AntsAntsArachnidsArachnids
BeesBeesBeetlesBeetles
BookliceBookliceBugsBugs
BumblebeesBumblebeesButterfliesButterflies
CicadasCicadasCrane fliesCrane flies
DragonfliesDragonfliesEarwigsEarwigs
FliesFliesIsopodsIsopods
LocustsLocustsMantisesMantises
MayfliesMayfliesMosquitoesMosquitoes
MyriapodsMyriapodsNet-wingedNet-winged
Plant-parasiticPlant-parasiticScorpion fliesScorpion flies
TermitesTermitesWaspsWasps
INSECTOIDGERENGDARK LIGHT
Home Arthropoda Imprint Contact
TAXFAMGENSPESUB
Checklist for Family Scarabaeidae Latreille, 1802 (Dung beetles)
and region Luzon (Philippines)


Class
 
Insecta
Linnaeus, 1758
Insects, True insects
322
2132
4816
Families
Genera
Species
 
Taxom
 

Subclass
 
Pterygota
Lang, 1888
Winged insects
322
2132
4816
Families
Genera
Species
 
Taxom
 

Infraclass
 
Neoptera
van der Wulp, 1890
Modern Wing-folding insects
307
2075
4704
Families
Genera
Species
 
Taxom
 

Superorder
 
Holometabola
Heider, 1889
Holometabolous Insects
209
1713
4061
Families
Genera
Species
 
Taxom
 

Order
 
Coleoptera
Linnaeus, 1758
Beetles, Coleopterans
60
670
1959
Families
Genera
Species
 
Taxom
 

Suborder
 
Polyphaga
Emery, 1886
Polyphags
56
642
1926
Families
Genera
Species
 
Taxom
 

Infraorder
 
Scarabaeiformia
Crowson, 1960
10
103
395
Families
Genera
Species
 
Taxom
 

Superfamily
 
Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles
10
103
395
Families
Genera
Species
 
Taxom
 
FamilyScarabaeidae
Latreille, 1802
Dung beetles, Flower beetles, Fruit chafers, Rain beetles
73
309
Genera
Species
 
