Checklist for Family Scarabaeidae Latreille, 1802 (Dung beetles)
and region Sabah (Malaysia)
Class
| Insecta
Linnaeus, 1758
Insects, True insects | 345
2906
6852 | Families
Genera
Species |
Taxom
|
Subclass
| Pterygota
Lang, 1888
Winged insects | 345
2906
6852 | Families
Genera
Species |
Taxom
|
Infraclass
| Neoptera
van der Wulp, 1890
Modern Wing-folding insects | 328
2841
6750 | Families
Genera
Species |
Taxom
|
Superorder
| Holometabola
Heider, 1889
Holometabolous Insects | 208
2176
5404 | Families
Genera
Species |
Taxom
|
Order
| Coleoptera
Linnaeus, 1758
Beetles, Coleopterans | 53
889
2843 | Families
Genera
Species |
Taxom
|
Suborder
| Polyphaga
Emery, 1886
Polyphags | 51
871
2825 | Families
Genera
Species |
Taxom
|
Infraorder
| Scarabaeiformia
Crowson, 1960
| 9
185
806 | Families
Genera
Species |
Taxom
|
Superfamily
| Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles | 9
185
806 | Families
Genera
Species |
Taxom
| | Family | Scarabaeidae
Latreille, 1802
Dung beetles, Flower beetles, Fruit chafers, Rain beetles | 131
548 | Genera
Species |
Taxom
|
Genera for Sabah
131 Genera with 548 Species #ABCDEFGHIKLMNOPRSTUVWX
| Acanthovalgus Kraatz, 1895 | | 1 | Species | Taxom | | Adebrattia Bordat & Howden, 1995 | | 1 | Species | Taxom | | Adoretus Laporte, 1932 | | 6 | Species | Taxom | | Agestrata Eschscholtz, 2010 | | 4 | Species | Taxom | | Agrilinus Mulsant & Rey, 1960 | | 1 | Species | Taxom | | Amphimallon Lepeletier &..., 1758 | European Chafers | 1 | Species | Taxom | | Anomala Samouelle, 1910 | Shining Leaf Chafer | 37 | Species | Taxom | | Anoplognathus Leach, 1890 | Christmas beetle | 1 | Species | Taxom | | Anthracophora Burmeister, 1908 | | 2 | Species | Taxom | | Apogonia Melichar, 1904 | | 20 | Species | Taxom | | Asactopholis Brenske, 1913 | | 1 | Species | Taxom | | Ataenius Harold, 1877 | | 3 | Species | Taxom | | Autoserica Brenske, 1915 | Garden beetle | 1 | Species | Taxom | | Blabephorus Fairmaire, 1898 | | 1 | Species | Taxom | | Bombodes Westwood, 1857 | | 1 | Species | Taxom | | Callistethus Blanchard, 0 | | 6 | Species | Taxom | | Campsiura Hope, 1864 | | 1 | Species | Taxom | | Catharsius Hope, 1996 | | 3 | Species | Taxom | | Ceroplophana Gestro, 1996 | | 1 | Species | Taxom | | Chalcosoma Hope, 1900 | | 3 | Species | Taxom | | Chalcothea Burmeister, 1858 | | 1 | Species | Taxom | | Chalcotheomima Miksic, 2020 | | 3 | Species | Taxom | | Chewia | | 1 | Species | Taxom | | Clerota Burmeister, 1917 | | 1 | Species | Taxom | | Clinteria Burmeister, 1911 | | 5 | Species | Taxom | | Clinterocera Motschulsky, 2010 | | 2 | Species | Taxom | | Clyster Arrow, 2002 | | 2 | Species | Taxom | | Coenochilus Burmeister &..., 1883 | | 3 | Species | Taxom | | Coilodera Hope, 1971 | | 3 | Species | Taxom | | Copris Geoffroy, 1921 | Mexican dung beetle | 6 | Species | Taxom | | Cyobius Sharp, 2006 | | 1 | Species | Taxom | | Danielssonia Bordat & Howden, 1995 | | 1 | Species | Taxom | | Dasyvalgus Kolbe, 1904 | | 19 | Species | Taxom | | Dichelomorpha Burmeister, 1893 | | 1 | Species | Taxom | | Dicheros Gory & Perch..., 1985 | | 2 | Species | Taxom | | Dichodontus Burmeister, 1912 | | 3 | Species | Taxom | | Eophileurus Arrow, 1993 | | 2 | Species | Taxom | | Eudicella White, 0 | | 1 | Species | Taxom | | Euselates J. Thomson, 1895 | | 4 | Species | Taxom | | Exopholis Motschoulsky, 1819 | | 2 | Species | Taxom | | Fruhstorferia Willemse, 1984 | | 2 | Species | Taxom | Gametis Burmeister, 1842 | | 0 | Species | Taxom | | Gilletianus Balthasar, 1963 | | 4 | Species | Taxom | | Glycosia Schoch, 1995 | | 2 | Species | Taxom | | Glycyphana Burmeister, 1914 | | 20 | Species | Taxom | | Glyptothea Bates, 1889 | | 2 | Species | Taxom | | Glyptotheomima Miksic, 1932 | | 1 | Species | Taxom | | Gnathocera Kirby, 1836 | | 1 | Species | Taxom | | Gymnetis MacLeay, 1875 | | 1 | Species | Taxom | Gymnopleurus Illiger, 1803 | | 0 | Species | Taxom | | Gyroplia Brenske, 1893 | | 1 | Species | Taxom | | Haroldius Boucomont, 1914 | | 4 | Species | Taxom | | Heterorhina Westwood, 1907 | | 3 | Species | Taxom | | Heterorrhina Westwood, 2010 | | 1 | Species | Taxom | | Heterovalgus Krikken, 1992 | | 1 | Species | Taxom | | Hoplia Illiger, 1893 | Monkey Beetles | 9 | Species | Taxom | | Hoplitovalgus