Checklist for Family Scarabaeidae Latreille, 1802 (Dung beetles)
and region Upala (Alajuela, Costa Rica)
Class
| Insecta
Linnaeus, 1758
Insects, True insects | 274
2826
3696 | Families
Genera
Species |
Taxom
|
Subclass
| Pterygota
Lang, 1888
Winged insects | 273
2825
3696 | Families
Genera
Species |
Taxom
|
Infraclass
| Neoptera
van der Wulp, 1890
Modern Wing-folding insects | 264
2806
3672 | Families
Genera
Species |
Taxom
|
Superorder
| Holometabola
Heider, 1889
Holometabolous Insects | 210
2535
3390 | Families
Genera
Species |
Taxom
|
Order
| Coleoptera
Linnaeus, 1758
Beetles, Coleopterans | 48
724
850 | Families
Genera
Species |
Taxom
|
Suborder
| Polyphaga
Emery, 1886
Polyphags | 45
666
790 | Families
Genera
Species |
Taxom
|
Infraorder
| Scarabaeiformia
Crowson, 1960
| 6
97
229 | Families
Genera
Species |
Taxom
|
Superfamily
| Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles | 6
97
229 | Families
Genera
Species |
Taxom
| | Family | Scarabaeidae
Latreille, 1802
Dung beetles, Flower beetles, Fruit chafers, Rain beetles | 76
199 | Genera
Species |
Taxom
|
Genera for Upala
76 Genera with 199 Species #ABCDEGHILMOPSTU
Aegidium Blanchard, 1845 | | 0 | Species | Taxom | | Amblyoproctus Kolbe, 2003 | | 1 | Species | Taxom | | Amithao Thomson, 1848 | | 2 | Species | Taxom | | Ancognatha Erichson, 1911 | | 2 | Species | Taxom | | Anomala Samouelle, 1844 | Shining Leaf Chafer | 19 | Species | Taxom | | Anomalorhina Jameson, Pau..., 1928 | | 1 | Species | Taxom | | Anomiopus Westwood, 1939 | | 1 | Species | Taxom | | Aphodius Hellwig, 1789 | | 1 | Species | Taxom | | Archophileurus Kolbe, 1888 | | 1 | Species | Taxom | | Argyripa Thomson, 1869 | | 1 | Species | Taxom | | Aspidolea Bates, 1888 | | 3 | Species | Taxom | Ataenius Harold, 1867 | | 0 | Species | Taxom | | Ateuchus Weber, 1996 | | 2 | Species | Taxom | | Barybas Burmeister, 1887 | | 1 | Species | Taxom | | Bdelyrus Harold, 1887 | | 1 | Species | Taxom | | Bradypodidium Vaz-de-Mello, 1928 | | 1 | Species | Taxom | | Callistethus Blanchard, 1888 | | 9 | Species | Taxom | | Calomacraspis Bates, 1869 | | 1 | Species | Taxom | | Canthidium Erichson, 1981 | | 9 | Species | Taxom | | Canthon Hoffmannsegg, 1867 | Tumblebug | 3 | Species | Taxom | Ceraspis Serville, 1825 | | 0 | Species | Taxom | Chasmodia MacLeay, 1819 | | 0 | Species | Taxom | | Chlorota Burmeister, 1912 | | 2 | Species | Taxom | | Chrysina Kirby, 1913 | Jewel scarabs | 5 | Species | Taxom | | Cnemida Kirby, 1888 | | 2 | Species | Taxom | | Coelosis Hope, 1767 | | 1 | Species | Taxom | | Copris Geoffroy, 1858 | Mexican dung beetle | 2 | Species | Taxom | | Coprophanaeus Olsoufieff, 2008 | | 5 | Species | Taxom | | Cotinis Burmeister, 1876 | Green June Beetles | 2 | Species | Taxom | | Cyclocephala Dejean, 1992 | Masked Chafers | 24 | Species | Taxom | | Deltochilum Eschscholtz, 1873 | | 3 | Species | Taxom | | Dialithus Parry, 1849 | | 1 | Species | Taxom | | Dichotomius Hope, 1867 | | 3 | Species | Taxom | | Disphysema Harold, 1873 | | 1 | Species | Taxom | | Dorysthetus Blanchard, 2002 | | 1 | Species | Taxom | | Dynastes MacLeay, 1758 | Hercules