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TAXFAMGENSPESUB
Checklist for Family Scarabaeidae Latreille, 1802 (Dung beetles)
and region Alajuela (Costa Rica)


Class
 
Insecta
Linnaeus, 1758
Insects, True insects
408
5803
10301
Families
Genera
Species
 
Taxom
 

Subclass
 
Pterygota
Lang, 1888
Winged insects
407
5802
10301
Families
Genera
Species
 
Taxom
 

Infraclass
 
Neoptera
van der Wulp, 1890
Modern Wing-folding insects
393
5734
10113
Families
Genera
Species
 
Taxom
 

Superorder
 
Holometabola
Heider, 1889
Holometabolous Insects
280
5013
9221
Families
Genera
Species
 
Taxom
 

Order
 
Coleoptera
Linnaeus, 1758
Beetles, Coleopterans
66
1524
2171
Families
Genera
Species
 
Taxom
 

Suborder
 
Polyphaga
Emery, 1886
Polyphags
62
1390
1993
Families
Genera
Species
 
Taxom
 

Infraorder
 
Scarabaeiformia
Crowson, 1960
7
147
489
Families
Genera
Species
 
Taxom
 

Superfamily
 
Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles
7
147
489
Families
Genera
Species
 
Taxom
 
FamilyScarabaeidae
Latreille, 1802
Dung beetles, Flower beetles, Fruit chafers, Rain beetles
120
429
Genera
Species
 