Taxom
 


Genera for Luzon
73 Genera with 309 Species
#ACDEGHILMNOPRSTX
Adoretus Laporte, 19122SpeciesTaxom
Agestrata Eschscholtz, 03SpeciesTaxom
Alissonotum Arrow, 18472SpeciesTaxom
Amphitrichia6SpeciesTaxom
Anomala Samouelle, 1899Shining Leaf Chafer48SpeciesTaxom
Anthracophora Burmeister, 20111SpeciesTaxom
Apogonia Melichar, 191624SpeciesTaxom
Callistethus Blanchard, 18691SpeciesTaxom
Callynomes Mohnike, 20168SpeciesTaxom
Chaetadoretus Ohaus, 19121SpeciesTaxom
Chalcosoma Hope, 18342SpeciesTaxom
Dasyvalgus Kolbe, 19173SpeciesTaxom
Digitonthophagus Balthasar, 17871SpeciesTaxom
Energetia Dalle Torre, 1912
0SpeciesTaxom
Eophileurus Arrow, 19951SpeciesTaxom
Euchirus Kirby, 18411SpeciesTaxom
Euglycyphana Miksic, 19171SpeciesTaxom
Euselates J. Thomson, 03SpeciesTaxom
Exopholis Motschoulsky, 18961SpeciesTaxom
Gilletianus Balthasar, 20106SpeciesTaxom
Glycosia Schoch, 19791SpeciesTaxom
Glycyphana Burmeister, 186712SpeciesTaxom
Haroldius Boucomont, 19542SpeciesTaxom
Hemiphaedimus Miksic, 19161SpeciesTaxom
Hemiprotaetia Miksic, 19172SpeciesTaxom
Heterorhina Westwood, 18732SpeciesTaxom
Heterorrhina Westwood, 18181SpeciesTaxom
Heterovalgus Krikken, 19941SpeciesTaxom
Holotrichia Hope, 19156SpeciesTaxom
Hoplia Illiger, 1910Monkey Beetles2SpeciesTaxom
Ixorida Thomson, 18676SpeciesTaxom
Lepidiota Kirby, 19121SpeciesTaxom
Leucopholis Blanchard, 19245SpeciesTaxom
Macronota Hoffmansegg, 03SpeciesTaxom
Maladera Mulsant, 19266SpeciesTaxom
Malaia Heller, 191216SpeciesTaxom
Microserica Brenske, 18944SpeciesTaxom
Microvalgus Kraatz, 19931SpeciesTaxom
Miksicus Özdikmen & T..., 18671SpeciesTaxom
Mimela Kirby, 19083SpeciesTaxom
Neohyphus Heller, 18831SpeciesTaxom
Neoserica Brenske, 18996SpeciesTaxom
Onitis Fabricius, 17861SpeciesTaxom
Onthophagus Latreille, 197719SpeciesTaxom
Oryctes Hellwig, 1758Coconut rhinoceros beetles2SpeciesTaxom
Panelus Lewis, 19812SpeciesTaxom
Papuana Arrow, 1934Taro beetles3SpeciesTaxom
Parascatonomus Paulian, 19925SpeciesTaxom
Parastasia Westwood, 184110SpeciesTaxom
Peltonotus Burmeister, 19101SpeciesTaxom
Phaedimus Westwood, 18937SpeciesTaxom
Philacelota Heller, 20191SpeciesTaxom
Phyllophaga Harris, 1888June beetles1SpeciesTaxom
Planolinoides Dellacasa an..., 19871SpeciesTaxom
Plectrone Wallace, 19833SpeciesTaxom
Pleuraphodius Schmidt, 19721SpeciesTaxom
Pleuronota Kraatz, 03SpeciesTaxom
Podopogonus Moser, 19171SpeciesTaxom
Popillia Leach, 19245SpeciesTaxom
Protaetia Burmeister, 187327SpeciesTaxom
Protaetiomorpha Miksic, 18731SpeciesTaxom
Pseudohomonyx Arrow, 20001SpeciesTaxom
Reineria Miksic, 193118SpeciesTaxom
Rhyparus Westwood, 19055SpeciesTaxom
Rhyssemodes Reitter, 19841SpeciesTaxom
Rhyssemus Mulsant, 18581SpeciesTaxom
Rutelarcha Waterhouse, 19151SpeciesTaxom
Saprosites Redtenbacher, 19441SpeciesTaxom
Saprovisca1SpeciesTaxom
Taeniodera Burmeister, 18732SpeciesTaxom
Thaumastopeus Kraatz, 18864SpeciesTaxom
Tibiodrepanus Krikken, 18683SpeciesTaxom
Xylotrupes Hope, 0Elephant beetles3SpeciesTaxom


