Checklist for Subfamily Scarabaeinae Latreille, 1802 (Dung Beetles)
and region Borneo (Indonesia)
Class
| Insecta
Linnaeus, 1758
Insects, True insects | 420
4676
14052 | Families
Genera
Species |
Taxom
|
Subclass
| Pterygota
Lang, 1888
Winged insects | 420
4676
14052 | Families
Genera
Species |
Taxom
|
Infraclass
| Neoptera
van der Wulp, 1890
Modern Wing-folding insects | 400
4569
13774 | Families
Genera
Species |
Taxom
|
Superorder
| Holometabola
Heider, 1889
Holometabolous Insects | 256
3371
10525 | Families
Genera
Species |
Taxom
|
Order
| Coleoptera
Linnaeus, 1758
Beetles, Coleopterans | 77
1597
5463 | Families
Genera
Species |
Taxom
|
Suborder
| Polyphaga
Emery, 1886
Polyphags | 73
1425
4831 | Families
Genera
Species |
Taxom
|
Infraorder
| Scarabaeiformia
Crowson, 1960
| 9
218
1268 | Families
Genera
Species |
Taxom
|
Superfamily
| Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles | 9
218
1268 | Families
Genera
Species |
Taxom
| | Family | Scarabaeidae
Latreille, 1802
Dung beetles, Flower beetles, Fruit chafers, Rain beetles | 157
879 | Genera
Species |
Taxom
| | Subfamily | Scarabaeinae
Latreille, 1802
Dung Beetles, True dung beetles | 20
177 | Genera
Species |
Taxom
|
Genera for Borneo
20 Genera with 177 Species | Anoctus Sharp, 1885 | | 1 | Species | Taxom | | Caccobius Thomson, 1877 | | 3 | Species | Taxom | Canthon Hoffmannsegg, 1817 | Tumblebug | 0 | Species | Taxom | | Catharsius Hope, 1996 | | 3 | Species | Taxom | | Copris Geoffroy, 1881 | Mexican dung beetle | 13 | Species | Taxom | | Cyobius Sharp, 1875 | | 4 | Species | Taxom | | Haroldius Boucomont, 1994 | | 6 | Species | Taxom | | Heliocopris Hope, 1859 | | 1 | Species | Taxom | | Liatongus Reitter, 1800 | | 1 | Species | Taxom | | Ochicanthon Vaz-de-Mello, 2006 | | 20 | Species | Taxom | | Oniticellus Serville, 1879 | | 2 | Species | Taxom | | Onitis Fabricius, 1931 | | 1 | Species | Taxom | | Onthophagus Latreille, 2006 | | 110 | Species | Taxom | | Panelus Lewis, 1992 | | 7 | Species | Taxom | | Paragymnopleurus Shipp, 1875 | | 6 | Species | Taxom | | Paraphytus Harold, 1880 | | 3 | Species | Taxom | | Proagoderus Lansberge, 2002 | | 2 | Species | Taxom | | Sisyphus Latreille, 0 | | 2 | Species | Taxom | | Synapsis Forster, 1874 | | 1 | Species | Taxom | | Yvescambefortius Ochi & Kon, 1926 | | 1 | Species | Taxom |
Species list for Borneo
177 Species and 12 Subspecies | Anoctus scatonomoides Lansberge, 1885 | | | Caccobius (Caccophilus) unicornis Fabricius, 1798 | | | Caccobius bawangensis Ochi & Kon &..., 1997 | | | Caccobius binodulus Harold, 1877 | | | Catharsius dayacus Lansberge, 1886 | | | Catharsius molossus Linnaeus, 1758 | | | Catharsius renaudpauliani Ochi & Kon, 1996 | | | Copris agnus Sharp, 1875 | | | Copris doriae Harold, 1877 | | | Copris fujiokai Ochi & Kon, 1996 | | | Copris fujiokai poringensis Ochi & Kon, 2005 | | | Copris gibbulus Lansberge, 1886 | | | Copris gibbulus borneensis