Taxom
 


Genera for Alajuela
120 Genera with 429 Species
#ABCDEFGHILMNOPRSTU
Aegidium Blanchard, 1845
0SpeciesTaxom
Agamopus Bates, 18871SpeciesTaxom
Amblyoproctus Kolbe, 20031SpeciesTaxom
Amithao Thomson, 18854SpeciesTaxom
Ancognatha Erichson, 19113SpeciesTaxom
Anomala Samouelle, 1888Shining Leaf Chafer38SpeciesTaxom
Anomalorhina Jameson, Pau..., 19281SpeciesTaxom
Anomiopus Westwood, 19392SpeciesTaxom
Aphodius Hellwig, 17892SpeciesTaxom
Archophileurus Kolbe, 18881SpeciesTaxom
Argyripa Thomson, 18691SpeciesTaxom
Aspidolea Bates, 18884SpeciesTaxom
Astaena Erichson, 1847
0SpeciesTaxom
Ataenius Harold, 1867
0SpeciesTaxom
Ateuchus Weber, 19966SpeciesTaxom
Balsameda Thomson, 18771SpeciesTaxom
Barybas Burmeister, 18871SpeciesTaxom
Bdelyrus Harold, 18871SpeciesTaxom
Bolax Fischer von ..., 18881SpeciesTaxom
Bradypodidium Vaz-de-Mello, 19281SpeciesTaxom
Caccobius Thomson, 19111SpeciesTaxom
Callistethus Blanchard, 188814SpeciesTaxom
Calomacraspis Bates, 18691SpeciesTaxom
Canthidium Erichson, 198116SpeciesTaxom
Canthon Hoffmannsegg, 1964Tumblebug13SpeciesTaxom
Ceraspis Serville, 1825
0SpeciesTaxom
Chasmodia MacLeay, 1819
0SpeciesTaxom
Chlaenobia Blanchard, 19921SpeciesTaxom
Chlorota Burmeister, 19122SpeciesTaxom
Chrysina Kirby, 1875Jewel scarabs10SpeciesTaxom
Cnemida Kirby, 18882SpeciesTaxom
Coelosis Hope, 17671SpeciesTaxom
Copris Geoffroy, 1910Mexican dung beetle6SpeciesTaxom
Coprophanaeus Olsoufieff, 20087SpeciesTaxom
Cotinis Burmeister, 1876Green June Beetles4SpeciesTaxom
Cyclocephala Dejean, 1992Masked Chafers37SpeciesTaxom
Daemonoplus Thomson, 1858
0SpeciesTaxom
Deltochilum Eschscholtz, 18735SpeciesTaxom
Dendropaemon Perty, 1830
0SpeciesTaxom
Dialithus Parry, 18491SpeciesTaxom
Dichotomius Hope, 18677SpeciesTaxom
Digitonthophagus Balthasar, 17871SpeciesTaxom
Diplotaxis Kirby, 1887Wall-Rockets5SpeciesTaxom
Disphysema Harold, 18731SpeciesTaxom
Dorysthetus Blanchard, 20021SpeciesTaxom
Dynastes MacLeay, 1985Hercules Beetles2SpeciesTaxom
Dyscinetus Harold, 1888Rice Beetles2SpeciesTaxom
Enema Hope, 18431SpeciesTaxom
Epectinaspis Blanchard, 20141SpeciesTaxom
Euetheola Bates, 18472SpeciesTaxom
Euparixia Brown, 19361SpeciesTaxom
Euphoria Burmeister, 18016SpeciesTaxom
Eurysternus Dalman, 18877SpeciesTaxom
Eutrichillum1SpeciesTaxom
Faula Blanchard, 18772SpeciesTaxom
Geniates Kirby, 19761SpeciesTaxom
Genuchinus Westwood, 19351SpeciesTaxom
Giesbertiolus1SpeciesTaxom
Golofa Hope, 20032SpeciesTaxom
Goniophileurus Kolbe, 18471SpeciesTaxom
Guatemalica Poll, 18701SpeciesTaxom
Gymnetis MacLeay, 18421SpeciesTaxom
Helictopleurus Orbigny, 20001SpeciesTaxom
Heliocopris Hope, 17751SpeciesTaxom
Hemiphileurus Kolbe, 19142SpeciesTaxom
Heterogomphus Burmeister, 18592SpeciesTaxom
Heterosternus Moure, 19011SpeciesTaxom
Homocopris Burmeister, 18671SpeciesTaxom
Homophileurus Kolbe, 19141SpeciesTaxom
Hoplia Illiger, 1887Monkey Beetles2SpeciesTaxom
Hoplopyga Thomson, 18332SpeciesTaxom
Hypaspidius Arrow, 20021SpeciesTaxom
Inca Lepeletier &..., 17921SpeciesTaxom
Isonychus Mannerheim, 19185SpeciesTaxom
Lagochile Hoffmannsegg, 03SpeciesTaxom
Leucothyreus MacLeay, 18441SpeciesTaxom
Liparetrus Guérin-Ménev..., 19801SpeciesTaxom
Luederwaldtinia Martínez, 19973SpeciesTaxom
Macraspis MacLeay, 18883SpeciesTaxom
Macrodactylus Dejean, 1887Rose Chafers6SpeciesTaxom
Macropoides Guérin-Ménev..., 19341SpeciesTaxom
Megaceras Hope, 18881SpeciesTaxom
Megasoma Kirby, 1775Elephant beetles1SpeciesTaxom
Melinopterus Mulsant, 17901SpeciesTaxom
Mimela Kirby, 19051SpeciesTaxom
Minidorysthetus2SpeciesTaxom
Nialaphodius Kolbe, 18011SpeciesTaxom
Nimbus Mulsant & Rey, 19942SpeciesTaxom
Ontherus Erichson, 19965SpeciesTaxom
Onthophagus Latreille, 200136SpeciesTaxom
Parachlorota Soula, 20021SpeciesTaxom
Parapucaya Prell, 19341SpeciesTaxom
Pedaridium Harold, 1868
0SpeciesTaxom
Pelidnota MacLeay, 1840Precious Metal Scarabs7SpeciesTaxom
Phalangogonia Burmeister, 18771SpeciesTaxom
Phanaeus MacLeay, 1863Burrowing Dung Beetles5SpeciesTaxom
Phileurus Sieg, 19886SpeciesTaxom
Phyllophaga Harris, 1934June beetles30SpeciesTaxom
Platycoelia Dejean, 19043SpeciesTaxom
Plectris Lepeletier a..., 19181SpeciesTaxom
Plusiotus
0SpeciesTaxom
Pseudocanthon Bates, 18471SpeciesTaxom
Ptenomela Bates, 19382SpeciesTaxom
Pucaya Ohaus, 19101SpeciesTaxom
Rutela Latreille, 19051SpeciesTaxom
Scatimus Erichson, 18622SpeciesTaxom
Serica MacLeay, 19202SpeciesTaxom
Sisyphus Latreille, 18331SpeciesTaxom
Smaragdesthes Kraatz, 18361SpeciesTaxom
Spodistes Burmeister, 18603SpeciesTaxom
Spodochlamys Burmeister, 18881SpeciesTaxom
Stenocrates Burmeister, 18703SpeciesTaxom
Strategus Kirby, 1847Ox Beetles2SpeciesTaxom
Strigoderma Burmeister, 18448SpeciesTaxom
Sulcophanaeus Olsoufieff, 18562SpeciesTaxom
Thyriochlorota Ohaus, 19051SpeciesTaxom
Tomarus Erichson, 1888Carrot Beetles4SpeciesTaxom
Trichillidium Vaz-de-Mello, 19481SpeciesTaxom
Trigonopeltastes Burmeister &..., 18892SpeciesTaxom
Uroxys Westwood, 198710SpeciesTaxom