Kolbe, 1904 | | 1 | Species | Taxom | Hybovalgus Kolbe, 1904 | | 0 | Species | Taxom | | Ischiopsopha Gestro, 1860 | | 1 | Species | Taxom | | Ixorida Thomson, 1899 | | 4 | Species | Taxom | | Kheper Janssens, 1843 | | 1 | Species | Taxom | | Latipalpus Moser, 1921 | | 4 | Species | Taxom | | Lepidiota Kirby, 1876 | | 3 | Species | Taxom | | Leucopholis Blanchard, 1892 | | 2 | Species | Taxom | | Liatongus Reitter, 1800 | | 1 | Species | Taxom | | Lorkovitschia Miksic, 1868 | | 2 | Species | Taxom | | Macronota Hoffmansegg, 1909 | | 8 | Species | Taxom | | Maladera Mulsant, 1899 | | 2 | Species | Taxom | | Mecinonota Kraatz, 1801 | | 1 | Species | Taxom | | Megasybacodes | | 1 | Species | Taxom | | Meroloba Thomson, 1913 | | 3 | Species | Taxom | | Microchalcothea Moser, 1910 | | 1 | Species | Taxom | | Microcopris Balthasar, 1787 | | 3 | Species | Taxom | | Microserica Brenske, 1915 | | 15 | Species | Taxom | | Microvalgus Kraatz, 1904 | | 2 | Species | Taxom | | Mimela Kirby, 2001 | | 8 | Species | Taxom | | Nematophylla Arrow, 1938 | | 1 | Species | Taxom | | Neoserica Brenske, 1899 | | 7 | Species | Taxom | | Neotrichiorhyssemus RakoviÄ & K..., 1901 | | 1 | Species | Taxom | | Neovalgus Miyake, 1989 | | 1 | Species | Taxom | | Ochicanthon Vaz-de-Mello, 2007 | | 11 | Species | Taxom | | Octoplasia Brenske, 1906 | | 3 | Species | Taxom | | Odochilus Harold, 2017 | | 3 | Species | Taxom | | Oniticellus Serville, 1879 | | 1 | Species | Taxom | | Onitis Fabricius, 1931 | | 1 | Species | Taxom | | Onthophagus Latreille, 2006 | | 59 | Species | Taxom | | Oreovalgus Kolbe, 1921 | | 1 | Species | Taxom | | Oryctes Hellwig, 1758 | Coconut rhinoceros beetles | 1 | Species | Taxom | | Oxyomus Dejean, 2016 | | 1 | Species | Taxom | | Panelus Lewis, 2016 | | 5 | Species | Taxom | | Paragymnopleurus Shipp, 1875 | | 4 | Species | Taxom | | Paraphytus Harold, 1880 | | 1 | Species | Taxom | | Paraprotaetia Moser, 1996 | | 3 | Species | Taxom | | Pararhabdotis Kraatz, 1897 | | 1 | Species | Taxom | | Parascatonomus Paulian, 2008 | | 20 | Species | Taxom | | Parastasia Westwood, 2008 | | 18 | Species | Taxom | | Peltonotus Burmeister, 1910 | | 7 | Species | Taxom | | Pentelia Kiriakoff, 1893 | | 3 | Species | Taxom | | Pharaphodius Reitter, 1781 | | 2 | Species | Taxom | | Philistina MacLeay, 1903 | | 2 | Species | Taxom | | Planolinoides Dellacasa an..., 1987 | | 1 | Species | Taxom | | Platysodes Westwood, 1874 | | 1 | Species | Taxom | | Plectrone Wallace, 1883 | | 2 | Species | Taxom | | Pleuraphodius Schmidt, 1875 | | 1 | Species | Taxom | | Pleuronota Kraatz, 2000 | | 5 | Species | Taxom | | Popillia Leach, 1838 | | 2 | Species | Taxom | | Proagoderus Lansberge, 2002 | | 2 | Species | Taxom | | Protaetia Burmeister, 1775 | | 13 | Species | Taxom | | Pseudochalcothea Ritsema, 1889 | | 9 | Species | Taxom | | Pseudochalcotheomima Miksic, 2010 | | 3 | Species | Taxom | | Pseudohomonyx Arrow, 1997 | | 3 | Species | Taxom | | Pseudostereomera Bordat & Howden, 1995 | | 1 | Species | Taxom | | Psilopholis Brenske, 1881 | | 3 | Species | Taxom | | Rhyparus Westwood, 1883 | | 14 | Species | Taxom | | Rutelarcha Waterhouse, 1874 | | 1 | Species | Taxom | | Saprosites Redtenbacher, 1909 | | 12 | Species | Taxom | | Sisyphus Latreille, 0 | | 1 | Species | Taxom | | Spinanomala Ohaus, 1910 | | 2 | Species | Taxom | Svobia Miksic, 1965 | | 0 | Species | Taxom | | Sybacodes Fairmaire, 2018 | | 1 | Species | Taxom | | Synapsis Forster, 1874 | | 1 | Species | Taxom | | Taeniodera Burmeister, 2004 | | 20 | Species | Taxom | | Termitodiellus Nakane, 1983 | | 1 | Species | Taxom | | Thaumastopeus Kraatz, 1995 | | 1 | Species | Taxom | | Theodosia J. Thomson, 1880 | | 17 | Species | Taxom | | Trichogomphus Burmeister, 1907 | | 4 | Species | Taxom | | Uenoeus Ochi & Kon, 2009 | | 1 | Species | Taxom | | Valgus Scriba, 1805 | | 1 | Species | Taxom | | Wadaia | | 1 | Species | Taxom | | Xenoloba Bates, 1889 | | 4 | Species | Taxom | | Xylotrupes Hope, 2004 | Elephant beetles | 5 | Species | Taxom |
Species list for Sabah