Species list for Luzon
309 Species and 32 Subspecies
Adoretus (Adoretus) luridus Blanchard, 1851
Adoretus semperi Ohaus, 1912
Agestrata luconica Eschscholtz, 1829
Agestrata semperi Mohnike, 1873
Agestrata semperi semperi
Alissonotum pauper Burmeister, 1847Sugarcane beetle
Alissonotum pauperum Burmeister, 1847
Amphitrichia bakeri Moser, 1924
Amphitrichia banahaoensis Moser, 1924
Amphitrichia imugana Moser, 1924
Amphitrichia luzonica Moser, 1926
Amphitrichia monticola Moser, 1915
Amphitrichia sorsogona Chapin, 1931
Anomala dubia aenea De Geer, 1774
Anomala abraensis Frey, 1976
Anomala albaya Arrow, 1915
Anomala andradei Heller, 1893
Anomala anoguttata Burmeister, 1844
Anomala anoguttata anoguttata
Anomala atrocyanea Burmeister, 1844
Anomala baeri Ohaus, 1910
Anomala bakeri Ohaus, 1914
Anomala calpurnia Ohaus, 1923
Anomala camarinensis Ohaus, 1910
Anomala catenatopunctata Ohaus, 1910
Anomala ceramopyga Ohaus, 1912
Anomala chalcoptera Burmeister, 1844
Anomala chalybaea Burmeister, 1844
Anomala chloropyga Burmeister, 1844
Anomala cobala Ohaus, 1916
Anomala dasypyga Burmeister, 1844
Anomala despumata Ohaus, 1910
Anomala ecbolima Ohaus, 1915
Anomala encausta Candeze, 1869
Anomala exanthematica Ohaus, 1914
Anomala flavoscutellata Ohaus, 1910
Anomala heteroglypha Ohaus, 1910
Anomala humeralis (Burmeister, 1844)⇒ Anomala walkeri Arrow, 1899
Anomala inconsueta Ohaus, 1910
Anomala infans Ohaus, 1910
Anomala inopinata Ohaus, 1914
Anomala laetabilis Ohaus, 1915
Anomala leotaudi Blanchard, 1851
Anomala macrophthalma Ohaus, 1910
Anomala maculifemorata Ohaus, 1912
Anomala misandria Ohaus, 1916
Anomala nocturna Ohaus, 1930
Anomala obesa Candeze, 1869
Anomala planata Candeze, 1869
Anomala polina Ohaus, 1923
Anomala praematura Ohaus, 1910
Anomala proctolasia Ohaus, 1910
Anomala sanchezi Ohaus, 1912
Anomala schultzeana Ohaus, 1910
Anomala semperiana Ohaus, 1910
Anomala smaragdina Eschscholtz, 1822
Anomala sulcatula Burmeister, 1844Cloister blenny
Anomala trigonopyga Ohaus, 1912
Anomala vietipennis Ohaus, 1910
Anomala vigilax Ohaus, 1924
Anomala walkeri Arrow, 1899
Anthracophora philippinica Jakl, 2011
Apogonia adoretoides Ritsema, 1897
Apogonia angusta Moser, 1926
Apogonia angustipes Moser, 1924
Apogonia bakeri Moser, 1915
Apogonia banosana Moser, 1924
Apogonia benguetana Moser, 1924
Apogonia callosifrons Moser, 1924
Apogonia depressifrons Kobayashi, 2013
Apogonia imugana Moser, 1924
Apogonia imugana imugana
Apogonia ligaoensis Kobayashi, 2013
Apogonia lutea Moser, 1910
Apogonia luzonica Moser, 1915
Apogonia maculipennis Moser, 1924
Apogonia monticola Moser, 1915
Apogonia nigrobrunnea Moser, 1910
Apogonia pectoralis Moser, 1917
Apogonia philippinica Moser, 1921
Apogonia philochlaenioides Moser, 1924
Apogonia polisana Moser, 1924
Apogonia rugipennis Moser, 1910
Apogonia viridana Moser, 1910
Apogonia viridifulva Heller, 1897
Apogonia viridimicans Moser, 1916
Callistethus picturatus Candeze, 1869
Callynomes borealis Kobayashi & ..., 2016
Callynomes brevispinus Kobayashi & ..., 2016
Callynomes fujiokai Sakai, 1997
Callynomes kobayashii Sasaki, 2019
Callynomes luzonica Schultze, 1916
Callynomes nishiguroi Kobayashi & ..., 2016
Callynomes parva Kobayashi & ..., 2016
Callynomes sakaii Kobayashi & ..., 2016
Chaetadoretus semperi Ohaus, 1912
Chalcosoma atlas Linnaeus, 1758Atlas beetle
Chalcosoma atlas hesperus Erichson, 1834
Dasyvalgus luzonicus Kraatz, 1883
Dasyvalgus nigritus Moser, 1914
Dasyvalgus obsoletus Moser, 1917
Digitonthophagus gazella Fabricius, 1787Brown dung beetle
Eophileurus iwasei Muramoto, 1995
Euchirus dupontianus Burmeister, 1841
Euglycyphana luzonica Moser, 1917
Euselates stictica Hope, 1847
Euselates stictica luzonica Miksic, 1974
Euselates stictica stictica
Exopholis philippinica Brenske, 1896
Gilletianus asinensis Ochi, Kon & ..., 2016
Gilletianus cavitensis Ochi, Kon & ..., 2016
Gilletianus kinabuhayan kinabuhayan
Gilletianus kinabuhayan lucenaensis Ochi, Kon & ..., 2016
Gilletianus luzonensis Ochi, Kon & ..., 2016
Gilletianus takeshii Ochi, Kon & ..., 2010
Glycosia intermedia Mikšić, 1979
Glycyphana aethiessida Wallace, 1867
Glycyphana cuculus Burmeister, 1842
