Checklist for Subfamily Scarabaeinae Latreille, 1802 (Dung Beetles)
and region Free State (Republic South Africa)
Class
| Insecta
Linnaeus, 1758
Insects, True insects | 283
1882
3253 | Families
Genera
Species |
Taxom
|
Subclass
| Pterygota
Lang, 1888
Winged insects | 282
1879
3243 | Families
Genera
Species |
Taxom
|
Infraclass
| Neoptera
van der Wulp, 1890
Modern Wing-folding insects | 265
1827
3170 | Families
Genera
Species |
Taxom
|
Superorder
| Holometabola
Heider, 1889
Holometabolous Insects | 163
1308
2273 | Families
Genera
Species |
Taxom
|
Order
| Coleoptera
Linnaeus, 1758
Beetles, Coleopterans | 39
413
714 | Families
Genera
Species |
Taxom
|
Suborder
| Polyphaga
Emery, 1886
Polyphags | 35
347
621 | Families
Genera
Species |
Taxom
|
Infraorder
| Scarabaeiformia
Crowson, 1960
| 5
135
359 | Families
Genera
Species |
Taxom
|
Superfamily
| Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles | 5
135
359 | Families
Genera
Species |
Taxom
| | Family | Scarabaeidae
Latreille, 1802
Dung beetles, Flower beetles, Fruit chafers, Rain beetles | 124
317 | Genera
Species |
Taxom
| | Subfamily | Scarabaeinae
Latreille, 1802
Dung Beetles, True dung beetles | 37
178 | Genera
Species |
Taxom
|
Genera for Free State
37 Genera with 178 Species #ACDEGHKLMNOPSTX
| Allogymnopleurus Janssens, 1855 | | 3 | Species | Taxom | | Caccobius Thomson, 1857 | | 3 | Species | Taxom | | Catharsius Hope, 1857 | | 5 | Species | Taxom | | Cheironitis van Lansberge, 1776 | | 2 | Species | Taxom | | Cleptocaccobius Cambefort, 1857 | | 1 | Species | Taxom | | Copris Geoffroy, 1875 | Mexican dung beetle | 18 | Species | Taxom | | Coptorhina Hope, 1833 | | 2 | Species | Taxom | | Digitonthophagus Balthasar, 1787 | | 1 | Species | Taxom | | Drepanocerus Kirby, 1923 | | 2 | Species | Taxom | | Eodrepanus Barbero, Pal..., 1901 | | 1 | Species | Taxom | | Epirinus Reiche, 1901 | | 7 | Species | Taxom | | Euoniticellus Janssens, 1873 | | 2 | Species | Taxom | | Euonthophagus Balthasar, 1855 | | 1 | Species | Taxom | | Garreta Janssens, 1857 | | 2 | Species | Taxom | | Gymnopleurus Illiger, 1843 | | 4 | Species | Taxom | | Heliocopris Hope, 1798 | | 5 | Species | Taxom | | Kheper Janssens, 1857 | | 2 | Species | Taxom | | Liatongus Reitter, 1840 | | 1 | Species | Taxom | | Litocopris Waterhouse, 1869 | | 1 | Species | Taxom | | Metacatharsius Paulian, 1857 | | 5 | Species | Taxom | | Neonitis Péringuey, 1918 | | 1 | Species | Taxom | | Neosisyphus Müller, 1818 | | 7 | Species | Taxom | | Odontoloma Boheman, 1868 | | 2 | Species | Taxom | | Oniticellus Serville, 1840 | | 3 | Species | Taxom | | Onitis Fabricius, 1857 | | 13 | Species | Taxom | | Onthophagus Latreille, 1843 | | 43 | Species | Taxom | | Pachylomera Griffith & P..., 1828 | | 1 | Species | Taxom | | Pedaria Laporte, 1857 | | 1 | Species | Taxom | | Phalops Erichson, 1867 | | 5 | Species | Taxom | | Proagoderus Lansberge, 1855 | | 4 | Species | Taxom | | Sarophorus Erichson, 1857 | | 1 | Species | Taxom | | Scarabaeus Linnaeus, 1940 | | 20 | Species | Taxom | | Sceliages Westwood, 1828 | | 1 | Species | Taxom | | Sisyphus Latreille, 1818 | | 8 | Species | Taxom | | Stiptopodius Harold, 1923 | | 1 | Species | Taxom | | Tiniocellus Péringuey, 2010 | | 1 | Species | Taxom | | Xinidium Harold, 1869 | | 1 | Species | Taxom |
