Checklist for Subfamily Scarabaeinae Latreille, 1802 (Dung Beetles)
and region Near East (West Asia)
Class
| Insecta
Linnaeus, 1758
Insects, True insects | 687
11472
52844 | Families
Genera
Species |
Taxom
|
Subclass
| Pterygota
Lang, 1888
Winged insects | 682
11445
52791 | Families
Genera
Species |
Taxom
|
Infraclass
| Neoptera
van der Wulp, 1890
Modern Wing-folding insects | 651
11312
52466 | Families
Genera
Species |
Taxom
|
Superorder
| Holometabola
Heider, 1889
Holometabolous Insects | 408
8936
44750 | Families
Genera
Species |
Taxom
|
Order
| Coleoptera
Linnaeus, 1758
Beetles, Coleopterans | 110
2924
17636 | Families
Genera
Species |
Taxom
|
Suborder
| Polyphaga
Emery, 1886
Polyphags | 102
2584
13895 | Families
Genera
Species |
Taxom
|
Infraorder
| Scarabaeiformia
Crowson, 1960
| 11
320
1727 | Families
Genera
Species |
Taxom
|
Superfamily
| Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles | 11
320
1727 | Families
Genera
Species |
Taxom
| | Family | Scarabaeidae
Latreille, 1802
Dung beetles, Flower beetles, Fruit chafers, Rain beetles | 264
1429 | Genera
Species |
Taxom
| | Subfamily | Scarabaeinae
Latreille, 1802
Dung Beetles, True dung beetles | 31
260 | Genera
Species |
Taxom
|
Genera for Near East
31 Genera with 260 Species #ABCDEGHKLMOPS
| Allogymnopleurus Janssens, 1845 | | 1 | Species | Taxom | | Bubas Mulsant, 1811 | | 2 | Species | Taxom | | Caccobius Thomson, 1767 | | 4 | Species | Taxom | | Canthon Hoffmannsegg, 1946 | Tumblebug | 3 | Species | Taxom | | Catharsius Hope, 1888 | | 3 | Species | Taxom | | Cheironitis van Lansberge, 1789 | | 14 | Species | Taxom | | Chironitis Lansberge, 1792 | | 1 | Species | Taxom | | Cleptocaccobius Cambefort, 1857 | | 1 | Species | Taxom | | Copris Geoffroy, 1901 | Mexican dung beetle | 9 | Species | Taxom | | Delopleurus Erichson, 2014 | | 1 | Species | Taxom | | Digitonthophagus Balthasar, 2017 | | 5 | Species | Taxom | | Epirinus Reiche, 1775 | | 1 | Species | Taxom | | Euoniticellus Janssens, 1781 | | 3 | Species | Taxom | | Euonthophagus Balthasar, 1946 | | 24 | Species | Taxom | | Gymnopleurus Illiger, 1843 | | 25 | Species | Taxom | | Heliocopris Hope, 1775 | | 5 | Species | Taxom | | Kheper Janssens, 0 | | 2 | Species | Taxom | | Liatongus Reitter, 1840 | | 1 | Species | Taxom | | Metacatharsius Paulian, 1840 | | 1 | Species | Taxom | | Mnematidium Ritsema, 1845 | | 2 | Species | Taxom | | Mnematium MacLeay, 1832 | | 1 | Species | Taxom | | Oniticellus Serville, 1879 | | 2 | Species | Taxom | | Onitis Fabricius, 1857 | | 13 | Species | Taxom | | Onthophagus Latreille, 1939 | | 133 | Species | Taxom | | Paroniticellus Balthasar, 1809 | | 1 | Species | Taxom | | Phalops Erichson, 1883 | | 3 | Species | Taxom | | Proagoderus Lansberge, 1851 | | 2 | Species | Taxom | | Scarabaeus Linnaeus, 2006 | | 23 | Species | Taxom | | Sinodrepanus Simonis, 1875 | | 1 | Species | Taxom | | Sisyphus Latreille, 1758 | | 3 | Species | Taxom | | Synapsis Forster, 1821 | | 1 | Species | Taxom |
Species list for Near East