Ochi & Kon, 2005 | | | Copris gibbulus gibbulus | | | Copris minutus Drury, 1773 | Small Black Dung Beetle | | Copris numa Lansberge, 1886 | Braken Bat Cave meshweaver | | Copris poggii Ochi & Kon, 2005 | | | Copris ramosiceps Gillet, 1921 | | | Copris reflexus (Fabricius, 1787) | ⇒ Copris minutus Drury, 1773 | | Copris spinator Harold, 1881 | | | Cyobius brendelli Ochi, Kon & ..., 2010 | | | Cyobius cheyi Ochi, Kon & ..., 2006 | | | Cyobius mendeli Ochi, Kon & ..., 2010 | | | Cyobius wallacei Sharp, 1875 | | | Haroldius borneensis Paulian, 1993 | | | Haroldius discoidalis Paulian, 1993 | | | Haroldius oharai Ochi, Kon & ..., 2009 | | | Haroldius pauliani Scheuern, 1995 | | | Haroldius rugatulus Boucomont, 1914 | | | Haroldius sumatranus Paulian & Sc..., 1994 | | | Heliocopris tyrannus Thomson, 1859 | | | Liatongus femoratus Illiger, 1800 | | | Ochicanthon cambeforti Ochi & Kon &..., 1997 | | | Ochicanthon crockermontis Krikken & Hu..., 2007 | | | Ochicanthon crypticus Krikken & Hu..., 2007 | | | Ochicanthon danum Krikken & Hu..., 2007 | | | Ochicanthon dulitmontis Krikken & Hu..., 2007 | | | Ochicanthon dytiscoides Boucomont, 1914 | | | Ochicanthon edmondsi Krikken & Hu..., 2007 | | | Ochicanthon gangkui Ochi & Kon &..., 1997 | | | Ochicanthon hanskii Krikken & Hu..., 2007 | | | Ochicanthon hikidai Ochi & Kon &..., 1997 | | | Ochicanthon kikutai Ochi, Ueda &..., 2006 | | | Ochicanthon kimanis Krikken & Hu..., 2007 | | | Ochicanthon maryatiae Ochi, Ueda &..., 2006 | | | Ochicanthon mulu Krikken & Hu..., 2007 | | | Ochicanthon neglectus Krikken & Hu..., 2007 | | | Ochicanthon parantisae Ochi & Kon &..., 1997 | | | Ochicanthon rombauti Krikken & Hu..., 2007 | | | Ochicanthon tambunan Krikken & Hu..., 2007 | | | Ochicanthon uedai Ochi, Kon, H..., 2008 | | | Ochicanthon woroae Ochi, Ueda &..., 2006 | | | Oniticellus cinctus Fabricius, 1775 | Bordered dung beetle | | Oniticellus tessellatus Harold, 1879 | | | Onitis subopacus Arrow, 1931 | | | Onthophagus (Furconthophagus) lilliputanus Lansberge, 1883 | | | Onthophagus (Furconthophagus) papulatus Boucomont, 1914 | | | Onthophagus (Gibbonthophagus) fujiii Ochi & Kon, 1995 | | | Onthophagus (Gibbonthophagus) kimioi Ochi, Kon & ..., 2011 | | | Onthophagus (Gibbonthophagus) limbatus Herbst, 1789 | | | Onthophagus (Gibbonthophagus) parviobscurior Ochi, Kon & ..., 2009 | | | Onthophagus (Indachorius) danumensis (Ochi, Kon & ..., 2009) | ⇒ Onthophagus danumicus Huijbregts &..., 2012 | | Onthophagus (Indachorius) masaoi Ochi, 1992 | | | Onthophagus (Indachorius) mendeli Ochi, Kon & ..., 2009 | | | Onthophagus (Indachorius) paramasaoi Ochi, Kon & ..., 2009 | | | Onthophagus (Indachorius) pseudoworoae Ochi, Kon & ..., 2009 | | | Onthophagus (Indachorius) semidanumensis Ochi, Kon & ..., 2009 | | | Onthophagus (Macronthophagus) diabolicus Harold, 1877 | | | Onthophagus uenoi borneomontanus Ochi, 2003 | | | Onthophagus (Onthophagus) angustatus