Species list for Free State
178 Species and 4 Subspecies | Allogymnopleurus consocius Péringuey, 1901 | | | Allogymnopleurus splendidus Bertoloni, 1849 | | | Allogymnopleurus thalassinus Klug, 1855 | | | Caccobius ferrugineus Fahraeus, 1857 | | | Caccobius histerinus Fahraeus, 1857 | | | Caccobius obtusus Fahraeus, 1857 | | | Catharsius marcellus Kolbe, 1893 | | | Catharsius sesostris Waterhouse, 1888 | | | Catharsius tricornutus DeGeer, 1778 | | | Catharsius ulysses Boheman, 1857 | | | Catharsius vitulus Boheman, 1857 | | | Cheironitis hoplosternus Harold, 1868 | | | Cheironitis scabrosus Fabricius, 1776 | | | Cleptocaccobius viridicollis Fahraeus, 1857 | | | Copris antares Bacescu & Ca..., 1947 | | | Copris caelatus Fabricius, 1794 | | | Copris cassius Péringuey, 1901 | | | Copris cornifrons Boheman, 1860 | | | Copris corniger (Sahlberg, 1823) | ⇒ Onthophagus taurus Schreber, 1759 | | Copris denticulatus Nguyen-phung, 1988 | | | Copris elphenor Klug, 1855 | | | Copris evanidus Klug, 1855 | | | Copris fallaciosus Gillet, 1907 | Kenyan dung beetle | | Copris fidius Olivier, 1789 | | | Copris inhalatus Quedenfeldt, 1884 | | | Copris inhalatus perturbator Péringuey, 1901 | | | Copris jacchoides Nguyen-phung..., 1987 | | | Copris jacchus Lang, 1971 | | | Copris mesacanthus Harold, 1878 | | | Copris mesacanthus transvaalensis Nguyen-phung, 1988 | | | Copris obesus Boheman, 1857 | | | Copris ritsemae Harold, 1875 | | | Coptorhina excavata Frolov, Akhm..., 2008 | | | Coptorhina klugii Hope, 1833 | | | Digitonthophagus gazella Fabricius, 1787 | Brown dung beetle | | Drepanocerus kirbyi Kirby, 1828 | | | Drepanocerus patrizii Boucomont, 1923 | | | Eodrepanus fastiditus Péringuey, 1901 | | | Epirinus asper Péringuey, 1901 | | | Epirinus flagellatus Fabricius, 1775 | | | Epirinus gratus Péringuey, 1901 | | | Epirinus obtusus Boheman, 1857 | | | Epirinus striatus Scholtz & Ho..., 1987 | | | Epirinus sulcipennis Boheman, 1857 | | | Epirinus validus Péringuey, 1901 | | | Euoniticellus africanus Harold, 1873 | Greater sandy dung beetle | | Euoniticellus triangulatus Harold, 1873 | | | Euonthophagus carbonarius Klug, 1855 | | | Garreta nitens Olivier, 1789 | | | Garreta unicolor Fahraeus, 1857 | | | Gymnopleurus aenescens Wiedemann, 1821 | | | Gymnopleurus humanus MacLeay, 1821 | | | Gymnopleurus leei Fabricius, 1792 | | | Gymnopleurus virens Erichson, 1843 | | | Heliocopris andersoni Bates, 1868 | | | Heliocopris faunus Boheman, 1857 | | | Heliocopris hamadryas Fabricius, 1775 | | | Heliocopris neptunus Boheman, 1857 | | | Heliocopris pirmal Fabricius, 1798 | | | Kheper lamarcki MacLeay, 1821 | | | Kheper nigroaeneus Boheman, 1857 | | | Liatongus militaris Castelnau, 1840 | Yellow Shouldered Dung Beetle | | Litocopris simplex Harold, 1869 | | | Metacatharsius latifrons