Species list for Alajuela
429 Species and 4 Subspecies
Agamopus lampros Bates, 1887
Amblyoproctus centroamericanus Ratcliffe, 2003
Amithao albopictus Poll, 1886
Amithao decemguttatus Waterhouse, 1876
Amithao haematopus Schaum, 1848
Amithao metallicus Janson, 1885
Ancognatha gracilis Endrödi, 1966
Ancognatha vexans Ratcliffe, 1992
Ancognatha vulgaris Arrow, 1911
Anomala aglaos Filippini, G..., 2015
Anomala calligrapha Bates, 1888
Anomala chiriquina Bates, 1888
Anomala clathrata Ohaus, 1930
Anomala coffea Filippini, G..., 2015
Anomala divisa Filippini, G..., 2015
Anomala dorsata Fåhraeus, 1857
Anomala estrella Filippini, G..., 2015
Anomala eulissa Bates, 1888
Anomala flavacoma Filippini, M..., 2013
Anomala gemella Say, 1835
Anomala hiata Filippini, M..., 2015
Anomala hoppi Ohaus, 1928
Anomala levicollis Filippini, M..., 2015
Anomala limon Filippini, M..., 2016
Anomala longisacculata Filippini, M..., 2015
Anomala moroni Filippini, M..., 2015
Anomala ochrogastra Bates, 1888
Anomala pincelada Filippini, G..., 2015
Anomala plurisulcata Benderitter, 1930
Anomala praecellens Bates, 1888
Anomala pseudoeucoma Filippini, M..., 2013
Anomala quiche Ohaus, 1897
Anomala robiginosa Filippini, G..., 2015
Anomala semicincta Bates, 1888
Anomala semilla Filippini, G..., 2014
Anomala solisi Filippini, G..., 2014
Anomala strigodermoides Filippini, G..., 2015
Anomala subaenea Nonfried, 1893
Anomala subusta Filippini, M..., 2015
Anomala sulana Zorn, 2006
Anomala tenoriensis Filippini, M..., 2015
Anomala testaceipennis (Blanchard, 1851)⇒ Anomala nitidula Blanchard, 1851
Anomala tuberculata Filippini, M..., 2015
Anomala unilineata Filippini, G..., 2015
Anomala valida (Burmeister, 1844)⇒ Anomala insularis Castelnau, 1840
Anomala vallisneria Filippini, M..., 2015
Anomala veraecrucis Bates, 1888
Anomalorhina turrialbana Ohaus, 1928
Anomiopus birai Canhedo, 2006
Anomiopus panamensis Paulian, 1939
Aphodius (Aphodius) foetidus Herbst, 1783
Aphodius (Nialus) lividus (Olivier, 1789)⇒ Acrossus luridus Fabricius, 1775
Archophileurus simplex Bates, 1888
Argyripa anomala Bates, 1869
Aspidolea fuliginea Burmeister, 1847
Aspidolea kuntzeni Höhne, 1922
Aspidolea notaticollis Höhne, 1922
Aspidolea singularis Bates, 1888
Ateuchus chrysopyge Bates, 1887
Ateuchus fetteri Kohlmann, 1996
Ateuchus hendrichsi Kohlmann, 1996
Ateuchus rodriguezi Preudhomme d..., 1886
Ateuchus solisi Kohlmann, 1996
Ateuchus zoebischi Kohlmann, 1996
Balsameda adspersa Sharp, 1877
Barybas idiota Bates, 1887
Bdelyrus seminudus Bates, 1887
Bolax flaveola Bates, 1888
Bradypodidium bradyporum Boucomont, 1928
Caccobius polygonus d´Orbigny, 1911
Callistethus chlorotoides Bates, 1888
Callistethus chontalensis Bates, 1888
Callistethus granulipygus Bates, 1888
Callistethus lativittis Filippini, G..., 2015
Callistethus macroxantholeus Filippini, G..., 2015
Callistethus microxantholeus Filippini, G..., 2015
Callistethus mimeloides Ohaus, 1902
Callistethus multiplicatus Filippini, G..., 2015
Callistethus nicoya Ohaus, 1928
Callistethus pseudocollaris Filippini, G..., 2015
Callistethus ruteloides Filippini, G..., 2015
Callistethus sulcans Bates, 1888
Callistethus vanpatteni Bates, 1888
Callistethus xiphostethus Bates, 1888
Calomacraspis haroldi Candeze, 1869
Canthidium angusticeps Bates, 1887
Canthidium annagabrielae Solis & Kohl..., 2004
Canthidium ardens Bates, 1887
Canthidium aurifex Bates, 1887
Canthidium centrale Boucomont, 1928
