1787
Rhysothorax rufus Fabricius, 1792Red sand scarab beetle
Rhyssemus algiricus meridionalis Reitter, 1890
Rhyssemus annaedicatus Pierotti, 1980
Rhyssemus berytensis Marseul, 1878
Rhyssemus germanus Linnaeus, 1767Germany small dung beetle
Rhyssemus insularis Pittino, 1983
Rhyssemus macedonicus Benard, 1923
Rhyssemus meridionalis (Reitter, 1890)⇒ Rhyssemus algiricus meridionalis Reitter, 1890
Rhyssemus plicatus Germar, 1817
Rhyssemus puncticollis (Brown, 1929)⇒ Rhyssemus germanus Linnaeus, 1767
Rhyssemus vinodolensis Petrovitz, 1963
Scabrostomus tanytarsus Skelley & Wo..., 1991Masked stargazer
Scarabaeus (Ateuchetus) semipunctatus Fabricius, 1792
Scarabaeus (Ateuchetus) variolosus (Fabricius, 1787)⇒ Ateuchetus semipunctatus Fabricius, 1792
Scarabaeus (Scarabaeus) pius Illiger, 1803
Scarabaeus (Scarabaeus) sacer (Linnaeus, 1758)⇒ Scarabaeus pius Illiger, 1803
Scarabaeus (Scarabaeus) typhon (Fischer von ..., 1823)⇒ Megasoma gyas Herbst, 1785
Scarabaeus indus (Linnaeus, 1758)⇒ Euphoria inda Linnaeus, 1758
Serica (Serica) brunnea (Linnaeus, 1758)⇒ Serica similis Lewis, 1895
Serica evidens Blatchley, 1919
Serica intermixta Blatchley, 1910
Serica sericea Illiger, 1802Onespot fringehead
Serica vespertina Gyllenhal, 1817
Sigorus porcus Fabricius, 1792Mahogany Ridgeback scarab
Sisyphus crispatus Gory, 1833
Sisyphus schaefferi Linnaeus, 1758
Sisyphus schaefferi boschniaki Fischer von ..., 1823
Sisyphus schaefferi schaefferi Linnaeus, 1758
Smaragdesthes africana smaragdina Chevrolat, 1836
Subrinus sturmi Harold, 1870
Tanyproctus reichei Rambur, 1843
Tanyproctus samai Piattella & ..., 1996
Temnorhynchus (Temnorhynchus) baal Reiche & Saulcy, 1856
Temnorhynchus baal baal Reiche & Saulcy, 1856
Temnorrhynchus baal Reiche & Saulcy, 1856
Teuchestes fossor Linnaeus, 1758Digger small dung beetle
Tibiodrepanus sulcicollis Castelnau, 1840
Tomarus gibbosus De Geer, 1774Carrot beetle
Tomarus neglectus LeConte, 1847
Tosevskiana inexpectata Pavicevic, 1985
Tosevskiana machackovae Sehnal, 2004
Tosevskiana sithoniensis Kral, 1998
Trichesthes tristis Fabricius, 1787
Trichiorhyssemus dalmatinus Petrovitz, 1967
Trichiotinus affinis Gory & Perch..., 1833Hairy Flower Scarab
Trichiotinus piger Fabricius, 1775Bee-like Flower Scarab
Trichius fasciatus Linnaeus, 1758Bee Beetle
Trichius gallicus (Dejean, 1821)⇒ Coilodera penicillata Hope, 1831
Trichius gallicus gallicus Dejean, 1821
Trichius orientalis Reitter, 1894
Trichius rosaceus Voet, 1769
Trichius sexualis Bedel, 1906
Trichonotulus scrofa Fabricius, 1787Sow small dung beetle
Triodontella aquila Laporte, 1840
Triodontella bucculenta Baraud, 1962
Triodontella dalmatica Baraud, 1962
Tropinota (Epicometis) hirta Poda, 1761Blossom feeder
Tropinota hirta heyrovskyi Obenberger, 1916
Tropinota hirta hirta Poda, 1761
Tropinota hirta suturalis Reitter, 1913
Tropinota (Tropinota) squalida Scopoli, 1783
Tropinota squalida squalida Scopoli, 1783
Valgus canaliculatus Olivier, 1789
Valgus hemipterus Linnaeus, 1758Bug flower chafer
Valgus hemipterus hemipterus
Valgus seticollis Palisot de B..., 1805Bristly-necked Valgus
Volinus sticticus Panzer, 1798

Subnodes of Scarabaeidae with occurrences in Southeast Europe
SubfamilyAphodiinae
Leach, 1815
Aphodiine Dung Beetles, Small dung beetles
74
206
Genera
Species

Taxom

SubfamilyCetoniinae
Leach, 1815
Flower beetles, Flower chafers, Flower scarabs, Fruit and flower chafers
23
73
Genera
Species

Taxom

SubfamilyDynastinae
MacLeay, 1819
Horn beetles, Rhinoceros beetles
13
22
Genera
Species

Taxom

SubfamilyEuchirinae
Hope, 1840
1
1
Genus
Species

Taxom

SubfamilyMelolonthinae
Samouelle, 1819
May Beetles and Junebugs, White grubs
35
188
Genera
Species

Taxom

SubfamilyOrphninae
Erichson, 1847
2
5
Genera
Species

Taxom

SubfamilyRutelinae
MacLeay, 1819
Shining Flower Chafers, Shining Leaf Chafers
14
61
Genera
Species

Taxom

SubfamilyScarabaeinae
Latreille, 1802
Dung Beetles, True dung beetles
25
114
Genera
Species

Taxom



Scarabaeidae (Latreille 1802)
Scarabaeidae (Latreille 1802)


Links, sources and further information
Scarabaeidae at artfakta.se
Scarabaeidae at dyntaxa.se
Scarabaeidae at gbif.org
Scarabaeidae at irmng.org
Scarabaeidae at itis.gov