Checklist for Family Scarabaeidae Latreille, 1802 (Dung beetles)
and region Sarawak (Malaysia)
Class
| Insecta
Linnaeus, 1758
Insects, True insects | 298
2355
4733 | Families
Genera
Species |
Taxom
|
Subclass
| Pterygota
Lang, 1888
Winged insects | 298
2355
4733 | Families
Genera
Species |
Taxom
|
Infraclass
| Neoptera
van der Wulp, 1890
Modern Wing-folding insects | 281
2265
4550 | Families
Genera
Species |
Taxom
|
Superorder
| Holometabola
Heider, 1889
Holometabolous Insects | 180
1636
3261 | Families
Genera
Species |
Taxom
|
Order
| Coleoptera
Linnaeus, 1758
Beetles, Coleopterans | 50
821
1783 | Families
Genera
Species |
Taxom
|
Suborder
| Polyphaga
Emery, 1886
Polyphags | 47
809
1772 | Families
Genera
Species |
Taxom
|
Infraorder
| Scarabaeiformia
Crowson, 1960
| 7
141
410 | Families
Genera
Species |
Taxom
|
Superfamily
| Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles | 7
141
410 | Families
Genera
Species |
Taxom
| | Family | Scarabaeidae
Latreille, 1802
Dung beetles, Flower beetles, Fruit chafers, Rain beetles | 98
236 | Genera
Species |
Taxom
|
Genera for Sarawak
98 Genera with 236 Species #ABCDEFGHILMNOPRSTXY
| Adoretus Laporte, 1914 | | 2 | Species | Taxom | | Agestrata Eschscholtz, 1867 | | 4 | Species | Taxom | | Alocoderus Schmidt, 1916 | | 1 | Species | Taxom | | Anomala Samouelle, 1911 | Shining Leaf Chafer | 17 | Species | Taxom | | Anoplognathus Leach, 1890 | Christmas beetle | 1 | Species | Taxom | | Apogonia Melichar, 1883 | | 16 | Species | Taxom | | Ataenius Harold, 1887 | | 1 | Species | Taxom | | Autoserica Brenske, 1915 | Garden beetle | 4 | Species | Taxom | | Bombodes Westwood, 1857 | | 1 | Species | Taxom | | Brahmina Blanchard, 1892 | | 1 | Species | Taxom | | Caccobius Thomson, 1877 | | 1 | Species | Taxom | | Callistethus Blanchard, 1910 | | 2 | Species | Taxom | Catharsius Hope, 1837 | | 0 | Species | Taxom | | Ceroplophana Gestro, 1911 | | 1 | Species | Taxom | | Chaetadoretus Ohaus, 1914 | | 1 | Species | Taxom | | Chalcosoma Hope, 1900 | | 3 | Species | Taxom | | Chalcotheomima Miksic, 2020 | | 1 | Species | Taxom | | Charitovalgus Kolbe, 1891 | | 1 | Species | Taxom | | Clerota Burmeister, 1917 | | 1 | Species | Taxom | | Clinteria Burmeister, 1911 | | 3 | Species | Taxom | | Clinterocera Motschulsky, 2010 | | 1 | Species | Taxom | | Clyster Arrow, 1911 | | 1 | Species | Taxom | | Coilodera Hope, 1972 | | 2 | Species | Taxom | | Copris Geoffroy, 1875 | Mexican dung beetle | 1 | Species | Taxom | | Cyobius Sharp, 1875 | | 1 | Species | Taxom | | Dasyvalgus Kolbe, 1911 | | 5 | Species | Taxom | | Dichelomorpha Burmeister, 1920 | | 1 | Species | Taxom | | Dicheros Gory & Perch..., 1817 | | 1 | Species | Taxom | | Dichodontus Burmeister, 1911 | | 1 | Species | Taxom | | Dipelicus Hope, 1911 | | 1 | Species | Taxom | | Eudicella White, 0 | | 1 | Species | Taxom | | Euryvalgus Moser, 1944 | | 1 | Species | Taxom | | Euselates J. Thomson, 