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Checklist for Family Scarabaeidae Latreille, 1802 (Dung beetles)
and region Pennsylvania (USA)


Class
 
Insecta
Linnaeus, 1758
Insects, True insects
554
5452
12530
Families
Genera
Species
 
Taxom
 

Subclass
 
Pterygota
Lang, 1888
Winged insects
553
5449
12526
Families
Genera
Species
 
Taxom
 

Infraclass
 
Neoptera
van der Wulp, 1890
Modern Wing-folding insects
525
5347
12273
Families
Genera
Species
 
Taxom
 

Superorder
 
Holometabola
Heider, 1889
Holometabolous Insects
355
4526
10429
Families
Genera
Species
 
Taxom
 

Order
 
Coleoptera
Linnaeus, 1758
Beetles, Coleopterans
102
1569
3485
Families
Genera
Species
 
Taxom
 

Suborder
 
Polyphaga
Emery, 1886
Polyphags
95
1388
2794
Families
Genera
Species
 
Taxom
 

Infraorder
 
Scarabaeiformia
Crowson, 1960
9
109
285
Families
Genera
Species
 
Taxom
 

Superfamily
 
Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles
9
109
285
Families
Genera
Species
 
Taxom
 
FamilyScarabaeidae
Latreille, 1802
Dung beetles, Flower beetles, Fruit chafers, Rain beetles
85
222
Genera
Species
 