260 Species and 33 Subspecies | Allogymnopleurus anthracinus Klug, 1845 | | | Bubas bubaloides Janssens, 1938 | | | Bubas bubalus Olivier, 1811 | | | Caccobius histeroides Ménétriés, 1832 | | | Caccobius mundus Ménétriés, 1838 | | | Caccobius pulicarius Harold, 1875 | | | Caccobius schreberi Linnaeus, 1767 | | | Canthon (Boreocanthon) ebenus Say, 1823 | | | Canthon (Boreocanthon) probus Germar, 1824 | | | Canthon (Canthon) imitator Brown, 1946 | | | Catharsius alpheus Gillet, 1932 | | | Catharsius eteocles (Laporte, 1840) | ⇒ Catharsius alpheus Gillet, 1932 | | Catharsius sesostris Waterhouse, 1888 | | | Cheironitis arrowi Janssens, 1937 | | | Cheironitis candezei Lansberge, 1875 | | | Cheironitis furcifer Rossi, 1792 | | | Cheironitis haroldi Ballion, 1870 | | | Cheironitis hungaricus Herbst, 1789 | | | Cheironitis irroratus Rossi, 1790 | | | Cheironitis klapperichi (Balthasar, 1955) | ⇒ Cheironitis furcifer Rossi, 1792 | | Cheironitis moeris Pallas, 1781 | | | Cheironitis osiridis Reiche, 1856 | | | Cheironitis pamphilus Ménétriés, 1849 | | | Cheironitis petraensis Coppo, 2021 | | | Cheironitis socotranus Gahan, 1900 | | | Cheironitis sterculius Ballion, 1870 | | | Cheironitis ungaricus Herbst, 1789 | | | Chironitis furcifer Rossi, 1792 | | | Cleptocaccobius viridicollis Fahraeus, 1857 | | | Copris armeniacus Falderman, 1835 | | | Copris felschei (Reitter, 1893) | ⇒ Copris armeniacus Falderman, 1835 | | Copris hispanus Linnaeus, 1764 | | | Copris hispanus cavolinii Petagna, 1792 | | | Copris hispanus hispanus Linné, 1764 | | | Copris hispanus persianus Miksic, 1958 | | | Copris incertus Say, 1835 | Black dung beetle | | Copris lunaris Linnaeus, 1758 | Horned Dung Beetle | | Copris umbilicatus Abeille de P..., 1901 | | | Delopleurus mencli Kral, 2014 | | | Digitonthophagus aksumensis Génier, 2017 | | | Digitonthophagus bonasus Fabricius, 1775 | | | Digitonthophagus eucatta Génier, 2017 | | | Digitonthophagus gazella Fabricius, 1787 | Brown dung beetle | | Digitonthophagus uks Génier, 2017 | | | Epirinus flagellatus Fabricius, 1775 | | | Euoniticellus fulvus Goeze, 1777 | | | Euoniticellus pallens Olivier, 1789 | | | Euoniticellus pallipes Fabricius, 1781 | Southern sandy dung beetle | | Euonthophagus aeneobrunneus Kabakov, 1977 | | | Euonthophagus amyntas Olivier, 1789 | | | Euonthophagus amyntas alces Fabricius, 1792 | | | Euonthophagus amyntas amyntas Olivier, 1789 | | | Euonthophagus amyntas auchenia Redtenbacher, 1850 | | | Euonthophagus amyntas subviolaceus Ménétries, 1832 | | | Euonthophagus atramentarius Ménétriés, 1832 | | | Euonthophagus bedeli Reitter, 1891 | | | Euonthophagus carbonarius Klug, 1855 | | | Euonthophagus conterminus Petrovitz, 1971 | | | Euonthophagus dorbignyi Olsoufieff, 1900 | | | Euonthophagus dorbignyi neutto Olsoufieff, 1900 | | | Euonthophagus gibbosus Scriba, 1790 | | | Euonthophagus koshantschikoffi Reitter, 1891 | | | Euonthophagus loeffleri Petrovitz, 1965 | | | Euonthophagus maindroni d´Orbigny, 1897 | | | Euonthophagus mostafatsairii Palestrini, ..., 1979 | | | Euonthophagus pentaceros Olsoufieff, 1900 | | | Euonthophagus pertinax Balthasar, 1959 | | | Euonthophagus