Canthidium discopygidiale Howden & Young, 1981
Canthidium guanacaste Howden & Gill, 1987
Canthidium haroldi Preudhomme d..., 1886
Canthidium hespenheidei Howden & Young, 1981
Canthidium marianelae Solis & Kohl..., 2004
Canthidium perceptibile Howden & Young, 1981
Canthidium planovultum Howden & Young, 1981
Canthidium priscillae Solis & Kohl..., 2004
Canthidium tenebrosum Howden & Young, 1981
Canthidium tuberifrons Howden & Young, 1981
Canthidium vespertinum Howden & Young, 1981
Canthon (Boreocanthon) ebenus Say, 1823
Canthon (Canthon) cyanellus LeConte, 1859
Canthon (Canthon) indigaceus (LeConte, 1866)⇒ Canthon humectus Say, 1831
Canthon aberrans Harold, 1868
Canthon angustatus Harold, 1867
Canthon euryscelis Bates, 1887
Canthon hartmanni Howden & Gill, 1987
Canthon meridionalis Martinez, Ha..., 1964
Canthon moniliatus (Bates, 1887)⇒ Scybalocanthon moniliatus Bates, 1887
Canthon silvaticus Solis & Kohl..., 2002
Canthon staigi Pereira, 1953
Canthon subhyalinus Harold, 1867
Canthon vazquezae Martinez, Ha..., 1964
Chlaenobia halffteriana Morón, 1992
Chlorota batesi Soula, 2002
Chlorota panamensis Ohaus, 1912
Chrysina aurigans Rothschildt ..., 1894
Chrysina aurora Boucard, 1875
Chrysina batesi Boucard, 1875
Chrysina boucardi Sallé, 1878
Chrysina bruyeai Hawks, 1999
Chrysina chrysargyrea Sallé, 1874
Chrysina cupreomarginata Bates, 1904
Chrysina limbata Rothschildt ..., 1894
Chrysina luteomarginata Ohaus, 1913
Chrysina resplendens Boucard, 1875
Cnemida aterrima Bates, 1888
Cnemida intermedia Bates, 1888
Coelosis biloba Linnaeus, 1767
Copris costaricensis Gahan, 1894
Copris incertus Say, 1835Black dung beetle
Copris laeviceps Harold, 1869
Copris lugubris Boheman, 1858
Copris remotus dicyrtus Matthews & H..., 1959
Copris subpunctatus Gillet, 1910
Coprophanaeus boucardi Nevinson, 1891
Coprophanaeus chiriquensis Olsufieff, 1924
Coprophanaeus kohlmanni (Arnaud, 2002)⇒ Coprophanaeus morenoi Arnaud, 1982
Coprophanaeus morenoi Arnaud, 1982
Coprophanaeus pecki Howden & Young, 1981
Coprophanaeus telamon Erichson, 1847
Coprophanaeus uhleri (Maly & Pokorny, 2008)⇒ Coprophanaeus chiriquensis Olsufieff, 1924
Cotinis (Cotinis) mutabilis Gory & Perch..., 1883Fig beetle
Cotinis lebasi (Gory & Perch..., 1833)⇒ Cotinorrhina lebasi Gory & Perch..., 1833
Cotinis olivia Bates, 1889
Cotinis polita Janson, 1876
Cyclocephala almitana Dechambre, 1992
Cyclocephala amazona Linnaeus, 1767
Cyclocephala amblyopsis Hurley, 1959
Cyclocephala ampliata Bates, 1888
Cyclocephala atripes Bates, 1888
Cyclocephala carbonaria Milne Edwards, 1848
Cyclocephala castaniella Bates, 1888
Cyclocephala cerea (Burmeister, 1847)⇒ Cyclocephala sanguinicollis Burmeister, 1847
Cyclocephala complanata Burmeister, 1847
Cyclocephala concolor Burmeister, 1847
Cyclocephala confusa Endrödi, 1966
Cyclocephala conspicua Sharp, 1877
Cyclocephala discolor Herbst, 1792
Cyclocephala erotylina Arrow, 1914
Cyclocephala fasciolata Bates, 1888
Cyclocephala fulgurata Burmeister, 1847
Cyclocephala fulvipennis Burmeister, 1847
Cyclocephala gravis Bates, 1888
Cyclocephala kaszabi Endrödi, 1964
Cyclocephala ligyrina Bates, 1888
Cyclocephala lunulata Burmeister, 1847Twister
Cyclocephala macrophylla Cohen, 1998
Cyclocephala maculiventris Höhne, 1923
Cyclocephala mafaffa Burmeister, 1847Twister
Cyclocephala melanocephala (Fabricius, 1775)⇒ Cyclocephala olivieri Arrow, 1911
Cyclocephala multiplex (Casey, 1915)⇒ Cyclocephala amazona Linnaeus, 1767