1831 | | 2 | Species | Taxom | | Exopholis Motschoulsky, 1819 | | 1 | Species | Taxom | | Fruhstorferia Willemse, 1984 | | 2 | Species | Taxom | Gametis Burmeister, 1842 | | 0 | Species | Taxom | | Glycyphana Burmeister, 1914 | | 8 | Species | Taxom | | Glyptothea Bates, 1889 | | 2 | Species | Taxom | | Gnathocera Kirby, 1836 | | 1 | Species | Taxom | | Gymnetis MacLeay, 1875 | | 1 | Species | Taxom | | Heptelia Brenske, 1944 | | 1 | Species | Taxom | | Heterovalgus Krikken, 1992 | | 1 | Species | Taxom | | Hoplia Illiger, 1912 | Monkey Beetles | 2 | Species | Taxom | Hybovalgus Kolbe, 1904 | | 0 | Species | Taxom | | Ischiopsopha Gestro, 1860 | | 1 | Species | Taxom | | Ixorida Thomson, 1899 | | 1 | Species | Taxom | | Latipalpus Moser, 1921 | | 4 | Species | Taxom | | Lepidiota Kirby, 1876 | | 3 | Species | Taxom | | Lepivalgus Moser, 1891 | | 1 | Species | Taxom | | Leucopholis Blanchard, 1892 | | 2 | Species | Taxom | | Lorkovitschia Miksic, 1868 | | 1 | Species | Taxom | | Maladera Mulsant, 1899 | | 2 | Species | Taxom | | Mecinonota Kraatz, 1801 | | 1 | Species | Taxom | | Meroloba Thomson, 1858 | | 1 | Species | Taxom | | Microcopris Balthasar, 1877 | | 1 | Species | Taxom | | Microserica Brenske, 1946 | | 9 | Species | Taxom | | Microvalgus Kraatz, 1993 | | 2 | Species | Taxom | | Mimela Kirby, 1908 | | 1 | Species | Taxom | | Miridiba Reitter, 1944 | | 2 | Species | Taxom | | Nematoserica Arrow, 1917 | | 1 | Species | Taxom | | Neoserica Brenske, 1915 | | 1 | Species | Taxom | | Ochicanthon Vaz-de-Mello, 2007 | | 6 | Species | Taxom | | Octoplasia Brenske, 1876 | | 2 | Species | Taxom | | Odochilus Harold, 1997 | | 3 | Species | Taxom | | Onthophagus Latreille, 1877 | | 12 | Species | Taxom | | Oreovalgus Kolbe, 1891 | | 2 | Species | Taxom | | Paragymnopleurus Shipp, 1875 | | 1 | Species | Taxom | | Paraphytus Harold, 1923 | | 2 | Species | Taxom | | Parascatonomus Paulian, 2000 | | 9 | Species | Taxom | | Parastasia Westwood, 1881 | | 6 | Species | Taxom | | Peltonotus Burmeister, 1910 | | 5 | Species | Taxom | | Pentelia Kiriakoff, 2005 | | 4 | Species | Taxom | | Philistina MacLeay, 2001 | | 1 | Species | Taxom | | Plectrone Wallace, 1883 | | 1 | Species | Taxom | | Pleuronota Kraatz, 1977 | | 2 | Species | Taxom | | Popillia Leach, 1923 | | 1 | Species | Taxom | | Protaetia Burmeister, 1841 | | 2 | Species | Taxom | | Pseudochalcothea Ritsema, 1889 | | 5 | Species | Taxom | | Pseudochalcotheomima Miksic, 1990 | | 1 | Species | Taxom | | Pseudohomonyx Arrow, 1999 | | 2 | Species | Taxom | | Psilopholis Brenske, 2010 | | 1 | Species | Taxom | | Pterobius Maruyama, 2010 | | 1 | Species | Taxom | | Rhyparus Westwood, 1866 | | 2 | Species | Taxom | | Rutelarcha Waterhouse, 1874 | | 1 | Species | Taxom | | Saprosites Redtenbacher, 1944 | | 5 | Species | Taxom | | Setylaides | | 1 | Species | Taxom | | Sophrops Fairmaire, 1892 | | 2 | Species | Taxom | | Spinanomala