Glycyphana cuculus cuculus
Glycyphana cuculus nigropygidialis Miksic, 1968
Glycyphana cuculus pilifera Miksic, 1967
Glycyphana georgijevici Miksic, 1967
Glycyphana immaculata Moser, 1925
Glycyphana luzonica (Moser, 1917)⇒ Euglycyphana luzonica Moser, 1917
Glycyphana luzonica banghaasi Bourgoin, 1930
Glycyphana luzonica luzonica
Glycyphana quadricolor rimskii Miksic, 1967
Glycyphana vernalis Wallace, 1867
Haroldius globosus Boucomont, 1925
Haroldius philippinensis Pereira, 1954
Hemiphaedimus inermis Moser, 1916
Hemiprotaetia dubia Wallace, 1867
Hemiprotaetia isarogensis Moser, 1917
Heterorhina macleayi Kirby, 1818
Heterorhina simillima Mohnike, 1873
Heterorrhina macleayi Kirby, 1818
Heterovalgus itohi Miyake, 1994
Holotrichia bakeri Moser, 1924
Holotrichia banahaoensis Moser, 1924
Holotrichia bipunctata Brenske, 1892
Holotrichia imugana Moser, 1924
Holotrichia luzonica Moser, 1926
Holotrichia monticola Moser, 1915
Hoplia luzonica Moser, 1924
Hoplia maculifera Moser, 1910
Ixorida albomaculata Moser, 1917
Ixorida mindanaoensis Mohnike, 1873
Ixorida philippinensis Waterhouse, 1841
Ixorida philippinensis nigripennis Miksic, 1970
Ixorida philippinensis philippinensis
Ixorida vidua Wallace, 1867
Lepidiota blanchardi Von Dalle Torre, 1912
Leucopholis irrorata (Chevrolat, 1841)⇒ Leucopholis jacquinoti Blanchard, 1851
Leucopholis jacquinoti Blanchard, 1851
Leucopholis pulverulenta Burmeister, 1855
Leucopholis pulvurelenta Burmeister, 1855
Leucopholis reflexa Moser, 1924
Macronota cupripes Waterhouse, 1841
Macronota inexpectatus inexpectatus
Macronota macgregori macgregori
Maladera analis Brenske, 1899
Maladera boettcheri Moser, 1926
Maladera fasciculata Moser, 1922
Maladera isarogensis Moser, 1922
Maladera kasigurana Moser, 1922
Maladera longiclava Moser, 1926
Malaia (Falsopopillia) scalpta Newman, 1841
Malaia (Pseudomalaia) pilifera Burmeister, 1844
Malaia alticola Ohaus, 1923
Malaia boettcheri Ohaus, 1916
Malaia cetrata Newman, 1841
Malaia conopyga Ohaus, 1905
Malaia depressiuscula Kraatz, 1892
Malaia furcula Ohaus, 1930
Malaia macronyx Ohaus, 1916
Malaia oculata Ohaus, 1910
Malaia pulla Ohaus, 1916
Malaia rugithorax Ohaus, 1916
Malaia semperi Kraatz, 1892
Malaia tagala Heller, 1891
Malaia thoracica Ohaus, 1910
Malaia whiteheadi Ohaus, 1912
Microserica balbalana Moser, 1922
Microserica fugax Erichson, 1834
Microserica humilis Brenske, 1899
Microserica semperi Brenske, 1894
Microvalgus luzonensis Antoine, 1993
Miksicus arrogans Wallace, 1867
Mimela blumei Hope, 1836
Mimela junii rugosula Fairmaire, 1859
Mimela maculicollis (Ohaus, 1908)⇒ Mimela junii rugosula Fairmaire, 1859
Neohyphus semivelutinus Fairmaire, 1883
Neoserica armata Moser, 1922
Neoserica fuscula Moser, 1921
Neoserica longiclava Moser, 1916
Neoserica luzonica Moser, 1915
Neoserica makilingica Moser, 1915
Neoserica uncinata Brenske, 1899
Onitis falcatus Wulfen, 1786
Onthophagus (Gibbonthophagus) sasajii Ochi & Kon, 2001
Onthophagus (Onthophagus) terminatus Eschscholtz, 1822
Onthophagus aereopictus aereopictus
Onthophagus armatus Blanchard, 1853
Onthophagus babirussa Eschscholtz, 1822
Onthophagus bakeri (Boucomont, 1919)⇒ Panelus bakeri Boucomont, 1921
Onthophagus bicolensis Ochi & Kon, 2006
Onthophagus bicolensis asin Ochi & Kon, 2006
Onthophagus citreum Boucomont, 1919
Onthophagus gestroi (Harold, 1877)⇒ Onthophagus bellus d´Orbigny, 1905
Onthophagus ifugaoensis Ochi & Kon, 2006
Onthophagus ohkuboi Ochi & Kon, 2006
Onthophagus poringensis Ochi & Kon, 2005
Onthophagus praedatus Harold, 1862
Onthophagus sexdentatus Boucomont, 1919
Onthophagus surdus Boucomont, 1925
Onthophagus taoi Ochi & Kon, 2006
Onthophagus unidentatus Boucomont, 1927
Onthophagus wiebesi Krikken, 1977
Oryctes gnu Mohnike, 1874Malaysian rhinoceros beetle
Oryctes rhinoceros Linnaeus, 1758Asiatic rhinoceros beetle
Panelus bakeri Boucomont, 1921
Panelus orousseti Paulian, 1981
Papuana cognata Endrödi, 1969
Papuana philippinica Arrow, 1937
Papuana uniformis Prell, 1934
Parascatonomus katoi (Ochi & Kon, 2005)⇒ Onthophagus poringensis Ochi & Kon, 2005
Parascatonomus pseudorudis Ochi & Araya, 1992
Parascatonomus quezonensis Ochi & Araya, 1992
Parascatonomus takedai Ochi & Araya, 1992