Cyclocephala mutata Harold, 1869
Cyclocephala nigerrima Bates, 1888
Cyclocephala nigritarsis Ratcliffe, 1992
Cyclocephala ovulum (Bates, 1888)⇒ Cyclocephala testacea Burmeister, 1847
Cyclocephala pan Ratcliffe, 1992
Cyclocephala porioni Dechambre, 1979
Cyclocephala sanguinicollis Burmeister, 1847Twister
Cyclocephala sexpunctata Castelnau, 1840
Cyclocephala stictica Burmeister, 1847
Cyclocephala weidneri Endrödi, 1964
Cyclocephala williami Ratcliffe, 1992
Deltochilum gibbosum Fabricius, 1775Humpback Dung Beetle
Deltochilum mexicanum Burmeister, 1848
Deltochilum parile Bates, 1887
Deltochilum pseudoparile Paulian, 1938
Deltochilum valgum Burmeister, 1873
Dialithus magnificus Parry, 1849
Dichotomius (Luederwaldtinia) centralis Harold, 1869
Dichotomius (Luederwaldtinia) danieli (Kohlmann & S..., 1997)⇒ Luederwaldtinia danieli Kohlmann & S..., 1997
Dichotomius (Luederwaldtinia) femoratus (Howden & Young, 1981)⇒ Luederwaldtinia femoratus Howden & Young, 1981
Dichotomius (Luederwaldtinia) rodrigoi (Kohlmann & S..., 1997)⇒ Luederwaldtinia rodrigoi Kohlmann & S..., 1997
Dichotomius (Luederwaldtinia) yucatanus Bates, 1887
Dichotomius annae Linnaeus, 1767
Dichotomius satanas Harold, 1867
Digitonthophagus gazella Fabricius, 1787Brown dung beetle
Diplotaxis denigrata Bates, 1889
Diplotaxis guatemalica Moser, 1918
Diplotaxis macrotarsus Vaurie, 1960
Diplotaxis mistura Vaurie, 1960
Diplotaxis poropyge Bates, 1887
Disphysema candezei Harold, 1873
Dorysthetus solisi Soula, 2002
Dynastes hercules Linnaeus, 1758Hercules beetle
Dynastes septentrionalis Lachaume, 1985
Dyscinetus dubius Olivier, 1789
Dyscinetus laevipunctatus Bates, 1888
Enema endymion Chevrolat, 1843Palehead blenny
Epectinaspis costaricensis Ramírez-Pon..., 2014
Euetheola bidentata Burmeister, 1847
Euetheola humilis Burmeister, 1847Spotted kelpfish
Euparixia costaricensis Hinton, 1936
Euphoria candezei Janson, 1875
Euphoria chontalensis Bates, 1889
Euphoria eximia Bates, 1889
Euphoria iridescens Schaum, 1841
Euphoria paradisiaca Orozco, 2012
Euphoria sepulcralis Fabricius, 1801Dark Flower Scarab
Eurysternus angustulus Harold, 1869
Eurysternus caribaeus Herbst, 1789
Eurysternus foedus Guérin-Ménev..., 1844
Eurysternus magnus Castelnau, 1840
Eurysternus mexicanus Harold, 1869
Eurysternus plebejus Harold, 1880
Eurysternus velutinus Bates, 1887
Eutrichillum arcus Solis & Kohl..., 2003
Faula brunneipennis Bates, 1887
Faula centralis Sharp, 1877
Geniates flaviventris Frey, 1976
Genuchinus nevermanni Schauer, 1935
Giesbertiolus ornatus Howden, 1988
Golofa hirsuta Ratcliffe, 2003
Golofa solisi Ratcliffe, 2003
Goniophileurus femoratus Burmeister, 1847
Guatemalica hueti Chevrolat, 1870
Gymnetis difficilis Burmeister, 1842
Helictopleurus neoamplicollis Krell, 2000
Heliocopris hamadryas Fabricius, 1775
Hemiphileurus dejeani Bates, 1888
Hemiphileurus simplex Prell, 1914
Heterogomphus chevrolati Burmeister, 1847
Heterogomphus mniszechi Thomson, 1859
Heterosternus oberthueri Ohaus, 1901
Homocopris achamas Harold, 1867
Homophileurus tricuspis Prell, 1914
Hoplia philanthus philanthus Fuesslin, 1775
Hoplia surata Bates, 1887
Hoplopyga liturata Olivier, 1789
Hoplopyga ocellata Gory & Perch..., 1833
Hypaspidius patrickarnaudi Soula, 2002
Inca clathratus Olivier, 1792
Isonychus chiriquinus Bates, 1887
Isonychus hirsutus Bates, 1887
Isonychus paradoxus Bates, 1887
Isonychus pictus Sharp, 1877
