Checklist for Subfamily Scarabaeinae Latreille, 1802 (Dung Beetles)
and region Costa Rica (Central America)
Class
| Insecta
Linnaeus, 1758
Insects, True insects | 611
11863
34633 | Families
Genera
Species |
Taxom
|
Subclass
| Pterygota
Lang, 1888
Winged insects | 608
11859
34631 | Families
Genera
Species |
Taxom
|
Infraclass
| Neoptera
van der Wulp, 1890
Modern Wing-folding insects | 579
11701
34157 | Families
Genera
Species |
Taxom
|
Superorder
| Holometabola
Heider, 1889
Holometabolous Insects | 379
9752
30015 | Families
Genera
Species |
Taxom
|
Order
| Coleoptera
Linnaeus, 1758
Beetles, Coleopterans | 108
3263
9002 | Families
Genera
Species |
Taxom
|
Suborder
| Polyphaga
Emery, 1886
Polyphags | 102
3021
8293 | Families
Genera
Species |
Taxom
|
Infraorder
| Scarabaeiformia
Crowson, 1960
| 9
271
1352 | Families
Genera
Species |
Taxom
|
Superfamily
| Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles | 9
271
1352 | Families
Genera
Species |
Taxom
| | Family | Scarabaeidae
Latreille, 1802
Dung beetles, Flower beetles, Fruit chafers, Rain beetles | 219
1104 | Genera
Species |
Taxom
| | Subfamily | Scarabaeinae
Latreille, 1802
Dung Beetles, True dung beetles | 44
277 | Genera
Species |
Taxom
|
Genera for Costa Rica
44 Genera with 277 Species #ABCDEHIMOPSTU
| Agamopus Bates, 1887 | | 1 | Species | Taxom | | Anomiopus Westwood, 1939 | | 2 | Species | Taxom | | Ateuchus Weber, 1996 | | 15 | Species | Taxom | | Bdelyrus Harold, 1887 | | 1 | Species | Taxom | | Bradypodidium Vaz-de-Mello, 1928 | | 1 | Species | Taxom | | Caccobius Thomson, 1911 | | 2 | Species | Taxom | | Canthidium Erichson, 1981 | | 27 | Species | Taxom | | Canthon Hoffmannsegg, 1964 | Tumblebug | 30 | Species | Taxom | | Copris Geoffroy, 2003 | Mexican dung beetle | 13 | Species | Taxom | | Coprophanaeus Olsoufieff, 2008 | | 9 | Species | Taxom | | Cryptocanthon Balthasar, 2002 | | 4 | Species | Taxom | | Deltochilum Eschscholtz, 1887 | | 14 | Species | Taxom | Dendropaemon Perty, 1830 | | 0 | Species | Taxom | | Dichotomius Hope, 2022 | | 27 | Species | Taxom | | Digitonthophagus Balthasar, 1787 | | 1 | Species | Taxom | | Disphysema Harold, 1873 | | 1 | Species | Taxom | | Euoniticellus Janssens, 1848 | | 2 | Species | Taxom | | Eurysternus Dalman, 1887 | | 10 | Species | Taxom | | Eutrichillum | | 1 | Species | Taxom | | Helictopleurus Orbigny, 2000 | | 1 | Species | Taxom | | Heliocopris Hope, 1775 | | 1 | Species | Taxom | | Homocopris Burmeister, 1867 | | 1 | Species | Taxom | | Isocopris Pereira & Ma..., 1931 | | 1 | Species | Taxom | | Malagoniella Martínez, 1863 | | 2 | Species | Taxom | | Megathoposoma Balthasar, 1873 | | 1 | Species | Taxom | | Onitis Fabricius, 1798 | | 1 | Species | Taxom | | Onoreidium Vaz-de-Mello, 1981 | | 1 | Species | Taxom | | Ontherus Erichson, 1996 | | 5 | Species | Taxom | | Onthophagus Latreille, 2003 | | 66 | Species | Taxom | | Oxysternon Laporte, 1840 | | 3 | Species | Taxom | | Pedaridium Harold, 1948 | | 1 | Species | Taxom | | Phanaeus MacLeay, 1863 | Burrowing Dung Beetles | 12 | Species | Taxom | | Pinotus Erichson, 1931 | | 1 | Species | Taxom | | Pseudocanthon Bates, 1847 | | 1 | Species | Taxom | | Scatimus Erichson, 1939 | | 3 | Species | Taxom | | Scybalocanthon Martínez, 1887 | | 1 | Species | Taxom | | Sisyphus Latreille, 1863 | | 3 | Species | Taxom | | Sulcophanaeus Olsoufieff, 1856 | | 5 | Species | Taxom | | Sylvicanthon Halffter & M..., 1868 | | 1 | Species | Taxom | | Synapsis Forster, 1821 | | 1 | Species | Taxom | | Tetraechma Agassiz, 1813 | | 1 | Species | Taxom | | Trichillidium Vaz-de-Mello, 1948 | | 1 | Species | Taxom | | Trichillum Harold, 1928 | | 1 | Species | Taxom | | Uroxys Westwood, 1987 | | 16 | Species | Taxom |