sulcicollis Reitter, 1892 | | | Euonthophagus tissoni Reitter, 1906 | | | Euonthophagus verberatus Ziani, 2006 | | | Euonthophagus yama Ziani, 2006 | | | Euonthophagus yemenicus Balthasar, 1946 | | | Gymnopleurus aciculatus Gebler, 1841 | | | Gymnopleurus aeruginosus Harold, 1867 | | | Gymnopleurus atratus Klug, 1845 | | | Gymnopleurus bicallosus Felsche, 1909 | | | Gymnopleurus elegans Klug, 1845 | | | Gymnopleurus flagellatus Fabricius, 1787 | | | Gymnopleurus flagellatus asperatus Motschulsky, 1849 | | | Gymnopleurus flagellatus calignosus Balthasar, 1934 | | | Gymnopleurus flagellatus flagellatus Fabricius, 1787 | | | Gymnopleurus flagellatus serratus Fischer von ..., 1821 | | | Gymnopleurus geoffroyi Fuessly, 1775 | | | Gymnopleurus geoffroyi cyanescens Motschulsky, 1849 | | | Gymnopleurus geoffroyi geoffroyi | | | Gymnopleurus lacunosus Klug, 1845 | | | Gymnopleurus mimus (Balthasar, 1934) | ⇒ Gymnopleurus geoffroyi Fuessly, 1775 | | Gymnopleurus mopsus Pallas, 1781 | | | Gymnopleurus mopsus mopsus | | | Gymnopleurus naviauxi Montreuil, 2009 | | | Gymnopleurus persianus (Reitter, 1909) | ⇒ Gymnopleurus mopsus Pallas, 1781 | | Gymnopleurus puncticollis (Gillet, 1909) | ⇒ Gymnopleurus virens Erichson, 1843 | | Gymnopleurus qurosh Montreuil, 2011 | | | Gymnopleurus serratus (Fischer von ..., 1821) | ⇒ Gymnopleurus flagellatus Fabricius, 1787 | | Gymnopleurus sturmi (MacLeay, 1821) | ⇒ Gymnopleurus sturmii MacLeay, 1821 | | Gymnopleurus sturmii MacLeay, 1821 | | | Gymnopleurus virens Erichson, 1843 | | | Heliocopris andersoni Bates, 1868 | | | Heliocopris gigas Linnaeus, 1758 | | | Heliocopris gigas gigas | | | Heliocopris isidis (Boheman, 1857) | ⇒ Heliocopris andersoni Bates, 1868 | | Heliocopris midas Fabricius, 1775 | | | Kheper aegyptiorum Latreille, 1823 | | | Kheper aegyptiorum aegyptiorum | | | Liatongus militaris Castelnau, 1840 | Yellow Shouldered Dung Beetle | | Metacatharsius inermis Castelnau, 1840 | | | Mnematidium multidentatum (Klug, 1845) | ⇒ Scarabaeus multidentatum Klug, 1845 | | Mnematidium multidentatus Klug, 1845 | | | Mnematium silenus Gray, 1832 | | | Oniticellus falsus (Sharp, 1875) | ⇒ Sinodrepanus falsus Sharp, 1875 | | Oniticellus tessellatus Harold, 1879 | | | Onitis alexis (Klug, 1835) | ⇒ Cheironitis ungaricus ungaricus Herbst, 1789 | | Onitis alexis alexis | | | Onitis alexis septentrionalis Balthasar, 1942 | | | Onitis damoetas Steven, 1806 | | | Onitis ezechias Reiche, 1856 | | | Onitis humerosus Pallas, 1771 | | | Onitis ion Olivier, 1789 | | | Onitis numida Castelnau, 1840 | | | Onitis pers Gusakov & Kl..., 2009 | | | Onitis sphinx Fabricius, 1775 | | | Onitis tingaudi Keith & Moretto, 2007 | | | Onitis tumidus Gillet, 1918 | | | Onitis viridulus (Boheman, 1857) | ⇒ Onitis pecuarius Lansberge, 1875 | | Onthophagus (Amphionthophagus) falzonii Goidanich, 1926 | | | Onthophagus (Euonthophagus) amyntas (Olivier, 1789) | ⇒ Euonthophagus amyntas Olivier, 1789 | | Onthophagus (Euonthophagus) atramentarius Ménétriés, 1832 | | | Onthophagus (Furconthophagus) furcatus (Fabricius, 1781) | ⇒ Onthophagus furcaticeps