Isonychus vicinus Moser, 1918
Lagochile chiriquina Bates, 1888
Lagochile divisa Ohaus, 1914
Lagochile gingearis
Leucothyreus femoratus Burmeister, 1844
Liparetrus paryphus Britton, 1980
Luederwaldtinia danieli Kohlmann & S..., 1997
Luederwaldtinia femoratus Howden & Young, 1981
Luederwaldtinia rodrigoi Kohlmann & S..., 1997
Macraspis chrysis Linnaeus, 1764
Macraspis costaricensis Soula, 1998
Macraspis hirtiventris H. Bates, 1888
Macrodactylus costaricensis Moser, 1919
Macrodactylus costulatus Bates, 1887
Macrodactylus sericinus Bates, 1887
Macrodactylus solisi Arce-Pérez &..., 2018
Macrodactylus suavis Bates, 1887
Macrodactylus sylphis Bates, 1887
Macropoides cribricollis Ohaus, 1934
Megaceras septentrionis Bates, 1888
Megasoma elephas Fabricius, 1775Elephant beetle
Melinopterus prodromus Brahm, 1790Spring small dung beetle
Mimela schneideri Ohaus, 1905
Minidorysthetus iodiellus Bates, 1888
Minidorysthetus solisi Soula, 2002
Nialaphodius nigrita Fabricius, 1801
Nimbus contaminatus Herbst, 1783
Nimbus proximus Adám, 1994
Ontherus azteca Harold, 1869
Ontherus brevicollis Kirsch, 1870
Ontherus brevipennis Harold, 1867
Ontherus pseudodidymus Génier, 1996
Ontherus sextuberculatus Génier, 1996
Onthophagus (Furconthophagus) furcatus (Fabricius, 1781)⇒ Onthophagus furcaticeps Masters, 1886
Onthophagus (Onthophagus) batesi Howden and C..., 1963
Onthophagus (Onthophagus) cyanellus Bates, 1887
Onthophagus (Onthophagus) incensus Say, 1835Dung beetle
Onthophagus (Onthophagus) landolti (Harold, 1880)⇒ Onthophagus schaefferi Howden and C..., 1963
Onthophagus (Palaeonthophagus) fracticornis (Preyssler, 1790)⇒ Onthophagus similis Scriba, 1790
Onthophagus (Palaeonthophagus) similis (Scriba, 1790)⇒ Onthophagus fracticornis Preyssler, 1790
Onthophagus acuminatus (Harold, 1880)⇒ Onthophagus embrikianus Paulian, 1936
Onthophagus andersoni Howden & Gill, 1987
Onthophagus anthracinus (Harold, 1873)⇒ Caccobius histeroides Ménétriés, 1832
Onthophagus atriglabrus Howden & Gill, 1987
Onthophagus atrosericeus Boucomont, 1932
Onthophagus carpophilus Pereira & Ha..., 1961
Onthophagus championi Bates, 1887
Onthophagus chryses Bates, 1887
Onthophagus coscineus Bates, 1887
Onthophagus crinitus (Harold, 1869)⇒ Onthophagus criniger d´Orbigny, 1904
Onthophagus cryptodicranius Kohlmann & S..., 2001
Onthophagus gazellinus Bates, 1887
Onthophagus genuinus Kohlmann & S..., 2001
Onthophagus hopfneri
Onthophagus limonensis Kohlmann & S..., 2001
Onthophagus marginicollis Harold, 1880
Onthophagus maya Zunino, 1981
Onthophagus nemorivagus Kohlmann & S..., 2001
Onthophagus notiodes Solis & Kohl..., 2003
Onthophagus nyctopus Bates, 1887
Onthophagus orphnoides (Bates, 1887)⇒ Onthophagus mirabilis Bates, 1887
Onthophagus praecellens Bates, 1887
Onthophagus propraecellens Howden & Gill, 1987
Onthophagus quetzalis Howden & Gill, 1993
Onthophagus rhinolophus Harold, 1869
Onthophagus solisi Howden & Gill, 1993
Onthophagus stockwelli Howden & Young, 1956
Onthophagus tapirus Sharp, 1877
Onthophagus viridivinosus Kohlmann & S..., 2001
Parachlorota uhligi Soula, 2002
Parapucaya amazonica Prell, 1934
Pelidnota belti Sharp, 1877
Pelidnota costaricensis Bates, 1888
Pelidnota frommeri Hardy, 1975
Pelidnota notata (Blanchard, 1851)⇒ Pelidnota punctatus Linnaeus, 1758
Pelidnota parallela Hardy, 1975
Pelidnota prolixa Sharp, 1877
Pelidnota strigosa Laporte, 1840
Phalangogonia sperata Sharp, 1877
Phanaeus (Notiophanaeus) endymion Harold, 1863