Ohaus, 1910 | | 4 | Species | Taxom | Svobia Miksic, 1965 | | 0 | Species | Taxom | | Taeniodera Burmeister, 1833 | | 6 | Species | Taxom | | Termitodiellus Nakane, 1987 | | 1 | Species | Taxom | Thaumastopeus Kraatz, 1885 | | 0 | Species | Taxom | | Theodosia J. Thomson, 1889 | | 5 | Species | Taxom | | Trichogomphus Burmeister, 1907 | | 1 | Species | Taxom | | Xenaclopus Arrow, 1915 | | 1 | Species | Taxom | | Xenoloba Bates, 1998 | | 1 | Species | Taxom | | Xylotrupes Hope, 1997 | Elephant beetles | 3 | Species | Taxom | | Yvescambefortius Ochi & Kon, 1926 | | 1 | Species | Taxom |
Species list for Sarawak
236 Species and 10 Subspecies | Adoretus moultoni Ohaus, 1914 | | | Adoretus trichostigma Ohaus, 1914 | | | Agestrata belitungana Miksic, 1973 | | | Agestrata dehaan Gory & Perch..., 1833 | | | Agestrata orichalca Linnaeus, 1789 | | | Agestrata parryi Wallace, 1867 | | | Alocoderus imitator Koshanschikov, 1916 | | | Anomala angulipennis Ohaus, 1910 | | | Anomala bifida Zorn, 2007 | | | Anomala concha Ohaus, 1916 | | | Anomala felicia Arrow, 1910 | | | Anomala forcipalis Ohaus, 1910 | | | Anomala furcula Ohaus, 1916 | | | Anomala lasiocnemis Ohaus, 1930 | | | Anomala manseri Zorn, 2007 | | | Anomala mascula Zorn, 2005 | | | Anomala matangensis Zorn, 2007 | | | Anomala parotidea Ohaus, 1910 | | | Anomala ponticula Ohaus, 1915 | | | Anomala posticalis Arrow, 1910 | | | Anomala saetipes Ohaus, 1916 | | | Anomala sarawakensis Ohaus, 1910 | | | Anomala ulcerata Ohaus, 1915 | | | Anomala ventriosa Ohaus, 1911 | | | Anoplognathus punctulatus Olliff, 1890 | Leaf green Christmas beetle | | Apogonia albomaculata Kobayashi, 2010 | | | Apogonia armadillo Arrow, 1938 | | | Apogonia bakunensis Kobayashi, 2013 | | | Apogonia crenulata Kobayashi, 2013 | | | Apogonia cuprina Moser, 1913 | | | Apogonia excedens Kobayashi, 2013 | | | Apogonia flavipes Gestro, 1883 | | | Apogonia hilaris Arrow, 1938 | | | Apogonia laeta Arrow, 1938 | | | Apogonia lobata (Ritsema, 1904) | ⇒ Dichecephala moseri Ritsema, 1912 | | Apogonia longispina Kobayashi, 2010 | | | Apogonia nobilis Arrow, 1938 | | | Apogonia opaca Gestro, 1883 | | | Apogonia sericea Gestro, 1883 | | | Apogonia tessellata Arrow, 1938 | | | Apogonia viridipennis Gestro, 1883 | | | Ataenius liogaster Bates, 1887 | | | Autoserica fasciata Moser, 1911 | | | Autoserica multimaculata Brenske, 1899 | | | Autoserica permagna Moser, 1915 | | | Autoserica sarawakensis Moser, 1915 | | | Bombodes westwoodi Thomson, 1857 | | | Brahmina rugulosa Brenske, 1892 | | | Caccobius binodulus Harold, 1877 | | | Callistethus morio Ohaus, 1913 | | | Callistethus moultoni Ohaus, 1910 | | | Ceroplophana modiglianii borneensis Ohaus, 1911 | | | Chaetadoretus trichostigma Ohaus, 1914 | | | Chalcosoma atlas Linnaeus, 1758 | Atlas beetle | | Chalcosoma atlas keyboh Nagai, 2004 | | | Chalcosoma moellenkampi Kolbe, 1900 | Moellenkampi beetle | | Chalcotheomima paramacrophylla