Taxom
 


Genera for Pennsylvania
85 Genera with 222 Species
#ABCDEGHILMNOPRSTVX
Acrossus Mulsant, 17583SpeciesTaxom
Aegialia Latreille, 18871SpeciesTaxom
Agoliinus Schmidt, 19283SpeciesTaxom
Alloblackburneus Bordat, 18701SpeciesTaxom
Amphimallon Lepeletier &..., 1789European Chafers1SpeciesTaxom
Anomala Samouelle, 1875Shining Leaf Chafer8SpeciesTaxom
Aphodius Hellwig, 18768SpeciesTaxom
Aphonus LeConte, 18233SpeciesTaxom
Apicencya1SpeciesTaxom
Ataenius Harold, 188714SpeciesTaxom
Ateuchus Weber, 18011SpeciesTaxom
Blackburneus Schmidt, 18451SpeciesTaxom
Bothynus Hope, 1837
0SpeciesTaxom
Caccobius Thomson, 19041SpeciesTaxom
Calamosternus Motschulsky, 17671SpeciesTaxom
Callistethus Blanchard, 17921SpeciesTaxom
Canthon Hoffmannsegg, 1858Tumblebug4SpeciesTaxom
Chasmatopterus Latreille, 18031SpeciesTaxom
Chilothorax Motschulsky, 17761SpeciesTaxom
Colobopterus Mulsant, 17581SpeciesTaxom
Copris Geoffroy, 1773Mexican dung beetle3SpeciesTaxom
Cotalpa Burmeister, 17581SpeciesTaxom
Cotinis Burmeister, 1758Green June Beetles1SpeciesTaxom
CotinusSmoketree1SpeciesTaxom
Cremastocheilus Knoch, 1827Anteater Scarab Beetles4SpeciesTaxom
Cryptoscatomaseter Gordon and S..., 20011SpeciesTaxom
Cyclocephala Dejean, 1992Masked Chafers7SpeciesTaxom
Dermolepida Arrow, 1941
0SpeciesTaxom
Dialytellus Brown, 19071SpeciesTaxom
Dialytes Harold, 18753SpeciesTaxom
Dichelonyx Harris, 18567SpeciesTaxom
Dichotomius Hope, 17671SpeciesTaxom
Diplotaxis Kirby, 1837Wall-Rockets7SpeciesTaxom
Dynastes MacLeay, 1766Hercules Beetles1SpeciesTaxom
Dyscinetus Harold, 1798Rice Beetles2SpeciesTaxom
Euetheola Bates, 18562SpeciesTaxom
Euphoria Burmeister, 18016SpeciesTaxom
Exomala Reitter, 18751SpeciesTaxom
Geomyphilus Gordon and S..., 19281SpeciesTaxom
Gnorimella Casey, 18011SpeciesTaxom
Hoplia Illiger, 1775Monkey Beetles6SpeciesTaxom
Irrasinus Gordon and S..., 18701SpeciesTaxom
Labarrus Mulsant and Rey, 19411SpeciesTaxom
Lachnosterna Hope, 1837
0SpeciesTaxom
Lechorodius Gordon and S..., 18231SpeciesTaxom
Leucopholis Blanchard, 18511SpeciesTaxom
Ligyrus Erichson, 18252SpeciesTaxom
Macrodactylus Dejean, 1775Rose Chafers1SpeciesTaxom
Maladera Mulsant, 18982SpeciesTaxom
Melanocanthon Halffter, 19411SpeciesTaxom
Melinopterus Mulsant, 17902SpeciesTaxom
Nipponoserica Nomura, 19381SpeciesTaxom
Onthophagus Latreille, 193014SpeciesTaxom
Oscarinus Gordon and S..., 18451SpeciesTaxom
Osmoderma Le Peletier ..., 18054SpeciesTaxom
Otophorus Mulsant, 17581SpeciesTaxom
Oxyomus Dejean, 17631SpeciesTaxom
Pachystethus Blanchard, 01SpeciesTaxom
Parastasia Westwood, 18561SpeciesTaxom
Pardalosus Gordon and S..., 18351SpeciesTaxom
Pelidnota MacLeay, 1922Precious Metal Scarabs2SpeciesTaxom
Phanaeus MacLeay, 1876Burrowing Dung Beetles2SpeciesTaxom
Phileurus Sieg, 17581SpeciesTaxom
Phyllophaga Harris, 1856June beetles38SpeciesTaxom
Planolinellus Dellacasa an..., 18251SpeciesTaxom
Pleurophorus Mulsant, 17961SpeciesTaxom
Polyphylla Harris, 1830Lined June Beetles2SpeciesTaxom
Popillia Leach, 18381SpeciesTaxom
Psammodius Fallen, 18441SpeciesTaxom
Pseudagolius Schmidt, 18231SpeciesTaxom
Rhyssemus Mulsant, 18481SpeciesTaxom
Scabrostomus Gordon and S..., 19911SpeciesTaxom
Serica MacLeay, 181711SpeciesTaxom
Stenotothorax Schmidt, 18451SpeciesTaxom
Strategus Kirby, 1773Ox Beetles2SpeciesTaxom
Strigoderma Burmeister, 17982SpeciesTaxom
Teuchestes Mulsant, 17581SpeciesTaxom
Tomarus Erichson, 1825Carrot Beetles2SpeciesTaxom
Trichesthes Erichson, 17871SpeciesTaxom
Trichiotinus Casey, 1775Bee-Like Flower Scarabs4SpeciesTaxom
Trichonotulus Bedel, 17871SpeciesTaxom
Trichopopillia Ohaus, 19231SpeciesTaxom
Trigonopeltastes Burmeister &..., 17711SpeciesTaxom
Valgus Scriba, 18052SpeciesTaxom
Xyloryctes Hope, 1773Rhinoceros Beetles1SpeciesTaxom