Beetles | 1 | Species | Taxom | | Dyscinetus Harold, 1789 | Rice Beetles | 1 | Species | Taxom | | Enema Hope, 1843 | | 1 | Species | Taxom | | Euphoria Burmeister, 1801 | | 2 | Species | Taxom | | Eurysternus Dalman, 1880 | | 4 | Species | Taxom | | Eutrichillum | | 1 | Species | Taxom | | Genuchinus Westwood, 1935 | | 1 | Species | Taxom | | Giesbertiolus | | 1 | Species | Taxom | | Guatemalica Poll, 1870 | | 1 | Species | Taxom | Gymnetis MacLeay, 1819 | | 0 | Species | Taxom | | Hemiphileurus Kolbe, 1888 | | 1 | Species | Taxom | | Hoplopyga Thomson, 1789 | | 1 | Species | Taxom | | Inca Lepeletier &..., 1792 | | 1 | Species | Taxom | | Isonychus Mannerheim, 1918 | | 3 | Species | Taxom | | Lagochile Hoffmannsegg, 1914 | | 2 | Species | Taxom | | Luederwaldtinia Martínez, 1997 | | 1 | Species | Taxom | | Macrodactylus Dejean, 1887 | Rose Chafers | 5 | Species | Taxom | | Megasoma Kirby, 1775 | Elephant beetles | 1 | Species | Taxom | | Mimela Kirby, 1905 | | 1 | Species | Taxom | | Minidorysthetus | | 1 | Species | Taxom | | Ontherus Erichson, 1996 | | 1 | Species | Taxom | | Onthophagus Latreille, 1877 | | 16 | Species | Taxom | | Pelidnota MacLeay, 1877 | Precious Metal Scarabs | 4 | Species | Taxom | | Phalangogonia Burmeister, 1877 | | 1 | Species | Taxom | | Phanaeus MacLeay, 1887 | Burrowing Dung Beetles | 2 | Species | Taxom | | Phileurus Sieg, 1988 | | 2 | Species | Taxom | | Phyllophaga Harris, 1888 | June beetles | 10 | Species | Taxom | | Platycoelia Dejean, 1904 | | 1 | Species | Taxom | | Plectris Lepeletier a..., 1918 | | 1 | Species | Taxom | | Ptenomela Bates, 1938 | | 2 | Species | Taxom | | Scatimus Erichson, 1862 | | 2 | Species | Taxom | | Serica MacLeay, 1933 | | 1 | Species | Taxom | | Spodistes Burmeister, 1860 | | 1 | Species | Taxom | | Spodochlamys Burmeister, 1888 | | 1 | Species | Taxom | | Strategus Kirby, 1847 | Ox Beetles | 1 | Species | Taxom | | Strigoderma Burmeister, 1893 | | 3 | Species | Taxom | | Sulcophanaeus Olsoufieff, 1887 | | 1 | Species | Taxom | | Tomarus Erichson, 1848 | Carrot Beetles | 2 | Species | Taxom | | Trichillidium Vaz-de-Mello, 1948 | | 1 | Species | Taxom | Trigonopeltastes Burmeister &..., 1840 | | 0 | Species | Taxom | | Uroxys Westwood, 1981 | | 5 | Species | Taxom |
Species list for Upala
199 Species | Amblyoproctus centroamericanus Ratcliffe, 2003 | | | Amithao decemguttatus Waterhouse, 1876 | | | Amithao haematopus Schaum, 1848 | | | Ancognatha vexans Ratcliffe, 1992 | | | Ancognatha vulgaris Arrow, 1911 | | | Anomala calligrapha Bates, 1888 | | | Anomala chiriquina Bates, 1888 | | | Anomala clathrata Ohaus, 1930 | | | Anomala eulissa Bates, 1888 | | | Anomala levicollis Filippini, M..., 2015 | | | Anomala longisacculata Filippini, M..., 2015 | | | Anomala pincelada Filippini, G..., 2015 | | | Anomala praecellens Bates, 1888 | | | Anomala pseudoeucoma Filippini, M..., 2013 | | | Anomala quiche Ohaus, 1897 | | | Anomala semicincta Bates, 1888 | | | Anomala semilla Filippini, G..., 2014 | | | Anomala solisi Filippini, G..., 2014 | | | Anomala strigodermoides Filippini, G..., 2015 | | | Anomala sulana