Phanaeus (Notiophanaeus) pyrois Bates, 1887
Phanaeus beltianus Bates, 1887
Phanaeus eximius Bates, 1887
Phanaeus wagneri Harold, 1863
Phileurus angustatus Kolbe, 1910
Phileurus carinatus Prell, 1914
Phileurus didymus Linnaeus, 1758
Phileurus truncatus Palisot de B..., 1807Triceratops Beetle
Phileurus valgus Linnaeus, 1758
Phileurus youngi Ratcliffe, 1988
Phyllophaga (Phyllophaga) setifera Burmeister, 1855Highfin blenny
Phyllophaga (Phytalus) obsoleta Blanchard, 1851
Phyllophaga (Phytalus) pruinosa Blanchard, 1851
Phyllophaga aequata Bates, 1888
Phyllophaga boruca Morón, 2003
Phyllophaga changuena Morón, 2003
Phyllophaga cometes Bates, 1888
Phyllophaga coronadis Saylor, 1941
Phyllophaga costaricensis Moser, 1918
Phyllophaga crena Saylor, 1941
Phyllophaga densata Moser, 1918
Phyllophaga gigantea
Phyllophaga guanacasteca Moron & Solis, 2000
Phyllophaga guapiles Saylor, 1941
Phyllophaga guapilesea Saylor, 1941
Phyllophaga hemilissa Bates, 1888
Phyllophaga kohlmanniana Moron & Solis, 2000
Phyllophaga lorencita Moron & Solis, 2001
Phyllophaga naranjina Moron & Solis, 2001
Phyllophaga nevermannea Saylor, 1941
Phyllophaga nevermanni Saylor, 1935
Phyllophaga orosina Moser, 1918
Phyllophaga panamana Chapin, 1935
Phyllophaga picadoana Moron & Solis, 2000
Phyllophaga prolixa Bates, 1888
Phyllophaga puntarenosa Moron & Solis, 2001
Phyllophaga solisiana Morón, 2003
Phyllophaga tapantina Moron & Solis, 2001
Phyllophaga tilarana Moron & Solis, 2001
Phyllophaga valeriana (Saylor, 1934)⇒ Phyllophaga sanjosicola Saylor, 1935
Platycoelia grandicula Smith, 2003
Platycoelia humeralis Bates, 1888
Platycoelia mesosternalis Ohaus, 1904
Plectris nigrita Moser, 1918
Pseudocanthon perplexus LeConte, 1847
Ptenomela gratiosa Sharp, 1877
Ptenomela hirtiventris Ohaus, 1938
Pucaya castanea Ohaus, 1910
Rutela pygidialis Ohaus, 1905
Scatimus erinnyos Kohlmann & S..., 1996
Scatimus ovatus Harold, 1862
Serica acontia Dawson, 1933
Serica perigonia perigonia Dawson, 1920
Sisyphus crispatus Gory, 1833
Smaragdesthes africana smaragdina Chevrolat, 1836
Spodistes batesi Arrow, 1902
Spodistes beltianus Bates, 1888
Spodistes mniszechi Thomson, 1860
Spodochlamys cupreola Bates, 1888
Stenocrates bicarinatus Robinson, 1947
Stenocrates hardyi Dechambre, 1985
Stenocrates laevicollis Kirsch, 1870
Strategus aloeus Linnaeus, 1758Coconut cockle
Strategus jugurtha Burmeister, 1847
Strigoderma auriventris Bates, 1888
Strigoderma biolleyi Ohaus, 1908
Strigoderma castor Newman, 1838
Strigoderma marginata Olivier, 1789
Strigoderma micans Nonfried, 1893
Strigoderma orbicularis Burmeister, 1855
Strigoderma rutelina Bates, 1888
Strigoderma vestita Burmeister, 1844Cloister blenny
Sulcophanaeus noctis Bates, 1887
Sulcophanaeus velutinus Murray, 1856
Thyriochlorota jordani Ohaus, 1905
Tomarus bituberculatus Palisot de B..., 1805
Tomarus gyas Erichson, 1848
Tomarus nasutus Burmeister, 1847Spotted kelpfish
Tomarus sallaei Bates, 1888
Trichillidium pilosum Robinson, 1948
Trigonopeltastes geometricus Schaum, 1841
Trigonopeltastes sallaei Bates, 1889Burrowers
Uroxys boneti Pereira & Ha..., 1961
Uroxys coarctatus Harold, 1867
Uroxys deavilai Delgado & Ko..., 2007
Uroxys dybasi Howden & Young, 1981
Uroxys gorgon Arrow, 1933
Uroxys microcularis Howden & Young, 1981
Uroxys micros Bates, 1887
Uroxys pauliani Balthasar, 1940
Uroxys platypyga Howden & Young, 1981
Uroxys transversifrons Howden & Gill, 1987