Boucomont, 1914 | | | Onthophagus (Onthophagus) ishiii Ochi & Kon, 1995 | | | Onthophagus (Onthophagus) javaecola Balthasar, 1959 | | | Onthophagus (Onthophagus) mulleri van Lansberge, 1883 | | | Onthophagus (Onthophagus) orientalis (Harold, 1868) | ⇒ Onthophagus fissinasus Fairmaire, 1895 | | Onthophagus (Onthophagus) trituber Wiedemann, 1823 | | | Onthophagus (Palaeonthophagus) muelleri (Lansberge, 1883) | ⇒ Onthophagus statzi Krell, 1990 | | Onthophagus (Palaeonthophagus) novaki Boucomont, 1927 | | | Onthophagus (Palaeonthophagus) olsoufieffi Boucomont, 1924 | | | Onthophagus (Palaeonthophagus) vacca Linnaeus, 1767 | | | Onthophagus (Paronthophagus) hidakai Ochi & Kon, 1995 | | | Onthophagus (Phanaeomorphus) johkii Ochi & Kon, 1994 | | | Onthophagus (Serrophorus) laevis (Harold, 1880) | ⇒ Onthophagus laevissimus d´Orbigny, 1905 | | Onthophagus laevis laevis | | | Onthophagus (Serrophorus) sagittarius Fabricius, 1775 | Sagittarius dung beetle | | Onthophagus aereopictus Boucomont, 1914 | | | Onthophagus aphodioides Lansberge, 1883 | | | Onthophagus armatus Blanchard, 1853 | | | Onthophagus avocetta Arrow, 1933 | | | Onthophagus azusae Ochi & Kon, 2006 | | | Onthophagus babirussa Eschscholtz, 1822 | | | Onthophagus bangueyensis Boucomont, 1914 | | | Onthophagus batillifer Harold, 1875 | | | Onthophagus bellus d´Orbigny, 1905 | | | Onthophagus bonorae Zunino, 1976 | | | Onthophagus borneensis Harold, 1877 | | | Onthophagus borneotagal Ochi, Kon & ..., 2016 | | | Onthophagus carpophilus Pereira & Ha..., 1961 | | | Onthophagus cervicapra Boucomont, 1914 | | | Onthophagus chandrai Ochi, 2007 | | | Onthophagus cheyi Ochi & Kon, 2006 | | | Onthophagus clivimerus Huijbregts &..., 2011 | | | Onthophagus collinsi Krikken & Hu..., 1987 | | | Onthophagus crassicollis Boucomont, 1913 | | | Onthophagus cupreopastillatus Ochi & Kon, 2006 | | | Onthophagus danumicus Huijbregts &..., 2012 | | | Onthophagus dayacus Boucomont, 1914 | | | Onthophagus deliensis bawangicus Ochi, Kon & ..., 2009 | | | Onthophagus deliensis deliensis | | | Onthophagus difficilis (Le Guillou, 1844) | ⇒ Onthophagus vacca Linnaeus, 1767 | | Onthophagus fissinasus Fairmaire, 1895 | | | Onthophagus foedus Boucomont, 1914 | | | Onthophagus fukuyamai Ochi, Kon & ..., 2018 | | | Onthophagus gestroi (Harold, 1877) | ⇒ Onthophagus bellus d´Orbigny, 1905 | | Onthophagus hartiniae Ochi & Kon, 2009 | | | Onthophagus hikidai Ochi & Kon, 2006 | | | Onthophagus hiroyukii Ochi, 2007 | | | Onthophagus incisus Harold, 1877 | | | Onthophagus infucatus Harold, 1877 | | | Onthophagus insulindicus Boucomont, 1914 | | | Onthophagus javanensis Balthasar, 1963 | | | Onthophagus kapitensis Frey, 1971 | | | Onthophagus kashizakii Kon & Ochi, 2005 | | | Onthophagus kawaharai Ochi & Kon, 2007 | | | Onthophagus keikoae Ochi & Kon, 2014 | | | Onthophagus koechlei Masumoto, Oc..., 2011 | | | Onthophagus koni Ochi, 2007 | | | Onthophagus