Parascatonomus takeshii Ochi & Araya, 1992
Parastasia canaliculata Westwood, 1841
Parastasia confluens Westwood, 1841
Parastasia discolor Westwood, 1841
Parastasia discolor discolor
Parastasia incurva Ohaus, 1923
Parastasia indica Ohaus, 1898
Parastasia luzonensis Wada, 2016
Parastasia nigroscutellata Ohaus, 1901
Parastasia westwoodi Westwood, 1841
Parastasia westwoodii Westwood, 1841
Peltonotus pruinosus Arrow, 1910
Phaedimus asinensis Ochi & Sakai, 2021
Phaedimus cumingi (Waterhouse, 1841)⇒ Phaedimus mohnikei Kraatz, 1893
Phaedimus inermis Moser, 1916
Phaedimus ishidai Nagai, 1998
Phaedimus jagori (Gerstaecker, 1862)⇒ Phaedimus mohnikei Kraatz, 1893
Phaedimus meridionalis Nagai, 1984
Phaedimus mohnikei Kraatz, 1893
Philacelota leucothea Prokofiev, 2019
Phyllophaga polyphylla Bates, 1888
Planolinoides aenictus Cooper & Gordon, 1987Benttooths
Plectrone ismaeli Arnaud, 2004
Plectrone nigrocoerulea Waterhouse, 1841
Plectrone nonveilleri Miksic, 1983
Pleuraphodius luzonensis Petrovitz, 1972
Pleuronota setosa Moser, 1914
Pleuronota setosa imuganica Miksic, 1972
Pleuronota setosa setosa
Podopogonus boettcheri Moser, 1917
Popillia macgregori Ohaus, 1910
Popillia mcgregori Ohaus, 1910
Popillia mutans Newman, 1838
Popillia variabilis Kraatz, 1892
Popillia varicolor Ohaus, 1924
Protaetia (Chalcoprotaetia) franzi Miksic, 1963
Protaetia (Chalcoprotaetia) philippensis Fabricius, 1775
Protaetia (Euprotaetia) mariae Arnaud, 2004
Protaetia (Euprotaetia) nox Janson, 1876
Protaetia (Euprotaetia) uhligi Arnaud, 1987
Protaetia (Heteroprotaetia) fusca Herbst, 1790Asian Mango Flower Beetle
Protaetia (Odontoprotaetia) butozani Miksic, 1963
Protaetia (Odontoprotaetia) cupriceps (Moser, 1917)⇒ Protaetia hiekeiana Miksic, 1970
Protaetia (Odontoprotaetia) hallegueni Arnaud, 2004
Protaetia (Odontoprotaetia) venerabilis Mohnike, 1873
Protaetia venerabilis niveomaculata Moser, 1918
Protaetia venerabilis niveomucalutus Moser, 1918
Protaetia venerabilis venerabilis
Protaetia (Protaetia) ambigua Chevrolat, 1841
Protaetia (Protaetia) ducalis Mohnike, 1873
Protaetia (Protaetia) hiekeiana Miksic, 1970
Protaetia (Protaetia) igorota Schultze, 1916
Protaetia (Protaetia) johani Arnaud, 1992
Protaetia (Protaetia) leucogramma Mohnike, 1873
Protaetia (Protaetia) nocturna Moser, 1909
Protaetia nocturna nocturna
Protaetia (Protaetia) plebeja Mohnike, 1873
Protaetia (Protaetia) subviridis Newman, 1841
Protaetia (Protaetiola) multiguttulata Mohnike, 1873
Protaetia arrogans (Wallace, 1867)⇒ Miksicus arrogans Wallace, 1867
Protaetia guerini Eydoux, 1839
Protaetia multoguttulata Mohnike, 1873
Protaetiomorpha flavovariegata Mohnike, 1873
Pseudohomonyx luzonicus Miyake & Yamaya, 2000
Reineria attenuata Mohnike, 1873
Reineria benguetia Schultze, 1916
Reineria benquetia Schultze, 1916
Reineria biguttulata Mohnike, 1873
Reineria coeruleosignata Miksic, 1979
Reineria eckeli Schürhoff, 1933
Reineria fasciata Schurhoff, 1935
Reineria flavopunctata Blanchard, 1850
Reineria flavopunctata flavopunctata
Reineria francolina bulusanica Miksic, 1981
Reineria intermedia Miksic, 1979
Reineria kuntzeni Schürhoff, 1933
Reineria luzonica Moser, 1922
Reineria megaspilota Wallace, 1867
Reineria moseri Bourgoin, 1931
Reineria punctiventris Moser, 1917
Reineria schaueri Schürhoff, 1933
Reineria simillima Bourgoin, 1931
Rhyparus barclayi Masumoto, Oc..., 2019
Rhyparus helophoroides Fairmaire, 1893
Rhyparus ifugaoensis Ochi & Kakizoe, 2021
Rhyparus luzonensis Ochi & Kakizoe, 2021
Rhyparus philippinensis Arrow, 1905
Rhyssemodes sindicus Pittino, 1984
Rhyssemus inscitus Walker, 1858
Rutelarcha bakeri Ohaus, 1915
Saprosites longethorax Paulian, 1944
Saprovisca sarangay Skelley, 2015
Taeniodera gratiosa Mohnike, 1873
Taeniodera tricolor Mohnike, 1873
Thaumastopeus cupripes Waterhouse, 1841
Thaumastopeus inexpectatus Miksic, 1977
Thaumastopeus macgregori Schultze, 1915
Thaumastopeus shangaicus Neervoort Va..., 1886
Tibiodrepanus hircus Wiedemann, 1823
Tibiodrepanus setosus Wiedemann, 1823
Tibiodrepanus sinicus Harold, 1868
Xylotrupes lumawigi Silvestre, 2002
Xylotrupes pubescens Waterhouse, 1841
Xylotrupes pubescens pubescens