548 Species and 48 Subspecies | Acanthovalgus marquardi Kraatz, 1895 | | | Adebrattia depressa Bordat & Howden, 1995 | | | Adoretus (Adoretus) sandakanus Ohaus, 1935 | | | Adoretus (Adoretus) speculator Ohaus, 1923 | | | Adoretus borneensis Kraatz, 1895 | | | Adoretus compressus Weber, 1801 | Brown cockchafer | | Adoretus furutai Kobayashi, 2007 | | | Adoretus klossi Ohaus, 1932 | | | Agestrata belitungana Miksic, 1973 | | | Agestrata dehaan Gory & Perch..., 1833 | | | Agestrata parryi Wallace, 1867 | | | Agestrata philipchewi Legrand & Ch..., 2010 | | | Agrilinus hasegawai aka Nakane, 1960 | | | Amphimallon (Amphimallon) solstitiale Linnaeus, 1758 | Cliff Beetle | | Anomala acromialis Ohaus, 1916 | | | Anomala aelia Ohaus, 1932 | | | Anomala ahlwarthi Ohaus, 1916 | | | Anomala bicolor (Fabricius, 1775) | ⇒ Anomala insularis Castelnau, 1840 | | Anomala chalcophysa Ohaus, 1916 | | | Anomala cochlearia Ohaus, 1932 | | | Anomala denticrus Ohaus, 1910 | | | Anomala diversicolor Ohaus, 1916 | | | Anomala eksae Prokofiev & ..., 2015 | | | Anomala fissula Ohaus, 1916 | | | Anomala fulvocalceata Ohaus, 1916 | | | Anomala fulvofusca Ohaus, 1915 | | | Anomala fuscula Sharp, 1881 | | | Anomala gordiana Ohaus, 1932 | | | Anomala heterostigma Ohaus, 1915 | | | Anomala kinabalensis Ohaus, 1910 | | | Anomala kudatina Ohaus, 1916 | | | Anomala latefemorata Ohaus, 1910 | | | Anomala matricula Ohaus, 1916 | | | Anomala metella Ohaus, 1923 | | | Anomala pallida Fabricius, 1775 | Cockchafer | | Anomala parastasioides Zorn, 2007 | | | Anomala prisca Ohaus, 1915 | | | Anomala prolixa Arrow, 1910 | | | Anomala pumilis Ohaus, 1932 | | | Anomala pumiloides Zorn, 2007 | | | Anomala quadrigemina Arrow, 1910 | | | Anomala rotundiceps Sharp, 1881 | | | Anomala sabana Ohaus, 1916 | | | Anomala silama Arrow, 1910 | | | Anomala snizeki Zorn, 2007 | | | Anomala soror Arrow, 1910 | | | Anomala teretina Ohaus, 1932 | | | Anomala uncata Zorn, 2005 | | | Anomala vorstmanni Heller, 1891 | | | Anomala waterstraati Ohaus, 1916 | | | Anomala whiteheadi (Arrow, 1910) | ⇒ Anomala albaya Arrow, 1915 | | Anoplognathus punctulatus Olliff, 1890 | Leaf green Christmas beetle | | Anthracophora borneensis Kraatz, 1898 | | | Anthracophora gigantea Moser, 1908 | | | Apogonia brunneipennis Moser, 1913 | | | Apogonia carinata Kobayashi, 2013 | | | Apogonia elongata Arrow, 1938 | | | Apogonia helleri Ritsema, 1904 | | | Apogonia heynei Moser, 1918 | | | Apogonia impressifrons Moser, 1913 | | | Apogonia insignis Brenske, 1893 | | | Apogonia irrorata Arrow, 1938 | | | Apogonia montana Moser, 1915 | | | Apogonia palawana Heller, 1896 | | | Apogonia perdix Arrow, 1938 | | | Apogonia producta Arrow, 1938 | | | Apogonia robusta Kobayashi, 2013 | | | Apogonia rudepunctata Moser, 1924 | | | Apogonia sabahensis Kobayashi, 2013 | | | Apogonia sandakana Moser, 1921 | | | Apogonia speculifera Ritsema, 1904 | | | Apogonia tortuosa Kobayashi, 2017 | | | Apogonia tuberculiventris Ritsema, 1897 | | | Apogonia waterstradti Ritsema, 1904 | | | Asactopholis sericea Moser, 1913 | | | Ataenius australasiae Boheman, 1858 | | | Ataenius borneensis Minkina & Be..., 2023 | | | Ataenius peregrinator Harold, 1877 | Birthworts | | Autoserica monticola Moser, 1915 | | | Blabephorus pinguis Fairmaire, 1898 | | | Bombodes westwoodi Thomson, 1857 | | | Callistethus dechambrei Wada, 1998 | | | Callistethus hauschildti Ohaus, 1903 | | | Callistethus mindanaoensis Wada, 2002 | | | Callistethus nigrosparsus Kobayashi & ..., 2021 | | | Callistethus stolidopygus Ohaus, 1915 | | | Callistethus waterstraati waterstraati | | | Campsiura flavoguttata Snellen van ..., 1864 | | | Catharsius dayacus Lansberge, 1886 | | | Catharsius molossus Linnaeus, 1758 | | | Catharsius renaudpauliani Ochi & Kon, 1996 | | | Ceroplophana malaysiana nobuyukii Nagai, 1996 | | | Chalcosoma atlas Linnaeus, 1758 | Atlas beetle | | Chalcosoma atlas keyboh Nagai, 2004 | | | Chalcosoma moellenkampi Kolbe, 1900 | Moellenkampi beetle | | Chalcothea affinis Snellen van ..., 1858 | | | Chalcotheomima laevis Arrow, 1932 | | | Chalcotheomima macrophylla Janson, 1912 | | | Chalcotheomima sakaii Kawahara, To..., 2020 | | | Chewia miniata Legrand, 2004 | | | Clerota rigifica Janson, 1917 | | | Clinteria atra Wiedemann, 1823 | | | Clinteria atra egens Snellen van ..., 1864 | | | Clinteria flavonotata nayani Legrand & Chew, 2004 | | | Clinteria jimlii Legrand & Ch..., 2010 | | | Clinteria moultoni Moser, 1911 | | | Clinterocera borneensis Krikken, 1982 | | | Clinterocera cyprilrusi Legrand & Ch..., 2010 | | | Clyster itys Olivier, 1789 | | | Clyster trinodosus Dechambre, 2002 | | | Coenochilus legrandi Krajcik, 2013 | | | Coenochilus leopoldi Bourgoin, 1931 | | | Coenochilus sumatranus Westwood, 1883 | | | Coilodera helleri Miksic, 1972 | | | Coilodera kalimantanica Miksic, 1972 | | | Coilodera pseudoalveata Miksic, 1971 | | | Copris agnus Sharp, 1875 | | | Copris gibbulus borneensis Ochi & Kon, 2005 | | | Copris hidakai Ochi & Kon, 1996 | | | Copris numa Lansberge, 1886 | Braken Bat Cave meshweaver | | Copris poggii Ochi & Kon, 2005 | | | Copris ramosiceps Gillet, 1921 | | | Cyobius cheyi Ochi, Kon & ..., 2006 | | | Danielssonia minuta Bordat & Howden, 1995 | | | Dasyvalgus bimaculatus Kraatz, 1896 | | | Dasyvalgus eucharis Kolbe, 1904 | | | Dasyvalgus flavomaculatus Miyake & Yamaya, 1993 | | | Dasyvalgus hauseri Ricchiardi, 2013 | | | Dasyvalgus inarcuatus Ricchiardi, 1995 | | | Dasyvalgus infuscatus Kolbe, 1904 | | | Dasyvalgus monachus monachus | | | Dasyvalgus pusillum Ricchiardi, 2013 | | | Dasyvalgus pusillus Ricchiardi, 2013 | | | Dasyvalgus pusio Kolbe, 1904 | | | Dasyvalgus pyrrhopygus Kraatz, 1883 | | | Dasyvalgus rollei Kolbe, 1904 | | | Dasyvalgus sabahi Ricchiardi, 2013 | | | Dasyvalgus spineus Miyake & Yamaya, 1993 | | | Dasyvalgus subnitidus Kolbe, 1904 | | | Dasyvalgus testaceus Kraatz, 1896 | | | Dasyvalgus trusmadii Ricchiardi, 2013 | | | Dasyvalgus vethi Ritsema, 1879 | | | Dasyvalgus waterstradti Kolbe, 1904 | | | Dichelomorpha nigra Brenske, 1893 | | | Dicheros bicornis Latreille, 1817 | | | Dicheros bicornis borneensis Nagai, 1985 | | | Dichodontus coronatus Burmeister, 1847 | | | Dichodontus grandis Ritsema, 1882 | | | Dichodontus pulchellus Prell, 1912 | | | Eophileurus borneensis Endrödi, 1977 | | | Eophileurus multidentatus Miyake & Yamaya, 1993 | | | Eudicella euthalia euthalia | | | Euselates cineracea Gory & Perch..., 1833 | | | Euselates conspersa Schoch, 1897 | | | Euselates quadrilineata Hope, 1831 | | | Euselates trifasciata bifasciata Kraatz, 1895 | | | Exopholis borneensis Brenske, 1893 | | | Exopholis hypoleuca Wiedemann, 1819 | | | Fruhstorferia flavipennis Nagai, 1984 | | | Fruhstorferia nigromuliebris Nagai, 1984 | | | Gilletianus miurai Minkina, 2022 | | | Gilletianus reichei Harold, 1859 | | | Gilletianus sundainsulanus Ochi, Kon & ..., 2014 | | | Gilletianus therondi Balthasar, 1963 | | | Glycosia alboguttata alboguttata | | | Glycosia borneensis Sakai, 1995 | | | Glycyphana binotata binotata | | | Glycyphana chewi Antoine & Le..., 2002 | | | Glycyphana cincta Wallace, 1867 | | | Glycyphana fadilae Miksic, 1967 | | | Glycyphana festiva Fabricius, 1792 | | | Glycyphana fruhstorferi Schoch, 1897 | | | Glycyphana illusa Janson, 1881 | | | Glycyphana inusta Mohnike, 1871 | | | Glycyphana inusta inusta | | | Glycyphana leucogastra Miksic, 1967 | | | Glycyphana macquarti sumatrensis Snellen van ..., 1864 | | | Glycyphana modesta Fabricius, 1792 | | | Glycyphana nicobarica Janson, 1877 | | | Glycyphana nigricollis Moser, 1908 | | | Glycyphana parvula Moser, 1914 | | | Glycyphana penanga Wallace, 1867 | | | Glycyphana pygmaea Mohnike, 1871 | | | Glycyphana quadricolor sinuata Wallace, 1867 | | | Glycyphana steveni Antoine & Le..., 2002 | | | Glycyphana tonkinensis Moser, 1914 | | | Glyptothea excavata Janson, 1912 | | | Glyptothea whiteheadi Bates, 1889 | | | Glyptotheomima pubescens Arrow, 1932 | | | Gnathocera petelli (Buquet, 1836) | ⇒ Dicheros bicornis Latreille, 1817 | | Gymnetis goryi Janson, 1875 | | | Gyroplia bimaculata Brenske, 1893 | | | Haroldius borneensis Paulian, 1993 | | | Haroldius discoidalis Paulian, 1993 | | | Haroldius pauliani Scheuern, 1995 | | | Haroldius rugatulus Boucomont, 1914 | | | Heterorhina borneensis borneensis | | | Heterorhina rahimi Legrand & Ch..., 2010 | | | Heterorhina simula Janson, 1907 | | | Heterorrhina rahimi Legrand & Ch..., 2010 | | | Heterovalgus antoinei Ricchiardi, 1992 | | | Hoplia aurata Waterhouse, 1877 | | | Hoplia aurifera Brenske, 1893 | | | Hoplia auromicans Brenske, 1893 | | | Hoplia fissipes Moser, 1912 | | | Hoplia imparilis Moser, 1912 | | | Hoplia magnifica Arrow, 1932 | | | Hoplia setifera Moser, 