Harold, 1868 | | | Metacatharsius marani Balthasar, 1940 | | | Metacatharsius pumilionis Wallengren, 1881 | | | Metacatharsius pusio Kolbe, 1908 | | | Metacatharsius troglodytes (Boheman, 1857) | ⇒ Metacatharsius anderseni Waterhouse, 1891 | | Neonitis nigritiae Gillet, 1918 | | | Neosisyphus barbarossa Wiedemann, 1823 | | | Neosisyphus calcaratus Klug, 1855 | | | Neosisyphus macrorubrus Paschalidis, 1974 | | | Neosisyphus rubripes Péringuey, 1901 | | | Neosisyphus rubrus Paschalidis, 1974 | | | Neosisyphus rugosus Gory, 1833 | | | Neosisyphus spinipes Thunberg, 1818 | Grey dungball roller | | Odontoloma louwi Howden & Sch..., 1987 | | | Odontoloma sculpturatum Harold, 1868 | | | Oniticellus formosus (Chevrolat, 1830) | ⇒ Oniticellus pictus Hausmann, 1807 | | Oniticellus pictus Hausmann, 1807 | | | Oniticellus planatus Castelnau, 1840 | | | Onitis alexis (Klug, 1835) | ⇒ Cheironitis ungaricus ungaricus Herbst, 1789 | | Onitis alexis alexis | | | Onitis aygulus Fabricius, 1781 | | | Onitis caffer Boheman, 1857 | | | Onitis confusus Boheman, 1860 | | | Onitis curvipes Lansberge, 1875 | | | Onitis parainflaticollis Ferreira, 1977 | | | Onitis pecuarius Lansberge, 1875 | | | Onitis perpunctatus Balthasar, 1963 | | | Onitis picticollis Boheman, 1857 | | | Onitis tortuosus Houston, 1983 | | | Onitis uncinatus Klug, 1855 | | | Onitis viridulus (Boheman, 1857) | ⇒ Onitis pecuarius Lansberge, 1875 | | Onthophagus (Afrostrandius) plebejus Klug, 1855 | | | Onthophagus (Proagoderus) pegesimallus Marcus, 1917 | | | Onthophagus absyrtus Balthasar, 1946 | | | Onthophagus aeruginosus Roth, 1851 | | | Onthophagus binodis Thunberg, 1818 | Humpbacked Dung Beetle | | Onthophagus bovinus Péringuey, 1892 | | | Onthophagus cyaneoniger d´Orbigny, 1902 | | | Onthophagus deterrens Péringuey, 1901 | | | Onthophagus ebenus Péringuey, 1888 | | | Onthophagus fimetarius Roth, 1851 | | | Onthophagus flavimargo d´Orbigny, 1902 | | | Onthophagus fritschi d´Orbigny, 1902 | | | Onthophagus granulifer (Harold, 1886) | ⇒ Onthophagus ponticus Harold, 1883 | | Onthophagus hyaena Fabricius, 1801 | | | Onthophagus impressicollis Boheman, 1860 | | | Onthophagus incisus Harold, 1877 | | | Onthophagus interstitialis (Fahraeus, 1857) | ⇒ Euonthophagus carbonarius Klug, 1855 | | Onthophagus lamnifer d´Orbigny, 1902 | | | Onthophagus leucopygus (Harold, 1867) | ⇒ Onthophagus albipodex d´Orbigny, 1902 | | Onthophagus lugens (Fahraeus, 1857) | ⇒ Onthophagus fimetarius Roth, 1851 | | Onthophagus lugubris (Fahraeus, 1857) | ⇒ Onthophagus lituratus Roth, 1851 | | Onthophagus modestus Harold, 1862 | | | Onthophagus monodon Fahraeus, 1857 | | | Onthophagus naso Fahraeus, 1857 | | | Onthophagus obtusicornis (Fahraeus, 1857) | ⇒ Onthophagus marginalis przewalski Kabakov, 2006 | | Onthophagus obtutus Péringuey, 1901 | | | Onthophagus pallidipennis (Fahraeus, 1857) | ⇒ Onthophagus rasipennis d´Orbigny, 1908 | | Onthophagus parumnotatus Fahraeus, 1857 | | | Onthophagus peringueyi Shipp, 1895 | | | Onthophagus pilosus (Fahraeus, 1857) | ⇒ Onthophagus