Zorn, 2006 | | | Anomala tenoriensis Filippini, M..., 2015 | | | Anomala testaceipennis (Blanchard, 1851) | ⇒ Anomala nitidula Blanchard, 1851 | | Anomala tuberculata Filippini, M..., 2015 | | | Anomala valida (Burmeister, 1844) | ⇒ Anomala insularis Castelnau, 1840 | | Anomalorhina turrialbana Ohaus, 1928 | | | Anomiopus panamensis Paulian, 1939 | | | Aphodius (Nialus) lividus (Olivier, 1789) | ⇒ Acrossus luridus Fabricius, 1775 | | Archophileurus simplex Bates, 1888 | | | Argyripa anomala Bates, 1869 | | | Aspidolea kuntzeni Höhne, 1922 | | | Aspidolea notaticollis Höhne, 1922 | | | Aspidolea singularis Bates, 1888 | | | Ateuchus fetteri Kohlmann, 1996 | | | Ateuchus zoebischi Kohlmann, 1996 | | | Barybas idiota Bates, 1887 | | | Bdelyrus seminudus Bates, 1887 | | | Bradypodidium bradyporum Boucomont, 1928 | | | Callistethus chontalensis Bates, 1888 | | | Callistethus lativittis Filippini, G..., 2015 | | | Callistethus macroxantholeus Filippini, G..., 2015 | | | Callistethus microxantholeus Filippini, G..., 2015 | | | Callistethus mimeloides Ohaus, 1902 | | | Callistethus multiplicatus Filippini, G..., 2015 | | | Callistethus pseudocollaris Filippini, G..., 2015 | | | Callistethus ruteloides Filippini, G..., 2015 | | | Callistethus sulcans Bates, 1888 | | | Calomacraspis haroldi Candeze, 1869 | | | Canthidium angusticeps Bates, 1887 | | | Canthidium annagabrielae Solis & Kohl..., 2004 | | | Canthidium ardens Bates, 1887 | | | Canthidium centrale Boucomont, 1928 | | | Canthidium discopygidiale Howden & Young, 1981 | | | Canthidium haroldi Preudhomme d..., 1886 | | | Canthidium hespenheidei Howden & Young, 1981 | | | Canthidium tuberifrons Howden & Young, 1981 | | | Canthidium vespertinum Howden & Young, 1981 | | | Canthon angustatus Harold, 1867 | | | Canthon silvaticus Solis & Kohl..., 2002 | | | Canthon subhyalinus Harold, 1867 | | | Chlorota batesi Soula, 2002 | | | Chlorota panamensis Ohaus, 1912 | | | Chrysina aurigans Rothschildt ..., 1894 | | | Chrysina aurora Boucard, 1875 | | | Chrysina bruyeai Hawks, 1999 | | | Chrysina chrysargyrea Sallé, 1874 | | | Chrysina luteomarginata Ohaus, 1913 | | | Cnemida aterrima Bates, 1888 | | | Cnemida intermedia Bates, 1888 | | | Coelosis biloba Linnaeus, 1767 | | | Copris incertus Say, 1835 | Black dung beetle | | Copris lugubris Boheman, 1858 | | | Coprophanaeus boucardi Nevinson, 1891 | | | Coprophanaeus kohlmanni (Arnaud, 2002) | ⇒ Coprophanaeus morenoi Arnaud, 1982 | | Coprophanaeus pecki Howden & Young, 1981 | | | Coprophanaeus telamon Erichson, 1847 | | | Coprophanaeus uhleri (Maly & Pokorny, 2008) | ⇒ Coprophanaeus chiriquensis Olsufieff, 1924 | | Cotinis lebasi (Gory & Perch..., 1833) | ⇒ Cotinorrhina lebasi Gory & Perch..., 1833 | | Cotinis polita Janson, 1876 | | | Cyclocephala almitana Dechambre, 1992 | | | Cyclocephala amblyopsis Hurley, 1959 | | | Cyclocephala carbonaria Milne Edwards, 1848 | | | Cyclocephala complanata Burmeister, 1847 | | | Cyclocephala concolor Burmeister, 1847 | | | Cyclocephala confusa Endrödi, 1966 | | | Cyclocephala conspicua Sharp, 1877 | | | Cyclocephala discolor Herbst, 1792 | | | Cyclocephala erotylina Arrow, 1914 | | | Cyclocephala fasciolata Bates, 1888 | | | Cyclocephala fulgurata Burmeister, 1847 | | | Cyclocephala gravis Bates, 1888 | | | Cyclocephala kaszabi Endrödi, 1964 | | | Cyclocephala ligyrina Bates, 1888 | | | Cyclocephala lunulata Burmeister, 1847 | Twister | | Cyclocephala maculiventris Höhne, 1923 | | | Cyclocephala mafaffa Burmeister, 1847 | Twister | | Cyclocephala melanocephala (Fabricius, 1775) | ⇒ Cyclocephala olivieri Arrow, 1911 | | Cyclocephala multiplex (Casey, 1915) | ⇒ Cyclocephala amazona Linnaeus, 1767 | | Cyclocephala ovulum (Bates, 1888) | ⇒ Cyclocephala testacea Burmeister, 1847 | | Cyclocephala porioni Dechambre, 1979 | | | Cyclocephala sexpunctata Castelnau, 1840 | | | Cyclocephala weidneri Endrödi, 1964 | | | Cyclocephala williami Ratcliffe, 1992 | | | Deltochilum gibbosum Fabricius, 1775 | Humpback Dung Beetle | | Deltochilum pseudoparile Paulian, 1938 | | | Deltochilum valgum Burmeister, 1873 | | | Dialithus magnificus Parry, 1849 | | | Dichotomius (Luederwaldtinia) danieli (Kohlmann & S..., 1997) | ⇒ Luederwaldtinia danieli Kohlmann & S..., 1997 | | Dichotomius annae Linnaeus, 1767 | | | Dichotomius satanas Harold, 1867 | | | Disphysema candezei Harold, 1873 | | | Dorysthetus solisi Soula, 2002 | | | Dynastes hercules Linnaeus, 1758 | Hercules beetle | | Dyscinetus dubius Olivier, 1789 | | | Enema endymion Chevrolat, 1843 | Palehead blenny | | Euphoria candezei Janson, 1875 | | | Euphoria sepulcralis Fabricius, 1801 | Dark Flower Scarab | | Eurysternus caribaeus Herbst, 1789 | | | Eurysternus magnus Castelnau, 1840 | | | Eurysternus mexicanus Harold, 1869 | | | Eurysternus plebejus Harold, 1880 | | | Eutrichillum arcus Solis & Kohl..., 2003 | | | Genuchinus nevermanni Schauer, 1935 | | | Giesbertiolus ornatus Howden, 1988 | | | Guatemalica hueti Chevrolat, 1870 | | | Hemiphileurus dejeani Bates, 1888 | | | Hoplopyga liturata Olivier, 1789 | | | Inca clathratus Olivier, 1792 | | | Isonychus chiriquinus Bates, 1887 | | | Isonychus pictus Sharp, 1877 | | | Isonychus vicinus Moser, 1918 | | | Lagochile chiriquina Bates, 1888 | | | Lagochile divisa Ohaus, 1914 | | | Luederwaldtinia danieli Kohlmann & S..., 1997 | | | Macrodactylus costaricensis Moser, 1919 | | | Macrodactylus costulatus Bates, 1887 | | | Macrodactylus sericinus Bates, 1887 | | | Macrodactylus suavis Bates, 1887 | | | Macrodactylus sylphis Bates, 1887 | | | Megasoma elephas Fabricius, 1775 | Elephant beetle | | Mimela schneideri Ohaus, 1905 | | | Minidorysthetus solisi Soula, 2002 | | | Ontherus pseudodidymus Génier, 1996 | | | Onthophagus (Onthophagus) batesi Howden and C..., 1963 | | | Onthophagus (Onthophagus) incensus Say, 1835 | Dung beetle | | Onthophagus (Onthophagus) landolti (Harold, 1880) | ⇒ Onthophagus schaefferi Howden and C..., 1963 | | Onthophagus acuminatus (Harold, 1880) | ⇒ Onthophagus embrikianus Paulian, 1936 | | Onthophagus carpophilus Pereira & Ha..., 1961 | | | Onthophagus chryses Bates, 1887 | | | Onthophagus coscineus Bates, 1887 | | | Onthophagus crinitus (Harold, 1869) | ⇒ Onthophagus criniger d´Orbigny, 1904 | | Onthophagus