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Checklist for Subfamily Scarabaeinae Latreille, 1802 (Dung Beetles)
and region Middle Miocene (Miocene)


Class
 
Insecta
Linnaeus, 1758
Insects, True insects
185
683
1400
Families
Genera
Species
 
Taxom
 

Subclass
 
Pterygota
Lang, 1888
Winged insects
185
683
1400
Families
Genera
Species
 
Taxom
 

Infraclass
 
Neoptera
van der Wulp, 1890
Modern Wing-folding insects
173
646
1347
Families
Genera
Species
 
Taxom
 

Superorder
 
Holometabola
Heider, 1889
Holometabolous Insects
108
459
992
Families
Genera
Species
 
Taxom
 

Order
 
Coleoptera
Linnaeus, 1758
Beetles, Coleopterans
41
238
455
Families
Genera
Species
 
Taxom
 

Suborder
 
Polyphaga
Emery, 1886
Polyphags
37
201
370
Families
Genera
Species
 
Taxom
 

Infraorder
 
Scarabaeiformia
Crowson, 1960
5
32
64
Families
Genera
Species
 
Taxom
 

Superfamily
 
Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles
5
32
64
Families
Genera
Species
 
Taxom
 
FamilyScarabaeidae
Latreille, 1802
Dung beetles, Flower beetles, Fruit chafers, Rain beetles
25
56
Genera
Species
 
Taxom
 
SubfamilyScarabaeinae
Latreille, 1802
Dung Beetles, True dung beetles
7
13
Genera
Species
 
Taxom
 


Genera for Middle Miocene
7 Genera with 13 Species
Copris Geoffroy, 1862Mexican dung beetle2SpeciesTaxom
Gymnopleurus Illiger, 18473SpeciesTaxom
Oniticellus Serville, 18621SpeciesTaxom
Onthophagus Latreille, 18675SpeciesTaxom
Phalops Erichson, 1847
0SpeciesTaxom
Scarabaeus Linnaeus, 18621SpeciesTaxom
Scelocopris Zhang, 19891SpeciesTaxom


Species list for Middle Miocene
13 Species
Copris druidum O. Heer, 1862
Copris subterranea O. Heer, 1862
Gymnopleurus deperditus O. Heer, 1862
Gymnopleurus rotundatus O. Heer, 1862
Gymnopleurus sisyphus O. Heer, 1847
Oniticellus amplicollis (O. Heer, 1862)⇒ Helictopleurus neoamplicollis Krell, 2000
Onthophagus bisontinus Heer, 1862
Onthophagus crassus (Heer, 1862)⇒ Parascatonomus tricornis Wiedemann, 1823
Onthophagus ovatulus Heer, 1847
Onthophagus prodromus Heer, 1862
Onthophagus urus (Harold, 1867)⇒ Onthophagus urusheeri Krell, 2000
Scarabaeus proserpinae O. Heer, 1862
Scelocopris enertheus J. F. Zhang, 1989

Subnodes of Scarabaeinae with occurrences in Middle Miocene
TribeCoprini
Kolbe, 1805
1
2
Genus
Species

Taxom

TribeGymnopleurini
Lacordaire, 1856
1
3
Genus
Species

Taxom

TribeOniticellini
d´Orbigny, 1916
1
1
Genus
Species

Taxom

TribeOnthophagini
Lacordaire, 1856
2
5
Genera
Species

Taxom

TribeScarabaeini
Latreille, 1802
1
1
Genus
Species

Taxom



Links, sources and further information
Scarabaeinae at artfakta.se
Scarabaeinae at dyntaxa.se
Scarabaeinae at itis.gov