Species list for Pennsylvania
222 Species and 11 Subspecies
Acrossus luridus Fabricius, 1775
Acrossus rubripennis Horn, 1870
Acrossus rufipes Linnaeus, 1758Night-Flying Dung Beetle
Aegialia (Aegialia) opifex Horn, 1887
Agoliinus bidentatus Schmidt, 1906Spotted weever
Agoliinus leopardus Horn, 1870
Agoliinus manitobensis Brown, 1928
Alloblackburneus lentus Horn, 1870
Amphimallon (Amphimallon) majale Razoumowsky, 1789European Chafer
Anomala adscita Robinson, 1941
Anomala bruchomorpha Arrow, 1912
Anomala incolumis Casey, 1915
Anomala innuba Fabricius, 1787
Anomala lucicola Fabricius, 1798
Anomala margina Lin, 1999
Anomala oblivia Horn, 1884Pine chafer
Anomala orientalis Waterhouse, 1875Oriental beetle
Aphodius (Acrossus) rubripennis Horn, 1870
Aphodius (Acrossus) rufipes (Linnaeus, 1758)⇒ Acrossus rufipes Linnaeus, 1758
Aphodius (Aphodius) fimetarius Linnaeus, 1758Common dung beetle
Aphodius (Aphodius) pedellus (DeGeer, 1774)⇒ Aphodius fimetarius Linnaeus, 1758
Aphodius (Blackburneus) stercorosus (Melsheimer, 1845)⇒ Blackburneus stercorosus Melsheimer, 1845
Aphodius (Nialus) lividus (Olivier, 1789)⇒ Acrossus luridus Fabricius, 1775
Aphodius (Oscarinus) rusicola (Melsheimer, 1845)⇒ Oscarinus rusicola Melsheimer, 1845
Aphodius precursor G. H. Horn, 1876
Aphonus castaneus Melsheimer, 1845
Aphonus densicauda Casey, 1915
Aphonus tridentatus Say, 1823
Apicencya apicalis Blanchard, 1851
Ataenius abditus Haldeman, 1848
Ataenius alternatus Melsheimer, 1845
Ataenius brevis Fall, 1930
Ataenius cognatus (LeConte, 1858)⇒ Ataenius liogaster Bates, 1887
Ataenius cylindrus (Horn, 1871)⇒ Ataenius ovatulus Horn, 1871
Ataenius gracilis Melsheimer, 1845
Ataenius imbricatus Melsheimer, 1845
Ataenius liogaster Bates, 1887
Ataenius ovatulus Horn, 1871Scratchbush
Ataenius spretulus Haldeman, 1848Black turfgrass ataenius
Ataenius strigatus Say, 1823Black turfgrass ataenius
Ataenius texanus Harold, 1874
Ataenius walterhorni Balthasar, 1938
Ataenius wenzeli Horn, 1887
Ateuchus histeroides Weber, 1801
Blackburneus stercorosus Melsheimer, 1845
Caccobius anthracites d´Orbigny, 1904
Calamosternus granarius Linnaeus, 1767Granular Small Dung Beetle
Callistethus marginatus Fabricius, 1792
Canthon (Canthon) chalcites Haldeman, 1843
Canthon (Canthon) imitator Brown, 1946
Canthon (Canthon) pilularius Linnaeus, 1758Dull tumblebug
Canthon (Canthon) vigilans LeConte, 1858Kalia
Chasmatopterus hirtulus Illiger, 1803
Chilothorax distinctus (Müller, 1776)⇒ Aphodius distinctus O.F. Müller, 1776
Colobopterus erraticus Linnaeus, 1758Erratic small dung beetle
Copris fricator Fabricius, 1787
Copris fricator fricator
Copris minutus Drury, 1773Small Black Dung Beetle
Cotalpa lanigera Linnaeus, 1758Goldsmith beetle
Cotinis (Cotinis) nitida Linnaeus, 1758Common Green June Beetle
Cotinus nitidaGreen June Beetle
Cremastocheilus (Cremastocheilus) canaliculatus Kirby, 1827Imitator blenny
Cremastocheilus (Cremastocheilus) castaneae (Knoch, 1801)⇒ 
Cremastocheilus (Cremastocheilus) variolosus Kirby, 1826Bowman stygobromid