1912 | | | Hoplia setosella Brenske, 1893 | | | Hoplia ventricosa Brenske, 1893 | | | Hoplitovalgus fallaciosus Kolbe, 1904 | | | Ischiopsopha olivacea (Thomson, 1860) | ⇒ Ischiopsopha jamesi Waterhouse, 1876 | | Ixorida gordoni Miksic, 1972 | | | Ixorida kinabaluensis Antoine, 1986 | | | Ixorida ornata borneensis Antoine, 1986 | | | Ixorida regia bicolor Kraatz, 1899 | | | Kheper prodigiosus Erichson, 1843 | | | Latipalpus elegans Itoh, 1999 | | | Latipalpus fujiokai Itoh, 1999 | | | Latipalpus maxillatus Brenske, 1892 | | | Latipalpus palpalis Moser, 1921 | | | Lepidiota flavimargo Arrow, 1919 | | | Lepidiota marginipennis Moser, 1913 | | | Lepidiota munda Sharp, 1876 | | | Leucopholis deplanata Moser, 1908 | | | Leucopholis staudingeri Brenske, 1892 | | | Liatongus femoratus Illiger, 1800 | | | Lorkovitschia douglasprimusensis Legrand & Ch..., 2004 | | | Lorkovitschia rustica Wallace, 1868 | | | Macronota agni Wallace, 1867 | | | Macronota chicheryi tenuigenius Miyake & Yamaya, 1995 | | | Macronota higginsi Janson, 1874 | | | Macronota miyashitai Alexis & Del..., 2001 | | | Macronota nigritus nigritus | | | Macronota striatus striatus | | | Macronota westwoodi Ritsema, 1893 | | | Macronota westwoodi borneensis Moser, 1909 | | | Maladera drescheri Moser, 1913 | | | Maladera kinabaluensis Brenske, 1899 | | | Mecinonota regia Fabricius, 1801 | | | Megasybacodes brevitarsis Kakizoe, Mar..., 2019 | | | Meroloba suturalis Snellen van ..., 1858 | | | Meroloba suturalis suturalis | | | Meroloba trivittata Moser, 1913 | | | Microchalcothea borneensis Moser, 1910 | | | Microcopris fujiokai poringensis Ochi & Kon, 2005 | | | Microcopris hidakai Ochi & Kon, 1996 | | | Microcopris reflexus Fabricius, 1787 | | | Microserica fascigera Moser, 1911 | | | Microserica flaveola Moser, 1911 | | | Microserica lineatipennis Moser, 1921 | | | Microserica lugens Moser, 1915 | | | Microserica magnifica Brenske, 1899 | | | Microserica minuscula Moser, 1915 | | | Microserica moultoni Moser, 1911 | | | Microserica nigriceps Arrow, 1946 | | | Microserica nigrolineata Moser, 1911 | | | Microserica nigrovittata Moser, 1911 | | | Microserica quinquelamellata Moser, 1920 | | | Microserica sandakana Moser, 1921 | | | Microserica strigata Brenske, 1899 | | | Microserica strigosa Brenske, 1899 | | | Microserica virgata Moser, 1915 | | | Microvalgus micros Kolbe, 1904 | | | Microvalgus parcus Kolbe, 1904 | | | Mimela insularis Ohaus, 1908 | | | Mimela maculicollis (Ohaus, 1908) | ⇒ Mimela junii rugosula Fairmaire, 1859 | | Mimela margarita Arrow, 1910 | | | Mimela piceoviridana Miyake, 1994 | | | Mimela prilca Ohaus, 1936 | | | Mimela suspecta Miyake, 1994 | | | Mimela uenoi Wada, 2001 | | | Mimela zorni Wada, 2001 | | | Nematophylla carinicollis Arrow, 1938 | | | Neoserica borneensis Brenske, 1899 | | | Neoserica curticrus Moser, 1915 | | | Neoserica fischeri Moser, 1911 | | | Neoserica montana Moser, 1915 | | | Neoserica nigrescens Moser, 1915 | | | Neoserica ovata Moser, 1915 | | | Neoserica squalida Brenske, 1899 | | | Neotrichiorhyssemus hirsutus Clouët, 1901 | | | Neovalgus scarletus Miyake, 1989 | | | Ochicanthon crockermontis Krikken & Hu..., 2007 | | | Ochicanthon danum Krikken & Hu..., 2007 | | | Ochicanthon dytiscoides Boucomont, 1914 | | | Ochicanthon gangkui Ochi & Kon &..., 1997 | | | Ochicanthon kikutai Ochi, Ueda &..., 2006 | | | Ochicanthon kimanis Krikken & Hu..., 2007 | | | Ochicanthon maryatiae Ochi, Ueda &..., 2006 | | | Ochicanthon masumotoi Ochi & Araya, 1996 | | | Ochicanthon parantisae Ochi & Kon &..., 1997 | | | Ochicanthon rombauti Krikken & Hu..., 2007 | | | Ochicanthon tambunan Krikken & Hu..., 2007 | | | Octoplasia castaneipennis Matsumoto, 2014 | | | Octoplasia princeps Sharp, 1876 | | | Octoplasia prolixa Arrow, 1906 | | | Odochilus bacchusi Rakovic, 1987 | | | Odochilus jani Bezdek & Krivan, 2001 | | | Odochilus kashizakii Ochi, Kon & ..., 2017 | | | Oniticellus tessellatus Harold, 1879 | | | Onitis subopacus Arrow, 1931 | | | Onthophagus (Gibbonthophagus) fujiii Ochi & Kon, 1995 | | | Onthophagus (Gibbonthophagus) kimioi Ochi, Kon & ..., 2011 | | | Onthophagus (Gibbonthophagus) limbatus Herbst, 1789 | | | Onthophagus (Gibbonthophagus) parviobscurior Ochi, Kon & ..., 2009 | | | Onthophagus (Indachorius) danumensis (Ochi, Kon & ..., 2009) | ⇒ Onthophagus danumicus Huijbregts &..., 2012 | | Onthophagus (Indachorius) masaoi Ochi, 1992 | | | Onthophagus (Indachorius) paramasaoi Ochi, Kon & ..., 2009 | | | Onthophagus (Indachorius) semidanumensis Ochi, Kon & ..., 2009 | | | Onthophagus (Matashia) anguliceps (Boucomont, 1914) | ⇒ Onthophagus avocetta Arrow, 1933 | | Onthophagus (Onthophagus) angustatus Boucomont, 1914 | | | Onthophagus (Onthophagus) ishiii Ochi & Kon, 1995 | | | Onthophagus (Onthophagus) mulleri van Lansberge, 1883 | | | Onthophagus (Palaeonthophagus) olsoufieffi Boucomont, 1924 | | | Onthophagus (Paronthophagus) hidakai Ochi & Kon, 1995 | | | Onthophagus (Phanaeomorphus) johkii Ochi & Kon, 1994 | | | Onthophagus laevis laevis | | | Onthophagus (Serrophorus) sagittarius Fabricius, 1775 | Sagittarius dung beetle | | Onthophagus aphodioides Lansberge, 1883 | | | Onthophagus arayai Ochi & Kon, 2007 | | | Onthophagus azusae Ochi & Kon, 2006 | | | Onthophagus bangueyensis Boucomont, 1914 | | | Onthophagus borneensis Harold, 1877 | | | Onthophagus cervicapra Boucomont, 1914 | | | Onthophagus cheyi Ochi & Kon, 2006 | | | Onthophagus clivimerus Huijbregts &..., 2011 | | | Onthophagus cupreopastillatus Ochi & Kon, 2006 | | | Onthophagus danumicus Huijbregts &..., 2012 | | | Onthophagus foedus Boucomont, 1914 | | | Onthophagus hikidai Ochi & Kon, 2006 | | | Onthophagus hiroyukii Ochi, 2007 | | | Onthophagus insulindicus Boucomont, 1914 | | | Onthophagus kashizakii Kon & Ochi, 2005 | | | Onthophagus koechlei Masumoto, Oc..., 2011 | | | Onthophagus koni Ochi, 2007 | | | Onthophagus liwagensis Ochi & Kon, 2006 | | | Onthophagus magnioculus Ochi & Kon, 2006 | | | Onthophagus maryatiae Ochi & Kon, 2005 | | | Onthophagus matsuii Ochi & Kon, 2006 | | | Onthophagus megapacificus Ochi & Kon, 2006 | | | Onthophagus obscurior Boucomont, 1914 | | | Onthophagus ochromerus Harold, 1877 | | | Onthophagus opacihartiniae Ochi & Kon, 2015 | | | Onthophagus otai Ochi & Kon, 2006 | | | Onthophagus pacificus Lansberge, 1885 | | | Onthophagus pastillatus Boucomont, 1919 | | | Onthophagus phillippsorum Krikken & Hu..., 1987 | | | Onthophagus quasijohkii Ochi & Kon, 2005 | | | Onthophagus quasitagal Ochi & Kon, 2005 | | | Onthophagus robertopoggii Ochi & Kon, 2006 | | | Onthophagus rutilans aborneensis Ochi, Kon & ..., 2009 | | | Onthophagus sabahensis Ochi & Kon, 2006 | | | Onthophagus sepilokensis Ochi & Kon, 2006 | | | Onthophagus sideki Krikken & Hu..., 1987 | | | Onthophagus subcornutus Boucomont, 1914 | | | Onthophagus taeniatus Boucomont, 1914 | | | Onthophagus variolaris Lansberge, 1883 | | | Onthophagus vulpes Harold, 1877 | | | Onthophagus waterstradti Boucomont, 1914 | | | Onthophagus yumotoi Ochi & Kon, 2006 | | | Oreovalgus sandakanus Moser, 1921 | | | Oryctes rhinoceros Linnaeus, 1758 | Asiatic rhinoceros beetle | | Oxyomus mencli Minkina, 2016 | | | Panelus bakeri Boucomont, 1921 | | | Panelus borneensis Paulian, 1992 | | | Panelus danumensis Ochi, Kon & ..., 2009 | | | Panelus sabahi Paulian, 1992 | | | Panelus saigusai Ochi & Kon, 2016 | | | Paragymnopleurus sparsus Sharp, 1875 | | | Paragymnopleurus sparsus sparsus | | | Paragymnopleurus spinotus Boucomont, 1914 | | | Paragymnopleurus striatus Sharp, 1875 | | | Paraphytus ritsemae Harold, 1880 | | | Paraprotaetia borneana Schoch, 1896 | | | Paraprotaetia fulva Schoch, 1896 | | | Paraprotaetia nielseni Antoine, 1996 | | | Pararhabdotis setigera Schoch, 1897 | | | Parascatonomus andrewdavisi Huijbregts &..., 2009 | | | Parascatonomus anitidus Ochi & Kon, 2005 | | | Parascatonomus aurifex Harold, 1877 | | | Parascatonomus brendelli Ochi, Kon & ..., 2008 | | | Parascatonomus bundutuhanensis Ochi, Kon & ..., 2008 | | | Parascatonomus danumcupreus Krikken & Hu..., 2009 | | | Parascatonomus dux Sharp, 1875 | | | Parascatonomus fujiokai Ochi & Araya, 1996 | | | Parascatonomus gunsalami Ochi & Kon, 2005 | | | Parascatonomus kikutai Ochi & Kon, 2005 | | | Parascatonomus liewi Ochi & Kon, 2005 | | | Parascatonomus monticupreus Krikken & Hu..., 2009 | | | Parascatonomus penicillatus Harold, 1879 | | | Parascatonomus poringensis Ochi & Kon, 2005 | | | Parascatonomus rudis Sharp, 1875 | | | Parascatonomus sarawacus Harold, 1877 | | | Parascatonomus sayapensis Ochi & Kon, 2005 | | | Parascatonomus semiaureus Lansberge, 1883 | | | Parascatonomus semicupreus Harold, 1877 | | | Parascatonomus taichii Ochi, Kon & ..., 2008 | | | Parastasia anomala Arrow, 1899 | | | Parastasia asahi Wada, 2008 | | | Parastasia bigibbosa Nonfried, 1891 | | | Parastasia burmeisteri Ohaus, 1898 | | | Parastasia confluens Westwood, 1841 | | | Parastasia discolor discolor | | | Parastasia diversipennis Ohaus, 1911 | | | Parastasia ephippium Snellen, 1864 | | | Parastasia glottidion Kuijten, 1992 | | | Parastasia kinibalensis Ohaus, 1901 | | | Parastasia kuijteni Wada, 1996 | | | Parastasia moseri Ohaus, 1903 | | | Parastasia pseudorufonigra Wada, 2013 | | | Parastasia rufonigra Ohaus, 1911 | | | Parastasia takahikoi Wada, 2007 | | | Parastasia tenomensis Wada, 2008 | | | Parastasia westwoodi Westwood, 1841 | | | Parastasia yasutoshii Wada, 2008 | | | Peltonotus brunnipennis Benderitter, 1934 | | | Peltonotus fujiokai Jameson & Wada, 2004 | | | Peltonotus mushiyaus Jameson & Wada, 2009 | | | Peltonotus nethis Jameson & Wada, 2004 | | | Peltonotus rubripennis Miyake & Yamaya, 1994 | | | Peltonotus similis Arrow, 1931 | | | Peltonotus vittatus Arrow, 1910 | | | Pentelia densa Arrow, 1944 | | | Pentelia kinabaluensis Brenske, 1896 | | | Pentelia serrulata Brenske, 1893 | | | Pharaphodius diadema Wiedemann, 1823 | | | Pharaphodius marginellus Fabricius, 1781 | | | Philistina bicoronatus Jordan, 1894 | | | Philistina inermis Janson, 1903 | | | Planolinoides aenictus Cooper & Gordon, 1987 | Benttooths | | Platysodes verlorenii Westwood, 1874 | | | Plectrone borneensis borneensis | | | Plectrone lugubris Janson, 1883 | | | Pleuraphodius lewisi Waterhouse, 1875 | | | Pleuronota aberrans Antoine, 2000 | | | Pleuronota allardi Antoine, 1990 | | | Pleuronota elongata Gory & Perch..., 1833 | | | Pleuronota pseudoelongata Legrand & Ch..., 2010 | | | Pleuronota sakaii Antoine, 2000 | | | Popillia dajaka Ohaus, 1923 | | | Popillia sandyx Newman, 1838 | | | Proagoderus schwaneri Snellen van ..., 1864 | | | Proagoderus watanabei Ochi & Kon, 2002 | | | Protaetia (Caloprotaetia) chewi Legrand, 2004 | | | Protaetia (Macroprotaetia) distincta Antoine & Pa..., 1994 | | | Protaetia (Macroprotaetia) gestroi Miksic, 1970 | | | Protaetia (Macroprotaetia) milani Antoine & Pa..., 1994 | | | Protaetia (Oxyperatex) spectabilis Schaum, 1841 | | | Protaetia spectabilis velutina Moser, 1907 | | | Protaetia spectabilis viridiopaca Moser, 1907 | | | Protaetia (Pachyprotaetia) ciliata Olivier, 1785 | | | Protaetia (Pachyprotaetia) medvedevi Miksic, 1965 | | | Protaetia (Pachyprotaetia) mixta Weber, 1801 | | | Protaetia (Pachyprotaetia) peterjacki Legrand & Ch..., 2010 | | | Protaetia (Protaetia) prevosti Legrand & Ch..., 2010 | | | Protaetia acuminata Fabricius, 1775 | | | Pseudochalcothea auripes Westwood, 1874 | | | Pseudochalcothea nagaii Sakai, 1993 | | | Pseudochalcothea planiuscula Bates, 1889 | | | Pseudochalcothea planiuscula planiuscula | | | Pseudochalcothea pomacea Bates, 1889 | | | Pseudochalcothea pomacea pomacea | | | Pseudochalcothea spathulifera Bates, 1889 | | | Pseudochalcothea staudingeri v. d. Poll, 1893 | | | Pseudochalcothea viridipes Bates, 1889 | | | Pseudochalcotheomima allardi Antoine, 1990 | | | Pseudochalcotheomima compacta Janson, 1912 | | | Pseudochalcotheomima sammananni Legrand & Ch..., 2010 | | | Pseudohomonyx borneensis Arrow, 1908 | | | Pseudohomonyx crassus Arrow, 1911 | | | Pseudohomonyx extradentatus Miyake & Yamaya, 1997 | | | Pseudostereomera mirabilis Bordat & Howden, 1995 | | | Psilopholis gigantea Matsumoto, 2010 | | | Psilopholis grandis Laporte, 1840 | | | Psilopholis vestita Sharp, 1881 | | | Rhyparus accessoricostatus Mencl, Rakov..., 2013 | | | Rhyparus adebratti Bordat, 1996 | | | Rhyparus approximans Fairmaire, 1893 | | | Rhyparus burckhardti Paulian, 1989 | | | Rhyparus danielssoni Bordat, 1996 | | | Rhyparus denticollis Fairmaire, 1893 | | | Rhyparus helophoroides Fairmaire, 1893 | | | Rhyparus kinabalu Paulian, 1989 | | | Rhyparus magnus Schmidt, 1911 | | | Rhyparus micros Bordat, 1996 | | | Rhyparus minor Paulian, 1989 | | | Rhyparus peninsularis Arrow, 1905 | | | Rhyparus pseudominor Bordat, 1996 | | | Rhyparus xanthi Frivaldszky, 