Checklist for Subfamily Scarabaeinae Latreille, 1802 (Dung Beetles)
and region Upala (Alajuela, Costa Rica)
Class
| Insecta
Linnaeus, 1758
Insects, True insects | 274
2826
3696 | Families
Genera
Species |
Taxom
|
Subclass
| Pterygota
Lang, 1888
Winged insects | 273
2825
3696 | Families
Genera
Species |
Taxom
|
Infraclass
| Neoptera
van der Wulp, 1890
Modern Wing-folding insects | 264
2806
3672 | Families
Genera
Species |
Taxom
|
Superorder
| Holometabola
Heider, 1889
Holometabolous Insects | 210
2535
3390 | Families
Genera
Species |
Taxom
|
Order
| Coleoptera
Linnaeus, 1758
Beetles, Coleopterans | 48
724
850 | Families
Genera
Species |
Taxom
|
Suborder
| Polyphaga
Emery, 1886
Polyphags | 45
666
790 | Families
Genera
Species |
Taxom
|
Infraorder
| Scarabaeiformia
Crowson, 1960
| 6
97
229 | Families
Genera
Species |
Taxom
|
Superfamily
| Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles | 6
97
229 | Families
Genera
Species |
Taxom
| | Family | Scarabaeidae
Latreille, 1802
Dung beetles, Flower beetles, Fruit chafers, Rain beetles | 76
199 | Genera
Species |
Taxom
| | Subfamily | Scarabaeinae
Latreille, 1802
Dung Beetles, True dung beetles | 20
64 | Genera
Species |
Taxom
|
Genera for Upala
20 Genera with 64 Species | Anomiopus Westwood, 1939 | | 1 | Species | Taxom | | Ateuchus Weber, 1996 | | 2 | Species | Taxom | | Bdelyrus Harold, 1887 | | 1 | Species | Taxom | | Bradypodidium Vaz-de-Mello, 1928 | | 1 | Species | Taxom | | Canthidium Erichson, 1981 | | 9 | Species | Taxom | | Canthon Hoffmannsegg, 1867 | Tumblebug | 3 | Species | Taxom | | Copris Geoffroy, 1858 | Mexican dung beetle | 2 | Species | Taxom | | Coprophanaeus Olsoufieff, 2008 | | 5 | Species | Taxom | | Deltochilum Eschscholtz, 1873 | | 3 | Species | Taxom | | Dichotomius Hope, 1867 | | 3 | Species | Taxom | | Disphysema Harold, 1873 | | 1 | Species | Taxom | | Eurysternus Dalman, 1880 | | 4 | Species | Taxom | | Eutrichillum | | 1 | Species | Taxom | | Ontherus Erichson, 1996 | | 1 | Species | Taxom | | Onthophagus Latreille, 1877 | | 16 | Species | Taxom | | Phanaeus MacLeay, 1887 | Burrowing Dung Beetles | 2 | Species | Taxom | | Scatimus Erichson, 1862 | | 2 | Species | Taxom | | Sulcophanaeus Olsoufieff, 1887 | | 1 | Species | Taxom | | Trichillidium Vaz-de-Mello, 1948 | | 1 | Species | Taxom | | Uroxys Westwood, 1981 | | 5 | Species | Taxom |
Species list for Upala
64 Species | Anomiopus panamensis Paulian, 1939 | | | Ateuchus fetteri Kohlmann, 1996 | | | Ateuchus zoebischi Kohlmann, 1996 | | | Bdelyrus seminudus Bates, 1887 | | | Bradypodidium bradyporum Boucomont, 1928 | | | Canthidium angusticeps Bates, 1887 | | | Canthidium annagabrielae Solis & Kohl..., 2004 | | | Canthidium ardens Bates, 1887 | | | Canthidium centrale Boucomont, 1928 | | | Canthidium discopygidiale Howden & Young, 1981 | | | Canthidium haroldi Preudhomme d..., 1886 | | | Canthidium hespenheidei Howden & Young, 1981 | | | Canthidium tuberifrons Howden & Young, 1981 | | | Canthidium vespertinum Howden & Young, 1981 | | | Canthon angustatus Harold, 1867 | | | Canthon silvaticus Solis & Kohl..., 2002 | | | Canthon subhyalinus Harold, 1867 | | | Copris incertus Say, 1835 | Black dung beetle | | Copris lugubris Boheman, 1858 | | | Coprophanaeus boucardi Nevinson, 1891 | | | Coprophanaeus kohlmanni (Arnaud, 2002) | ⇒ Coprophanaeus morenoi Arnaud, 1982 | | Coprophanaeus pecki Howden & Young, 1981 | | | Coprophanaeus telamon Erichson, 1847 | | | Coprophanaeus uhleri (Maly & Pokorny, 2008) | ⇒ Coprophanaeus chiriquensis Olsufieff, 1924 | | Deltochilum gibbosum Fabricius, 1775 | Humpback Dung Beetle | | Deltochilum pseudoparile Paulian, 1938 | | | Deltochilum valgum Burmeister, 1873 | | | Dichotomius (Luederwaldtinia) danieli (Kohlmann & S..., 1997) | ⇒ Luederwaldtinia danieli Kohlmann & S..., 1997 | | Dichotomius annae Linnaeus, 1767 | | | Dichotomius satanas Harold, 1867 | | | Disphysema candezei Harold, 1873 | | | Eurysternus caribaeus Herbst, 1789 | | | Eurysternus magnus Castelnau, 1840 | | | Eurysternus mexicanus Harold, 1869 | | | Eurysternus plebejus Harold, 1880 | | | Eutrichillum arcus Solis & Kohl..., 2003 | | | Ontherus pseudodidymus Génier, 1996 | | | Onthophagus (Onthophagus) batesi Howden and C..., 1963 | | | Onthophagus (Onthophagus) incensus Say, 1835 | Dung beetle | | Onthophagus (Onthophagus) landolti (Harold, 1880) | ⇒ Onthophagus schaefferi Howden and C..., 1963 | | Onthophagus acuminatus (Harold, 1880) | ⇒ Onthophagus embrikianus Paulian, 1936 | | Onthophagus carpophilus Pereira & Ha..., 1961 | | | Onthophagus chryses Bates, 1887 | | | Onthophagus coscineus Bates, 1887 | | | Onthophagus crinitus (Harold, 1869) | ⇒ Onthophagus criniger d´Orbigny, 1904 | | Onthophagus gazellinus Bates, 1887 | | | Onthophagus genuinus Kohlmann & S..., 2001 | | | Onthophagus nemorivagus Kohlmann & S..., 2001 | | | Onthophagus nyctopus Bates, 1887 | | | Onthophagus praecellens Bates, 1887 | | | Onthophagus solisi Howden & Gill, 1993 | | | Onthophagus stockwelli Howden & Young, 1956 | | | Onthophagus tapirus Sharp, 1877 | | | Phanaeus (Notiophanaeus) pyrois Bates, 1887 | | | Phanaeus beltianus Bates, 1887 | | | Scatimus erinnyos Kohlmann & S..., 1996 | | | Scatimus ovatus Harold, 1862 | | | Sulcophanaeus noctis Bates, 1887 | | | Trichillidium pilosum Robinson, 1948 | | | Uroxys boneti Pereira & Ha..., 1961 | | | Uroxys dybasi Howden & Young, 1981 | | | Uroxys gorgon Arrow, 1933 | | | Uroxys pauliani Balthasar, 1940 | | | Uroxys platypyga Howden & Young, 1981 | |
Subnodes of Scarabaeinae with occurrences in Upala
Links, sources and further information Scarabaeinae at artfakta.se
Scarabaeinae at dyntaxa.se
Scarabaeinae at itis.gov
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