Cremastocheilus harrisi Kirby, 1827
Cryptoscatomaseter gambrinus Skelley & Go..., 2001Warteye stargazer
Cyclocephala borealis Arrow, 1911Northern masked chafer
Cyclocephala immaculata Olivier, 1789Southern masked chafer
Cyclocephala lurida (Bland, 1863)⇒ Cyclocephala immaculata Olivier, 1789
Cyclocephala lurida lurida Bland, 1863Southern masked chafer
Cyclocephala pasadenae Casey, 1915Pasadena masked chafer
Cyclocephala villosa (Blanchard, 1846)⇒ Cyclocephala borealis Arrow, 1911
Cyclocephala zodion Ratcliffe, 1992
Dialytellus dialytoides Fall & Cocke..., 1907
Dialytes striatulus Say, 1825
Dialytes truncatus Melsheimer, 1845
Dialytes ulkei Horn, 1875
Dichelonyx albicollis Burmeister, 1855Lamellicorn Beetle
Dichelonyx canadensis Horn, 1876
Dichelonyx diluta Fall, 1901
Dichelonyx elongata Fabricius, 1792
Dichelonyx elongatula Schönherr, 1817Solf-winged flower beetles
Dichelonyx fuscula LeConte, 1856Aquatic oligochaete worm
Dichelonyx subvittata LeConte, 1856Aquatic oligochaete worm
Dichotomius carolinus Linnaeus, 1767Carolina copris
Diplotaxis atlantis Fall, 1909
Diplotaxis frondicola Say, 1825
Diplotaxis harperi Blanchard, 1851
Diplotaxis liberta Germar, 1824
Diplotaxis sordida Say, 1825
Diplotaxis subcostata Blanchard, 1851
Diplotaxis tristis Kirby, 1837
Dynastes tityus Linnaeus, 1766Eastern Hercules beetle
Dyscinetus dubius Olivier, 1789
Dyscinetus morator Fabricius, 1798Rice Beetle
Euetheola humilis Burmeister, 1847Spotted kelpfish
Euetheola rugiceps (LeConte, 1856)⇒ Euetheola humilis Burmeister, 1847
Euphoria areata Fabricius, 1775
Euphoria fulgida (Fabricius, 1775)⇒ Neomyia cornicina Fabricius, 1781
Euphoria herbacea Olivier, 1789Olive Euphoria
Euphoria inda Linnaeus, 1758Brown Fruit Chafer
Euphoria lurida Fabricius, 1775
Euphoria sepulcralis Fabricius, 1801Dark Flower Scarab
Exomala orientalis C. O. Waterh..., 1875Asiatic beetle
Geomyphilus leptotarsis Brown, 1928
Gnorimella maculosa Knoch, 1801Maculate flower beetle
Hoplia (Hoplia) equina LeConte, 1880
Hoplia (Hoplia) modesta Haldemann, 1843
Hoplia (Hoplia) trifasciata Say, 1823Three lined Hoplia
Hoplia (Hoplia) trivialis Harold, 1869Dark Hoplia
Hoplia fernsi
Hoplia philanthus philanthus Fuesslin, 1775
Irrasinus stupidus Horn, 1870
Labarrus pseudolividus Balthasar, 1941
Lechorodius terminalis Say, 1823
Leucopholis lepidophora (Blanchard, 1851)⇒ Leucopholis burmeisteri Brenske, 1894
Ligyrus gibbosus DeGeer, 1774Carrot beetle
Ligyrus relictus Say, 1825
Macrodactylus subspinosus Fabricius, 1775American Rose Chafer
Maladera (Cephaloserica) castanea Arrow, 1913Asian garden beetle
Maladera formosae Brenske, 1898Asiatic Garden Beetle
Melanocanthon bispinatus Robinson, 1941
Melinopterus femoralis Say, 1823
Melinopterus prodromus Brahm, 1790Spring small dung beetle
Nipponoserica peregrina Chapin, 1938Picture rockskipper
Onthophagus (Onthophagus) concinnus (Laporte, 1840)⇒ Onthophagus striatulus Palisot de B..., 1809
Onthophagus (Onthophagus) hecate Panzer, 1794Scooped Scarab