Kawahara, To..., 2020 | | | Charitovalgus doriae Gestro, 1891 | | | Clerota rigifica Janson, 1917 | | | Clinteria atra Wiedemann, 1823 | | | Clinteria hageni borneoana Jákl, 2020 | | | Clinteria moultoni Moser, 1911 | | | Clinterocera cyprilrusi Legrand & Ch..., 2010 | | | Clyster trachypygus Prell, 1911 | | | Coilodera chungi Legrand & Ch..., 2010 | | | Coilodera kalimantanica Miksic, 1972 | | | Copris agnus Sharp, 1875 | | | Cyobius wallacei Sharp, 1875 | | | Dasyvalgus eucharis Kolbe, 1904 | | | Dasyvalgus gilleti Moser, 1906 | | | Dasyvalgus nigrofasciculatus Moser, 1906 | | | Dasyvalgus rufipennis Arrow, 1944 | | | Dasyvalgus sarawakensis Moser, 1911 | | | Dichelomorpha aranea Arrow, 1920 | | | Dicheros bicornis Latreille, 1817 | | | Dichodontus fissicornis Arrow, 1911 | | | Dipelicus borneensis Arrow, 1911 | | | Eudicella euthalia euthalia | | | Euryvalgus tridens Arrow, 1944 | | | Euselates cineracea Gory & Perch..., 1833 | | | Euselates quadrilineata Hope, 1831 | | | Exopholis hypoleuca Wiedemann, 1819 | | | Fruhstorferia flavipennis Nagai, 1984 | | | Fruhstorferia nigromuliebris Nagai, 1984 | | | Glycyphana cincta Wallace, 1867 | | | Glycyphana festiva Fabricius, 1792 | | | Glycyphana illusa Janson, 1881 | | | Glycyphana inusta Mohnike, 1871 | | | Glycyphana nicobarica Janson, 1877 | | | Glycyphana penanga Wallace, 1867 | | | Glycyphana sarawakensis Moser, 1914 | | | Glycyphana tonkinensis Moser, 1914 | | | Glyptothea moultoni Janson, 1912 | | | Glyptothea whiteheadi Bates, 1889 | | | Gnathocera petelli (Buquet, 1836) | ⇒ Dicheros bicornis Latreille, 1817 | | Gymnetis goryi Janson, 1875 | | | Heptelia polyphylla Arrow, 1944 | | | Heterovalgus alberti Ricchiardi, 1992 | | | Hoplia aurata Waterhouse, 1877 | | | Hoplia moultoni Moser, 1912 | | | Ischiopsopha olivacea (Thomson, 1860) | ⇒ Ischiopsopha jamesi Waterhouse, 1876 | | Ixorida apicalis Kraatz, 1899 | | | Latipalpus elegans Itoh, 1999 | | | Latipalpus fujiokai Itoh, 1999 | | | Latipalpus maxillatus Brenske, 1892 | | | Latipalpus palpalis Moser, 1921 | | | Lepidiota flavimargo Arrow, 1919 | | | Lepidiota marginipennis Moser, 1913 | | | Lepidiota munda Sharp, 1876 | | | Lepivalgus borneensis Gestro, 1891 | | | Leucopholis armata Sharp, 1876 | | | Leucopholis staudingeri Brenske, 1892 | | | Lorkovitschia rustica Wallace, 1868 | | | Maladera drescheri Moser, 1913 | | | Maladera spissa Brenske, 1899 | | | Mecinonota regia Fabricius, 1801 | | | Meroloba suturalis Snellen van ..., 1858 | | | Microcopris doriae Harold, 1877 | | | Microserica fascipennis Moser, 1911 | | | Microserica fulvovittata Moser, 1911 | | | Microserica imitatrix Moser, 1915 | | | Microserica latefemorata Moser, 1915 | | | Microserica lineolata Moser, 1915 | | | Microserica nigrosuturata Moser, 1911 | | | Microserica sanguineicollis Moser, 1911 | | | Microserica sarawakana Moser, 1920 | | | Microserica shelfordi Arrow, 1946 | | | Microvalgus