Subnodes of Scarabaeidae with occurrences in Luzon
SubfamilyAphodiinae
Leach, 1815
Aphodiine Dung Beetles, Small dung beetles
8
16
Genera
Species

Taxom

SubfamilyCetoniinae
Leach, 1815
Flower beetles, Flower chafers, Flower scarabs, Fruit and flower chafers
25
98
Genera
Species

Taxom

SubfamilyDynastinae
MacLeay, 1819
Horn beetles, Rhinoceros beetles
8
13
Genera
Species

Taxom

SubfamilyEuchirinae
Hope, 1840
1
1
Genus
Species

Taxom

SubfamilyMelolonthinae
Samouelle, 1819
May Beetles and Junebugs, White grubs
13
62
Genera
Species

Taxom

SubfamilyRutelinae
MacLeay, 1819
Shining Flower Chafers, Shining Leaf Chafers
9
85
Genera
Species

Taxom

SubfamilyScarabaeinae
Latreille, 1802
Dung Beetles, True dung beetles
6
27
Genera
Species

Taxom



Scarabaeidae (Latreille 1802)
Scarabaeidae (Latreille 1802)


Links, sources and further information
Scarabaeidae at artfakta.se
Scarabaeidae at dyntaxa.se
Scarabaeidae at gbif.org
Scarabaeidae at irmng.org
Scarabaeidae at itis.gov