Onthophagus hecate hecate Panzer, 1794
Onthophagus (Onthophagus) orpheus Panzer, 1794
Onthophagus orpheus canadensis Fabricius, 1801
Onthophagus orpheus orpheus Panzer, 1794
Onthophagus (Onthophagus) pennsylvanicus Harold, 1871
Onthophagus (Onthophagus) striatulus Palisot de B..., 1809Surf blenny
Onthophagus (Onthophagus) subaeneus (Palisot de B..., 1811)⇒ Onthophagus striatulus Palisot de B..., 1809
Onthophagus (Onthophagus) taurus Schreber, 1759Bull Headed Dung Beetle
Onthophagus (Palaeonthophagus) nuchicornis Linnaeus, 1758Mottled dung beetle
Onthophagus (Palaeonthophagus) ovatus Linnaeus, 1767
Onthophagus brucei cyanopterus (d´Orbigny, 1905)⇒ Proagoderus cyanopterus d´Orbigny, 1905
Onthophagus malthinus Gillet, 1930
Oscarinus rusicola Melsheimer, 1845
Osmoderma eremicola Knoch, 1801Hermit Flower Beetle
Osmoderma rugosa
Osmoderma scabra (Palisot de B..., 1805)⇒ Osmoderma scabrum Palisot de B..., 1805
Osmoderma scabrum Palisot de B..., 1805
Otophorus haemorrhoidalis Linnaeus, 1758Bloody small dung beetle
Oxyomus sylvestris Scopoli, 1763Forest small dung beetle
Pachystethus incolumis
Parastasia brevipes LeConte, 1856
Pardalosus serval Say, 1835
Pelidnota (Pelidnota) punctata Linnaeus, 1758Grapevine beetle
Pelidnota glabra Ohaus, 1922
Phanaeus (Phanaeus) vindex MacLeay, 1819Laguna Madre fiddler
Phanaeus antiquus G. H. Horn, 1876
Phileurus valgus Linnaeus, 1758
Phyllophaga (Phyllophaga) anxia LeConte, 1850Cranberry White Grub
Phyllophaga (Phyllophaga) balia Say, 1825
Phyllophaga (Phyllophaga) corrosa LeConte, 1856
Phyllophaga (Phyllophaga) crassissima Blanchard, 1851
Phyllophaga (Phyllophaga) crenulata Frölich, 1792
Phyllophaga (Phyllophaga) cribrosa LeConte, 1853
Phyllophaga (Phyllophaga) diffinis Blanchard, 1851
Phyllophaga (Phyllophaga) drakii Kirby, 1837
Phyllophaga (Phyllophaga) ephilida Say, 1825Bearded lupine
Phyllophaga ephilida ephilida Say, 1825
Phyllophaga (Phyllophaga) fervida Fabricius, 1775
Phyllophaga (Phyllophaga) forsteri Burmeister, 1855Highfin blenny
Phyllophaga (Phyllophaga) fraterna Harris, 1842Bluespotted blenny
Phyllophaga (Phyllophaga) fusca Frölich, 1792Northern June beetle
Phyllophaga (Phyllophaga) futilis LeConte, 1850Gibbous June beetle
Phyllophaga (Phyllophaga) glaberrima Blanchard, 1851
Phyllophaga (Phyllophaga) gracilis Burmeister, 1855Highfin blenny
Phyllophaga gracilis gracilis
Phyllophaga (Phyllophaga) hirsuta Knoch, 1801
Phyllophaga (Phyllophaga) hirticula Knoch, 1801
Phyllophaga (Phyllophaga) hornii Smith, 1889
Phyllophaga (Phyllophaga) ilicis Knoch, 1801
Phyllophaga (Phyllophaga) implicita Horn, 1887
Phyllophaga (Phyllophaga) inversa Horn, 1887Southern June beetle
Phyllophaga (Phyllophaga) knochii Schönherr an..., 1817
Phyllophaga (Phyllophaga) longispina Smith, 1889
Phyllophaga (Phyllophaga) marginalis LeConte, 1856Sand stargazer
Phyllophaga (Phyllophaga) micans Knoch, 1801
Phyllophaga (Phyllophaga) nitida LeConte, 1856
Phyllophaga (Phyllophaga) panorpa Sanderson, 1950
Phyllophaga (Phyllophaga) quercus Knoch, 1801