gazellinus Bates, 1887 | | | Onthophagus genuinus Kohlmann & S..., 2001 | | | Onthophagus nemorivagus Kohlmann & S..., 2001 | | | Onthophagus nyctopus Bates, 1887 | | | Onthophagus praecellens Bates, 1887 | | | Onthophagus solisi Howden & Gill, 1993 | | | Onthophagus stockwelli Howden & Young, 1956 | | | Onthophagus tapirus Sharp, 1877 | | | Pelidnota frommeri Hardy, 1975 | | | Pelidnota notata (Blanchard, 1851) | ⇒ Pelidnota punctatus Linnaeus, 1758 | | Pelidnota parallela Hardy, 1975 | | | Pelidnota prolixa Sharp, 1877 | | | Phalangogonia sperata Sharp, 1877 | | | Phanaeus (Notiophanaeus) pyrois Bates, 1887 | | | Phanaeus beltianus Bates, 1887 | | | Phileurus didymus Linnaeus, 1758 | | | Phileurus youngi Ratcliffe, 1988 | | | Phyllophaga (Phytalus) obsoleta Blanchard, 1851 | | | Phyllophaga boruca Morón, 2003 | | | Phyllophaga gigantea | | | Phyllophaga guanacasteca Moron & Solis, 2000 | | | Phyllophaga guapiles Saylor, 1941 | | | Phyllophaga guapilesea Saylor, 1941 | | | Phyllophaga nevermanni Saylor, 1935 | | | Phyllophaga orosina Moser, 1918 | | | Phyllophaga panamana Chapin, 1935 | | | Phyllophaga prolixa Bates, 1888 | | | Platycoelia mesosternalis Ohaus, 1904 | | | Plectris nigrita Moser, 1918 | | | Ptenomela gratiosa Sharp, 1877 | | | Ptenomela hirtiventris Ohaus, 1938 | | | Scatimus erinnyos Kohlmann & S..., 1996 | | | Scatimus ovatus Harold, 1862 | | | Serica acontia Dawson, 1933 | | | Spodistes mniszechi Thomson, 1860 | | | Spodochlamys cupreola Bates, 1888 | | | Strategus jugurtha Burmeister, 1847 | | | Strigoderma auriventris Bates, 1888 | | | Strigoderma marginata Olivier, 1789 | | | Strigoderma micans Nonfried, 1893 | | | Sulcophanaeus noctis Bates, 1887 | | | Tomarus bituberculatus Palisot de B..., 1805 | | | Tomarus gyas Erichson, 1848 | | | Trichillidium pilosum Robinson, 1948 | | | Uroxys boneti Pereira & Ha..., 1961 | | | Uroxys dybasi Howden & Young, 1981 | | | Uroxys gorgon Arrow, 1933 | | | Uroxys pauliani Balthasar, 1940 | | | Uroxys platypyga Howden & Young, 1981 | |
Subnodes of Scarabaeidae with occurrences in Upala | Subfamily | Aphodiinae
Leach, 1815
Aphodiine Dung Beetles, Small dung beetles | 2
1 | Genera
Species |
Taxom
| | Subfamily | Cetoniinae
Leach, 1815
Flower beetles, Flower chafers, Flower scarabs, Fruit and flower chafers | 12
13 | Genera
Species |
Taxom
| | Subfamily | Dynastinae
MacLeay, 1819
Horn beetles, Rhinoceros beetles | 15
43 | Genera
Species |
Taxom
| | Subfamily | Melolonthinae
Samouelle, 1819
May Beetles and Junebugs, White grubs | 7
21 | Genera
Species |
Taxom
| | Subfamily | Orphninae
Erichson, 1847
| 1
0 | Genus
Species |
Taxom
| | Subfamily | Rutelinae
MacLeay, 1819
Shining Flower Chafers, Shining Leaf Chafers | 18
56 | Genera
Species |
Taxom
| | Subfamily | Scarabaeinae
Latreille, 1802
Dung Beetles, True dung beetles | 20
64 | Genera
Species |
Taxom
|
 | | Scarabaeidae (Latreille 1802) |
Links, sources and further information Scarabaeidae at artfakta.se
Scarabaeidae at dyntaxa.se
Scarabaeidae at gbif.org
Scarabaeidae at irmng.org
Scarabaeidae at itis.gov
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