Masters, 1886 | | Onthophagus (Onthophagus) amirus Kabakov, 1982 | | | Onthophagus (Onthophagus) illyricus Scopoli, 1763 | | | Onthophagus (Onthophagus) margaritifer d´Orbigny, 1898 | | | Onthophagus (Onthophagus) oklahomensis Brown, 1927 | Oklahoma dung beetle | | Onthophagus (Onthophagus) pennsylvanicus Harold, 1871 | | | Onthophagus (Onthophagus) taurus Schreber, 1759 | Bull Headed Dung Beetle | | Onthophagus (Palaeonthophagus) angorensis Petrovitz, 1963 | | | Onthophagus (Palaeonthophagus) bytinskii Balthasar, 1960 | | | Onthophagus (Palaeonthophagus) coenobita Herbst, 1783 | | | Onthophagus (Palaeonthophagus) conspersus Reitter, 1893 | | | Onthophagus (Palaeonthophagus) cruciatus (Ménétriés, 1832) | ⇒ Onthophagus lemur Fabricius, 1781 | | Onthophagus (Palaeonthophagus) dacatrai Pittino, 2004 | | | Onthophagus (Palaeonthophagus) dellacasai Pittino & Ma..., 1981 | | | Onthophagus (Palaeonthophagus) excisus Reiche & Saulcy, 1856 | | | Onthophagus (Palaeonthophagus) excubitor Ziani & Gudenzi, 2006 | | | Onthophagus (Palaeonthophagus) fissicornis Steven, 1809 | | | Onthophagus (Palaeonthophagus) fortigibber Reitter, 1909 | | | Onthophagus (Palaeonthophagus) fracticornis (Preyssler, 1790) | ⇒ Onthophagus similis Scriba, 1790 | | Onthophagus (Palaeonthophagus) furciceps Marseul, 1869 | | | Onthophagus (Palaeonthophagus) gibbulus Pallas, 1781 | | | Onthophagus gibbulus gibbulus | | | Onthophagus gibbulus rostrifer Reitter, 1892 | | | Onthophagus (Palaeonthophagus) grossepunctatus Reitter, 1905 | | | Onthophagus (Palaeonthophagus) haroldi Ballion, 1870 | | | Onthophagus (Palaeonthophagus) isikdagensis Pittino, 2004 | | | Onthophagus (Palaeonthophagus) kindermanni Harold, 1877 | | | Onthophagus (Palaeonthophagus) lemur Fabricius, 1781 | | | Onthophagus (Palaeonthophagus) leucostigma Steven, 1811 | | | Onthophagus (Palaeonthophagus) lucidus Illiger, 1800 | | | Onthophagus (Palaeonthophagus) marginalis Gebler, 1817 | | | Onthophagus marginalis marginalis | | | Onthophagus (Palaeonthophagus) medius (Kugelann, 1792) | ⇒ Onthophagus vacca Linnaeus, 1767 | | Onthophagus (Palaeonthophagus) muelleri (Lansberge, 1883) | ⇒ Onthophagus statzi Krell, 1990 | | Onthophagus (Palaeonthophagus) novaki Boucomont, 1927 | | | Onthophagus (Palaeonthophagus) nuchicornis Linnaeus, 1758 | Mottled dung beetle | | Onthophagus (Palaeonthophagus) opacicollis Reitter, 1893 | | | Onthophagus (Palaeonthophagus) ovatus Linnaeus, 1767 | | | Onthophagus (Palaeonthophagus) parmatus Reitter, 1892 | | | Onthophagus (Palaeonthophagus) petrovitzi (Frey, 1975) | ⇒ Onthophagus petrovitzianus Pittino, 2006 | | Onthophagus (Palaeonthophagus) petrovitzianus Pittino, 2006 | | | Onthophagus (Palaeonthophagus) ponticus Harold, 1883 | | | Onthophagus (Palaeonthophagus) psychopompus Ziani & Ghar..., 2010 | | | Onthophagus (Palaeonthophagus) pygargus Motschulsky, 1845 | | | Onthophagus (Palaeonthophagus) rakovici Pittino, 2004 | | | Onthophagus (Palaeonthophagus) ruficapillus Brullé, 1832 | | | Onthophagus ruficapillus guilanensis (Pittino, 1982) | ⇒ Onthophagus ruficapillus Brullé, 1832 | | Onthophagus ruficapillus ruficapillus Brullé, 1832 | | | Onthophagus (Palaeonthophagus) samai Ziani, 2011 | | | Onthophagus (Palaeonthophagus) semicornis Panzer, 1798 | | | Onthophagus (Palaeonthophagus) sericatus Reitter, 1893 | | | Onthophagus (Palaeonthophagus) similis (Scriba, 1790) | ⇒ Onthophagus fracticornis Preyssler, 1790 | | Onthophagus (Palaeonthophagus) speculifer Solsky, 1876 | | | Onthophagus (Palaeonthophagus) strabo Reitter, 1892 | | | Onthophagus (Palaeonthophagus) suturellus Brullé, 1832 | | | Onthophagus (Palaeonthophagus) truchmenus Kolenati, 1846 | | | Onthophagus truchmenus iranicus Kabakov, 2006 | | | Onthophagus truchmenus truchmenus | | | Onthophagus truchmenus xerxes Petrovitz, 1965 | | | Onthophagus (Palaeonthophagus) vacca Linnaeus, 1767 | | | Onthophagus (Palaeonthophagus) verticicornis (Laicharting, 1781) | ⇒ Onthophagus nuchicornis Linnaeus, 1758 | | Onthophagus (Palaeonthophagus) viridis Ménétriés, 1832 | | | Onthophagus (Palaeonthophagus) vlasovi Medvedev, 1958 | | | Onthophagus andalusicus persianus Olsoufieff, 1900 | | | Onthophagus (Paleonthophagus) nuchicornis Linnaeus, 1758 | | | Onthophagus (Paleonthophagus) similis Scriba, 1790 | | | Onthophagus (Paleonthophagus) vacca Linnaeus, 1767 | | | Onthophagus (Phalops) tricuspis (Semenov, 1900) | ⇒ Onthophagus fortigibber Reitter, 1909 | | Onthophagus (Strandius) obliquus Olivier, 1789 | | | Onthophagus aeneopiceus d´Orbigny, 1902 | | | Onthophagus aerarius Reitter, 1892 | | | Onthophagus aethiopicus d´Orbigny, 1902 | | | Onthophagus aleppensis Redtenbacher, 1843 | | | Onthophagus aleppensis aleppensis | | | Onthophagus aleppensis hyrcanicus Kabakov, 1982 | | | Onthophagus amphicoma Boucomont, 1914 | | | Onthophagus anatolicus Petrovitz, 1962 | | | Onthophagus basipustulatus Heyden, 1879 | | | Onthophagus bicolor (Raffray, 1877) | ⇒ Onthophagus flavolimbatus Klug, 1855 | | Onthophagus bicristatus d´Orbigny, 1905 | | | Onthophagus blanchardi Harold, 1869 | | | Onthophagus bonsae Zunino, 1976 | | | Onthophagus brittoni Paulian, 1948 | | | Onthophagus carpanetoi Pittino, 1982 | | | Onthophagus carpophilus Pereira & Ha..., 1961 | | | Onthophagus circulator Reitter, 1891 | | | Onthophagus dorsosignatus d´Orbigny, 1898 | | | Onthophagus falsus Gillet, 1925 | | | Onthophagus filicornis Harold, 1873 | | | Onthophagus fissinasus Fairmaire, 1895 | | | Onthophagus flagrans Reitter, 1893 | | | Onthophagus formaneki Reitter, 1897 | | | Onthophagus frontalis Raffray, 1877 | | | Onthophagus furcaticeps Masters, 1886 | | | Onthophagus hermonensis Baraud, 1982 | | | Onthophagus heydeni Harold, 1875 | | | Onthophagus infuscatus Klug, 1845 | | | Onthophagus jugicola d´Orbigny, 1902 | | | Onthophagus latipennis d´Orbigny, 1897 | | | Onthophagus lemuroides d´Orbigny, 1898 | | | Onthophagus liothorax Koshantschikov, 1894 | | | Onthophagus malthinus Gillet, 1930 | | | Onthophagus melanocephalus Klug, 1845 | | | Onthophagus millingeni d´Orbigny, 1900 | | | Onthophagus moreleti Baraud, 1980 | | | Onthophagus nebulosus Reiche, 1864 | | | Onthophagus nitidulus Klug, 1845 | | | Onthophagus oberthueri d´Orbigny, 1898 | | | Onthophagus oberthuri d´Orbigny, 1898 | | | Onthophagus ochreatus d´Orbigny, 1897 | | | Onthophagus orcas (Ménétriés, 1838) | ⇒ Onthophagus atramentarius Ménétriés, 1832 | | Onthophagus osellai Pittino, 1982 | | | Onthophagus pictus (Reitter, 1893) | ⇒ Oniticellus tessellatus Harold, 1879 | | Onthophagus roessneri Ziani, 2016 | | | Onthophagus rugulipennis Fairmaire, 1887 | | | Onthophagus sacharovskii Olsoufieff, 1918 | | | Onthophagus sellatus Klug, 1845 | | | Onthophagus shaykh Montanaro & ..., 2022 | | | Onthophagus simius Reitter, 1893 | | | Onthophagus sparsulus Reitter, 1893 | | | Onthophagus statzi ♰ Krell, 1990 | | | Onthophagus sticticus Harold, 1867 | | | Onthophagus suermelii Petrovitz, 1963 | | | Onthophagus tessulatus Harold, 1871 | | | Onthophagus transcaspicus Koenig, 1888 | Gazella scarab | | Onthophagus trapezicornis d´Orbigny, 1902 | | | Onthophagus variegatus (Fabricius, 1798) | ⇒ Onthophagus filicornis Harold, 1873 | | Onthophagus viriditinctus Reitter, 1893 | | | Onthophagus zagrosicus Kabakov, 2006 | | | Onthophagus zuvandi Qaryagdy, 1939 | | | Paroniticellus festivus Steven, 1809 | | | Phalops chalybeus Klug, 1845 | | | Phalops iphis Olivier, 1789 | | | Phalops olivaceus Lansberge, 1883 | | | Proagoderus quadriarmatus Fairmaire, 1892 | | | Proagoderus worsissa Roth, 1851 | | | Scarabaeus (Ateuchetus) armeniacus Ménétriés, 1832 | | | Scarabaeus (Ateuchetus) puncticollis (Latreille, 1819) | ⇒ Geotrupes spiniger Marsham, 1802 | | Scarabaeus (Ateuchetus) semipunctatus Fabricius, 1792 | | | Scarabaeus (Ateuchetus) variolosus (Fabricius, 1787) | ⇒ Ateuchetus semipunctatus Fabricius, 1792 | | Scarabaeus (Scarabaeus) babori Balthasar, 1934 | | | Scarabaeus (Scarabaeus) pius Illiger, 1803 | | | Scarabaeus (Scarabaeus) sacer (Linnaeus, 1758) | ⇒ Scarabaeus pius Illiger, 1803 | | Scarabaeus (Scarabaeus) typhon (Fischer von ..., 1823) | ⇒ Megasoma gyas Herbst, 1785 | | Scarabaeus (Scarabaeus) winkleri Stolfa, 1938 | | | Scarabaeus acuticollis Motschulsky, 1849 | | | Scarabaeus aegyptiacus Stolfa, 1938 | | | Scarabaeus aegyptiorum Latreille, 1827 | | | Scarabaeus cristatus villiersi Baraud, 1973 | | | Scarabaeus gangeticus Castelnau, 1840 | | | Scarabaeus irakensis Stolfa, 1938 | | | Scarabaeus isidis Castelnau, 1840 | | | Scarabaeus krutai Maly & Montr..., 2011 | | | Scarabaeus multidentatum Klug, 1845 | | | Scarabaeus pseudogangeticus Moretto, 2023 | | | Scarabaeus sennaariensis Castelnau, 1840 | | | Scarabaeus transcaspicus Stolfa, 1938 | | | Scarabaeus wilsoni Waterhouse, 1890 | | | Scarabaeus zagrosensis Montreuil, 2006 | | | Sinodrepanus falsus Sharp, 1875 | | | Sisyphus schaefferi Linnaeus, 1758 | | | Sisyphus schaefferi boschniaki Fischer von ..., 1823 | | | Sisyphus schaefferi schaefferi Linnaeus, 1758 | | | Synapsis tmolus Fischer von ..., 1821 | |
Subnodes of Scarabaeinae with occurrences in Near East
Links, sources and further information Scarabaeinae at artfakta.se
Scarabaeinae at dyntaxa.se
Scarabaeinae at itis.gov
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