laevissimus d´Orbigny, 1905 | | | Onthophagus liwagensis Ochi & Kon, 2006 | | | Onthophagus magnioculus Ochi & Kon, 2006 | | | Onthophagus maryatiae Ochi & Kon, 2005 | | | Onthophagus matsuii Ochi & Kon, 2006 | | | Onthophagus megapacificus Ochi & Kon, 2006 | | | Onthophagus obscurior Boucomont, 1914 | | | Onthophagus ochromerus Harold, 1877 | | | Onthophagus otai Ochi & Kon, 2006 | | | Onthophagus pacificus Lansberge, 1885 | | | Onthophagus pastillatus Boucomont, 1919 | | | Onthophagus pavidus Harold, 1877 | | | Onthophagus phillippsorum Krikken & Hu..., 1987 | | | Onthophagus quasijohkii Ochi & Kon, 2005 | | | Onthophagus quasitagal Ochi & Kon, 2005 | | | Onthophagus robertopoggii Ochi & Kon, 2006 | | | Onthophagus rorarius Harold, 1877 | | | Onthophagus rouyeri Boucomont, 1914 | | | Onthophagus rutilans (Sharp, 1875) | ⇒ Onthophagus borneensis Harold, 1877 | | Onthophagus rutilans aborneensis Ochi, Kon & ..., 2009 | | | Onthophagus rutilans rutilans | | | Onthophagus sabahensis Ochi & Kon, 2006 | | | Onthophagus semipacificus Ochi & Kon, 2006 | | | Onthophagus sepilokensis Ochi & Kon, 2006 | | | Onthophagus sideki Krikken & Hu..., 1987 | | | Onthophagus simboroni Ochi & Kon, 2006 | | | Onthophagus statzi ♰ Krell, 1990 | | | Onthophagus subcornutus Boucomont, 1914 | | | Onthophagus sugihartoi Ochi, 2007 | | | Onthophagus sumatranus Lansberge, 1883 | | | Onthophagus taeniatus Boucomont, 1914 | | | Onthophagus tridentitibialis Ochi & Kon, 2008 | | | Onthophagus uedai Ochi & Kon, 2006 | | | Onthophagus vethi Krikken, 1977 | | | Onthophagus vulpes Harold, 1877 | | | Onthophagus waterstradti Boucomont, 1914 | | | Onthophagus woroae Ochi & Kon, 2006 | | | Onthophagus woroae woroae | | | Onthophagus yumotoi Ochi & Kon, 2006 | | | Panelus bakeri Boucomont, 1921 | | | Panelus borneensis Paulian, 1992 | | | Panelus brendelli Ochi, Kon & ..., 2009 | | | Panelus danumensis Ochi, Kon & ..., 2009 | | | Panelus kalimantanicus Ochi, Kon & ..., 2009 | | | Panelus mendeli Ochi, Kon & ..., 2009 | | | Panelus sabahi Paulian, 1992 | | | Paragymnopleurus maurus Sharp, 1875 | | | Paragymnopleurus sinuatus Olivier, 1789 | | | Paragymnopleurus sinuatus productus Sharp, 1875 | | | Paragymnopleurus sparsus Sharp, 1875 | | | Paragymnopleurus spinotus Boucomont, 1914 | | | Paragymnopleurus striatus Sharp, 1875 | | | Paraphytus doriae Harold, 1877 | | | Paraphytus foveatus Boucomont, 1923 | | | Paraphytus ritsemae Harold, 1880 | | | Proagoderus schwaneri Snellen van ..., 1864 | | | Proagoderus watanabei Ochi & Kon, 2002 | | | Sisyphus thoracicus Sharp, 1875 | | | Sisyphus thoracicus thoracicus | | | Synapsis ritsemae Lansberge, 1874 | | | Yvescambefortius sarawacus Gillet, 1926 | |
Subnodes of Scarabaeinae with occurrences in Borneo
Links, sources and further information Scarabaeinae at artfakta.se
Scarabaeinae at dyntaxa.se
Scarabaeinae at itis.gov
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