excavatus Antoine, 1993 | | | Microvalgus pachyphallus Antoine, 1993 | | | Mimela maculicollis (Ohaus, 1908) | ⇒ Mimela junii rugosula Fairmaire, 1859 | | Miridiba borneensis Moser, 1918 | | | Miridiba coxalis Arrow, 1944 | | | Nematoserica coerulea Arrow, 1917 | | | Neoserica limbangica Moser, 1915 | | | Ochicanthon crypticus Krikken & Hu..., 2007 | | | Ochicanthon dulitmontis Krikken & Hu..., 2007 | | | Ochicanthon edmondsi Krikken & Hu..., 2007 | | | Ochicanthon hanskii Krikken & Hu..., 2007 | | | Ochicanthon mulu Krikken & Hu..., 2007 | | | Ochicanthon neglectus Krikken & Hu..., 2007 | | | Octoplasia lineata Sharp, 1876 | | | Octoplasia princeps Sharp, 1876 | | | Odochilus borneensis RakoviÄ� & A..., 2021 | | | Odochilus haroldi Rakovic, 1987 | | | Odochilus sarawakensis Rakovic, 1997 | | | Onthophagus (Macronthophagus) diabolicus Harold, 1877 | | | Onthophagus (Paronthophagus) hidakai Ochi & Kon, 1995 | | | Onthophagus baramtagal Ochi, Kon & ..., 2016 | | | Onthophagus collinsi Krikken & Hu..., 1987 | | | Onthophagus incisus Harold, 1877 | | | Onthophagus infucatus Harold, 1877 | | | Onthophagus kapitensis Frey, 1971 | | | Onthophagus keikoae Ochi & Kon, 2014 | | | Onthophagus mulutagal Ochi, Kon & ..., 2016 | | | Onthophagus ochromerus Harold, 1877 | | | Onthophagus pavidus Harold, 1877 | | | Onthophagus rorarius Harold, 1877 | | | Oreovalgus bryanti Arrow, 1944 | | | Oreovalgus cristatus Gestro, 1891 | | | Paragymnopleurus maurus Sharp, 1875 | | | Paraphytus doriae Harold, 1877 | | | Paraphytus foveatus Boucomont, 1923 | | | Parascatonomus aurifex Harold, 1877 | | | Parascatonomus discedens Sharp, 1875 | | | Parascatonomus dux Sharp, 1875 | | | Parascatonomus fujiokai Ochi & Araya, 1996 | | | Parascatonomus itiokai Ochi, Kon & ..., 2010 | | | Parascatonomus sarawacus Harold, 1877 | | | Parascatonomus semicupreus Harold, 1877 | | | Parascatonomus serapiensis Ochi, Kon & ..., 2008 | | | Parascatonomus tamijii Kon, Sakai &..., 2000 | | | Parastasia bigibbosa Nonfried, 1891 | | | Parastasia confluens Westwood, 1841 | | | Parastasia discolor discolor | | | Parastasia gestroi Ohaus, 1900 | | | Parastasia moultoni Ohaus, 1911 | | | Parastasia nigripennis Sharp, 1881 | | | Peltonotus malayensis Arrow, 1910 | | | Peltonotus silvanus Jameson & Wada, 2004 | | | Peltonotus similis Arrow, 1931 | | | Peltonotus suehirogarus Jameson & Wada, 2004 | | | Peltonotus vittatus Arrow, 1910 | | | Pentelia coxalis Arrow, 1944 | | | Pentelia crinifrons Moser, 1912 | | | Pentelia desiderata Brenske, 1894 | | | Pentelia malaccensis apicalis Matsumoto, 2005 | | | Philistina sakaii Alexis & Del..., 2001 | | | Plectrone lugubris Janson, 1883 | | | Pleuronota allardi Antoine, 1990 | | | Pleuronota alleni Krikken, 1977 | | | Popillia dajaka Ohaus, 1923 | | | Protaetia (Macroprotaetia) gestroi Miksic, 1970 | | | Protaetia (Oxyperatex) spectabilis Schaum, 1841 | | | Pseudochalcothea auripes Westwood, 