Phyllophaga (Phyllophaga) rugosa Melsheimer, 1845Rugose June beetle
Phyllophaga (Phyllophaga) spreta Horn, 1887
Phyllophaga (Phyllophaga) subpruinosa Casey, 1884
Phyllophaga (Phyllophaga) subtonsa LeConte, 1856
Phyllophaga (Phyllophaga) vilifrons LeConte, 1856
Phyllophaga knochi Gyllenhall, 1817
Phyllophaga villifrons LeConte, 1856
Planolinellus vittatus Say, 1825
Pleurophorus caesus Creutzer, 1796Cut small dung beetle
Polyphylla (Polyphylla) hammondi LeConte, 1856Aquatic oligochaete worm
Polyphylla (Polyphylla) variolosa Hentz, 1830Variegated June Beetle
Popillia japonica Newman, 1838Japanese beetle
Psammodius laevipennis Costa, 1844Awkward-winged small dung beetle
Pseudagolius bicolor Say, 1823
Rhyssemus scaber Haldeman, 1848
Scabrostomus tanytarsus Skelley & Wo..., 1991Masked stargazer
Serica acontia Dawson, 1933
Serica atracapilla Kirby, 1837
Serica blatchleyi Dawson, 1932Rippled blenny
Serica georgiana Leng, 1911
Serica georgiana georgiana Leng, 1911
Serica georgiana lecontei Dawson, 1921
Serica iricolor Say, 1824
Serica mystaca Dawson, 1922
Serica parallela Casey, 1884
Serica sericea Illiger, 1802Onespot fringehead
Serica vespertina Gyllenhal, 1817
Stenotothorax badipes Melsheimer, 1845
Strategus aloeus Linnaeus, 1758Coconut cockle
Strategus antaeus Drury, 1773Ox beetle
Strigoderma arboricola Fabricius, 1792
Strigoderma pygmaea Fabricius, 1798Pygmy Chafer
Teuchestes fossor Linnaeus, 1758Digger small dung beetle
Tomarus gibbosus De Geer, 1774Carrot beetle
Tomarus relictus Say, 1825
Trichesthes tristis Fabricius, 1787
Trichiotinus affinis Gory & Perch..., 1833Hairy Flower Scarab
Trichiotinus assimilis Kirby, 1837Bee-Mimic Beetle
Trichiotinus bibens Fabricius, 1775
Trichiotinus piger Fabricius, 1775Bee-like Flower Scarab
Trichonotulus scrofa Fabricius, 1787Sow small dung beetle
Trichopopillia sexmaculata Benderitter, 1923
Trigonopeltastes delta Forster, 1771Delta Flower Scarab
Valgus canaliculatus Olivier, 1789
Valgus seticollis Palisot de B..., 1805Bristly-necked Valgus
Xyloryctes jamaicensis Drury, 1773Eastern Rhinoceros Beetle

Subnodes of Scarabaeidae with occurrences in Pennsylvania
SubfamilyAphodiinae
Leach, 1815
Aphodiine Dung Beetles, Small dung beetles
31
58
Genera
Species

Taxom

SubfamilyCetoniinae
Leach, 1815
Flower beetles, Flower chafers, Flower scarabs, Fruit and flower chafers
8
23
Genera
Species

Taxom

SubfamilyDynastinae
MacLeay, 1819
Horn beetles, Rhinoceros beetles
11
22
Genera
Species

Taxom

SubfamilyMelolonthinae
Samouelle, 1819
May Beetles and Junebugs, White grubs
16
76
Genera
Species

Taxom

SubfamilyRutelinae
MacLeay, 1819
Shining Flower Chafers, Shining Leaf Chafers
10
19
Genera
Species

Taxom

SubfamilyScarabaeinae
Latreille, 1802
Dung Beetles, True dung beetles
8
23
Genera
Species

Taxom



Scarabaeidae (Latreille 1802)
Scarabaeidae (Latreille 1802)


Links, sources and further information
Scarabaeidae at artfakta.se
Scarabaeidae at dyntaxa.se
Scarabaeidae at gbif.org
Scarabaeidae at irmng.org
Scarabaeidae at itis.gov