Subnodes of Scarabaeidae with occurrences in Alajuela
SubfamilyAphodiinae
Leach, 1815
Aphodiine Dung Beetles, Small dung beetles
6
7
Genera
Species

Taxom

SubfamilyCetoniinae
Leach, 1815
Flower beetles, Flower chafers, Flower scarabs, Fruit and flower chafers
14
27
Genera
Species

Taxom

SubfamilyDynastinae
MacLeay, 1819
Horn beetles, Rhinoceros beetles
25
84
Genera
Species

Taxom

SubfamilyMelolonthinae
Samouelle, 1819
May Beetles and Junebugs, White grubs
13
56
Genera
Species

Taxom

SubfamilyOrphninae
Erichson, 1847
1
0
Genus
Species

Taxom

SubfamilyRutelinae
MacLeay, 1819
Shining Flower Chafers, Shining Leaf Chafers
29
110
Genera
Species

Taxom

SubfamilyScarabaeinae
Latreille, 1802
Dung Beetles, True dung beetles
30
142
Genera
Species

Taxom



Scarabaeidae (Latreille 1802)
Scarabaeidae (Latreille 1802)


Links, sources and further information
Scarabaeidae at artfakta.se
Scarabaeidae at dyntaxa.se
Scarabaeidae at gbif.org
Scarabaeidae at irmng.org
Scarabaeidae at itis.gov