1874 | | | Pseudochalcothea planiuscula Bates, 1889 | | | Pseudochalcothea pomacea Bates, 1889 | | | Pseudochalcothea shelfordi Janson, 1912 | | | Pseudochalcothea spathulifera Bates, 1889 | | | Pseudochalcotheomima allardi Antoine, 1990 | | | Pseudohomonyx borneensis Arrow, 1908 | | | Pseudohomonyx similis Miyake & Yamaya, 1999 | | | Psilopholis gigantea Matsumoto, 2010 | | | Pterobius itiokai Maruyama, 2010 | | | Rhyparus pseudominor Bordat, 1996 | | | Rhyparus saundersi Pascoe, 1866 | | | Rutelarcha quadrimaculata Waterhouse, 1874 | | | Saprosites kapitensis Petrovitz, 1973 | | | Saprosites marchionalis (Harold, 1877) | ⇒ Saprosites cycloporum Paulian, 1944 | | Saprosites matangi Stebnicka, 2012 | | | Saprosites penrisseni Stebnicka, 2012 | | | Saprosites rufus Paulian, 1944 | | | Setylaides monstrosus Harold, 1877 | | | Sophrops rugulosus rugulosus | | | Sophrops sharpi Brenske, 1892 | | | Spinanomala bispinosa Ohaus, 1910 | | | Spinanomala gracilis Kobayashi, 2012 | | | Spinanomala psilopyga Ohaus, 1910 | | | Spinanomala unispinosa Ohaus, 1910 | | | Taeniodera aurantiaca Snellen van ..., 1858 | | | Taeniodera cervina Wallace, 1867 | | | Taeniodera inermis Krikken, 1982 | | | Taeniodera marmorata Wallace, 1867 | | | Taeniodera modesta modesta | | | Taeniodera monacha Gory & Perch..., 1833 | | | Termitodiellus hammondi Krikken & Hu..., 1987 | | | Theodosia howitti Castelnau, 1873 | | | Theodosia magnifica bawangensis Nagai, 1998 | | | Theodosia nagai Legrand & Ch..., 2010 | | | Theodosia telifer Bates, 1889 | | | Theodosia viridiaurata Bates, 1889 | | | Trichogomphus zangi Sternberg, 1907 | | | Xenaclopus borneensis Arrow, 1915 | | | Xenoloba nagaii Sakai, 1998 | | | Xylotrupes gideon Linnaeus, 1767 | Brown rhinoceros beetle | | Xylotrupes pauliani dayakorum Silvestre, 2004 | | | Xylotrupes wiltrudae Silvestre, 1997 | | | Yvescambefortius sarawacus Gillet, 1926 | |
Subnodes of Scarabaeidae with occurrences in Sarawak | Subfamily | Aphodiinae
Leach, 1815
Aphodiine Dung Beetles, Small dung beetles | 8
15 | Genera
Species |
Taxom
| | Subfamily | Cetoniinae
Leach, 1815
Flower beetles, Flower chafers, Flower scarabs, Fruit and flower chafers | 39
71 | Genera
Species |
Taxom
| | Subfamily | Dynastinae
MacLeay, 1819
Horn beetles, Rhinoceros beetles | 7
11 | Genera
Species |
Taxom
| | Subfamily | Melolonthinae
Samouelle, 1819
May Beetles and Junebugs, White grubs | 20
60 | Genera
Species |
Taxom
| | Subfamily | Rutelinae
MacLeay, 1819
Shining Flower Chafers, Shining Leaf Chafers | 12
43 | Genera
Species |
Taxom
| | Subfamily | Scarabaeinae
Latreille, 1802
Dung Beetles, True dung beetles | 9
25 | Genera
Species |
Taxom
|
 | | Scarabaeidae (Latreille 1802) |
Links, sources and further information Scarabaeidae at artfakta.se
Scarabaeidae at dyntaxa.se
Scarabaeidae at gbif.org
Scarabaeidae at irmng.org
Scarabaeidae at itis.gov
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