echinus Boucomont, 1914 | | Onthophagus pugionatus Fahraeus, 1857 | | | Onthophagus pullus (Roth, 1851) | ⇒ Onthophagus parenthesis Boucomont, 1913 | | Onthophagus quadraticeps Harold, 1867 | | | Onthophagus rasipennis d´Orbigny, 1908 | | | Onthophagus rubens d´Orbigny, 1902 | | | Onthophagus setosus Fahraeus, 1857 | | | Onthophagus stellio Erichson, 1843 | | | Onthophagus sugillatus Klug, 1855 | | | Onthophagus suturalis (Péringuey, 1888) | ⇒ Onthophagus trituber Wiedemann, 1823 | | Onthophagus variegatus (Fabricius, 1798) | ⇒ Onthophagus filicornis Harold, 1873 | | Onthophagus variolosus d´Orbigny, 1902 | | | Onthophagus verticalis (Fahraeus, 1857) | ⇒ Onthophagus praedatus Harold, 1862 | | Onthophagus vinctus Erichson, 1843 | | | Pachylomera femoralis Kirby, 1828 | | | Pedaria picea Fahraeus, 1857 | | | Phalops bubalus Harold, 1867 | | | Phalops dregei Harold, 1867 | | | Phalops flavocinctus Klug, 1855 | | | Phalops rufosignatus (Lansberge, 1885) | ⇒ Phalops dregei Harold, 1867 | | Phalops wittei Harold, 1867 | | | Proagoderus brucei brucei | | | Proagoderus chalcostolus d´Orbigny, 1902 | | | Proagoderus lanista Castelnau, 1840 | | | Proagoderus loricatus Klug, 1855 | | | Sarophorus costatus Fahraeus, 1857 | | | Scarabaeus ambiguus (Boheman, 1857) | ⇒ Gnorimus variabilis Linnaeus, 1758 | | Scarabaeus anderseni Waterhouse, 1890 | | | Scarabaeus basuto zur Strassen, 1962 | | | Scarabaeus bohemani Harold, 1868 | | | Scarabaeus costatus Wiedemann, 1823 | | | Scarabaeus flavicornis Boheman, 1860 | | | Scarabaeus fritschi Harold, 1868 | | | Scarabaeus funebris Boheman, 1857 | | | Scarabaeus galenus Westwood, 1847 | | | Scarabaeus goryi Castelnau, 1840 | | | Scarabaeus heqvisti zur Strassen, 1962 | | | Scarabaeus inguisitus | | | Scarabaeus inoportunus Ferreira, 1953 | | | Scarabaeus inquisitus Péringuey, 1908 | | | Scarabaeus interstitialis Boheman, 1857 | | | Scarabaeus intricatus Fabricius, 1801 | | | Scarabaeus karrooensis zur Strassen, 1961 | | | Scarabaeus pabulator Péringuey, 1908 | | | Scarabaeus viator Péringuey, 1901 | | | Scarabaeus vicinus (Janssens, 1940) | ⇒ Lucanus cervus Linnaeus, 1758 | | Sceliages adamastor Lepeletier &..., 1828 | | | Sisyphus barbarossa (Wiedemann, 1823) | ⇒ Neosisyphus spinipes Thunberg, 1818 | | Sisyphus caffer Boheman, 1857 | | | Sisyphus costatus Thunberg, 1818 | | | Sisyphus crispatus Gory, 1833 | | | Sisyphus goryi (Harold, 1859) | ⇒ Sisyphus caffer goryi Harold, 1859 | | Sisyphus macrorubrus (Paschalidis, 1974) | ⇒ Neosisyphus macrorubrus Paschalidis, 1974 | | Sisyphus rubrus (Paschalidis, 1974) | ⇒ Neosisyphus rubrus Paschalidis, 1974 | | Sisyphus spinipes (Thunberg, 1818) | ⇒ Neosisyphus spinipes Thunberg, 1818 | | Stiptopodius patrizii Boucomont, 1923 | | | Tiniocellus eurypygus Branco, 2010 | | | Xinidium dentilabris Harold, 1869 | |
Subnodes of Scarabaeinae with occurrences in Free State
Links, sources and further information Scarabaeinae at artfakta.se
Scarabaeinae at dyntaxa.se
Scarabaeinae at itis.gov
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