1883 | | | Rutelarcha quadrimaculata Waterhouse, 1874 | | | Saprosites celebicus Paulian, 1944 | | | Saprosites crockerensis Stebnicka, 1991 | | | Saprosites difficilis Harold, 1877 | | | Saprosites japonicus Waterhouse, 1875 | | | Saprosites kinabalu Pittino, 2008 | | | Saprosites laeviceps Harold, 1877 | | | Saprosites laticollis Pittino, 2008 | | | Saprosites marchionalis (Harold, 1877) | ⇒ Saprosites cycloporum Paulian, 1944 | | Saprosites raffrayi Paulian, 1944 | | | Saprosites rufopolitus Gillet, 1924 | | | Saprosites titschacki Balthasar, 1941 | | | Saprosites verecundus Schmidt, 1909 | | | Sisyphus thoracicus thoracicus | | | Spinanomala bispinosa Ohaus, 1910 | | | Spinanomala unispinosa Ohaus, 1910 | | | Sybacodes borneensis Minkina, Men..., 2018 | | | Synapsis ritsemae Lansberge, 1874 | | | Taeniodera aurantiaca Snellen van ..., 1858 | | | Taeniodera aurantiaca aurantiaca | | | Taeniodera bisignata Moser, 1913 | | | Taeniodera borneensis borneensis | | | Taeniodera cervina Wallace, 1867 | | | Taeniodera chewi Legrand, 2000 | | | Taeniodera cordata Schoch, 1897 | | | Taeniodera corticalis corticalis | | | Taeniodera fenestrata Arrow, 1946 | | | Taeniodera inermis Krikken, 1982 | | | Taeniodera jirouxi Legrand, 2000 | | | Taeniodera kinabaluana Arrow, 1932 | | | Taeniodera krikkeni Legrand & Ch..., 2010 | | | Taeniodera marmorata Wallace, 1867 | | | Taeniodera modesta modesta | | | Taeniodera monacha Gory & Perch..., 1833 | | | Taeniodera oudemansi Valck Lucassen, 1932 | | | Taeniodera rafflesiana Westwood, 1842 | | | Taeniodera sakaii Antoine, 2000 | | | Taeniodera trusmadiana Legrand, 2004 | | | Termitodiellus besucheti Paulian, 1983 | | | Thaumastopeus chicheryi tenuigenius Miyake & Yamaya, 1995 | | | Theodosia antoinei Nagai, 1998 | | | Theodosia chewi Ochi, 1993 | | | Theodosia dinoni Legrand & Ch..., 2010 | | | Theodosia howitti Castelnau, 1873 | | | Theodosia katsurai Sakai, 1994 | | | Theodosia magnifica Rotschild & ..., 1893 | | | Theodosia magnifica magnifica | | | Theodosia miyashitai Sakai, 1997 | | | Theodosia nagai Legrand & Ch..., 2010 | | | Theodosia nobuyukii dinoni Legrand & Ch..., 2010 | | | Theodosia nobuyukii nobuyukii | | | Theodosia normanwongi Legrand & Bo..., 2015 | | | Theodosia perakensis tanggilaon Nagai, 1998 | | | Theodosia rothschildi (Janson, 1903) | ⇒ Theodosia telifer Bates, 1889 | | Theodosia telifer Bates, 1889 | | | Theodosia viridiaurata Bates, 1889 | | | Theodosia westwoodi (Thomson, 1880) | ⇒ Theodosia magnifica Rotschild & ..., 1893 | | Trichogomphus bronchus Herbst, 1785 | | | Trichogomphus lunicollis Burmeister, 1847 | | | Trichogomphus simson simson | | | Trichogomphus zangi Sternberg, 1907 | | | Uenoeus andrewdavisi Huijbregts &..., 2009 | | | Valgus seticollis Palisot de B..., 1805 | Bristly-necked Valgus | | Wadaia kaorui Itoh, 1993 | | | Xenoloba elegans Sakai, 1998 | | | Xenoloba lobata Arrow, 1932 | | | Xenoloba nagaii Sakai, 1998 | | | Xenoloba speciosa Bates, 1889 | | | Xylotrupes beckeri intermedius Silvestre, 2004 | | | Xylotrupes gideon Linnaeus, 1767 | Brown rhinoceros beetle | | Xylotrupes gideon borneensis Minck, 1920 | | | Xylotrupes pachycera Rowland, 2006 | | | Xylotrupes pauliani dayakorum Silvestre, 2004 | |
Subnodes of Scarabaeidae with occurrences in Sabah | Subfamily | Aphodiinae
Leach, 1815
Aphodiine Dung Beetles, Small dung beetles | 16
48 | Genera
Species |
Taxom
| | Subfamily | Cetoniinae
Leach, 1815
Flower beetles, Flower chafers, Flower scarabs, Fruit and flower chafers | 54
181 | Genera
Species |
Taxom
| | Subfamily | Dynastinae
MacLeay, 1819
Horn beetles, Rhinoceros beetles | 9
22 | Genera
Species |
Taxom
| | Subfamily | Melolonthinae
Samouelle, 1819
May Beetles and Junebugs, White grubs | 19
80 | Genera
Species |
Taxom
| | Subfamily | Rutelinae
MacLeay, 1819
Shining Flower Chafers, Shining Leaf Chafers | 12
91 | Genera
Species |
Taxom
| | Subfamily | Scarabaeinae
Latreille, 1802
Dung Beetles, True dung beetles | 17
101 | Genera
Species |
Taxom
|
 | | Scarabaeidae (Latreille 1802) |
Links, sources and further information Scarabaeidae at artfakta.se
Scarabaeidae at dyntaxa.se
Scarabaeidae at gbif.org
Scarabaeidae at irmng.org
Scarabaeidae at itis.gov
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