Species list for Costa Rica
277 Species and 17 Subspecies | Agamopus lampros Bates, 1887 | | | Anomiopus birai Canhedo, 2006 | | | Anomiopus panamensis Paulian, 1939 | | | Ateuchus aeneomicans Harold, 1868 | | | Ateuchus alutacius Kohlmann & S..., 2012 | | | Ateuchus candezei Harold, 1868 | | | Ateuchus chrysopyge Bates, 1887 | | | Ateuchus earthorus Kohlmann & S..., 2009 | | | Ateuchus fetteri Kohlmann, 1996 | | | Ateuchus ginae Kohlmann, 1996 | | | Ateuchus hendrichsi Kohlmann, 1996 | | | Ateuchus howdeni Kohlmann, 1996 | | | Ateuchus illaesum Harold, 1868 | | | Ateuchus illaesus Harold, 1868 | | | Ateuchus pygidialis Harold, 1868 | | | Ateuchus rodriguezi Preudhomme d..., 1886 | | | Ateuchus solisi Kohlmann, 1996 | | | Ateuchus zoebischi Kohlmann, 1996 | | | Bdelyrus seminudus Bates, 1887 | | | Bradypodidium bradyporum Boucomont, 1928 | | | Caccobius histeroides Ménétriés, 1832 | | | Caccobius polygonus d´Orbigny, 1911 | | | Canthidium (Eucanthidium) laetum (Harold, 1867) | ⇒ Eucanthidium laetum Harold, 1867 | | Canthidium (Eucanthidium) pseudopuncticolle Solis & Kohl..., 2004 | | | Canthidium angusticeps Bates, 1887 | | | Canthidium annagabrielae Solis & Kohl..., 2004 | | | Canthidium ardens Bates, 1887 | | | Canthidium aurifex Bates, 1887 | | | Canthidium centrale Boucomont, 1928 | | | Canthidium darwini Kohlmann & S..., 2009 | | | Canthidium discopygidiale Howden & Young, 1981 | | | Canthidium emoryi Solis & Kohl..., 2004 | | | Canthidium guanacaste Howden & Gill, 1987 | | | Canthidium haroldi Preudhomme d..., 1886 | | | Canthidium hespenheidei Howden & Young, 1981 | | | Canthidium leucopterum Howden & Young, 1981 | | | Canthidium macroculare Howden & Gill, 1987 | | | Canthidium marianelae Solis & Kohl..., 2004 | | | Canthidium marielae Solis & Kohl..., 2004 | | | Canthidium pallidoalatum Howden & Young, 1981 | | | Canthidium perceptibile Howden & Young, 1981 | | | Canthidium planovultum Howden & Young, 1981 | | | Canthidium priscillae Solis & Kohl..., 2004 | | | Canthidium puncticolle Harold, 1867 | | | Canthidium rufinum Harold, 1867 | | | Canthidium tenebrosum Howden & Young, 1981 | | | Canthidium tuberifrons Howden & Young, 1981 | | | Canthidium variolosum Howden & Young, 1981 | | | Canthidium vespertinum Howden & Young, 1981 | | | Canthon (Boreocanthon) ebenus Say, 1823 | | | Canthon (Canthon) cyanellus LeConte, 1859 | | | Canthon (Canthon) humectus Say, 1831 | Longhorn blenny | | Canthon (Canthon) indigaceus (LeConte, 1866) | ⇒ Canthon humectus Say, 1831 | | Canthon indigaceus chevrolati Harold, 1868 | | | Canthon (Glaphyrocanthon) femoralis Chevrolat, 1834 | | | Canthon aberrans Harold, 1868 | | | Canthon aequinoctialis Harold, 1868 | | | Canthon angustatus Harold, 1867 | | | Canthon caelius Bates, 1887 | | | Canthon championi (Bates, 1887) | ⇒ Glaphyrocanthon championi Bates, 1887 | | Canthon deyrollei Harold, 1868 | | | Canthon euryscelis Bates, 1887 | | | Canthon hartmanni Howden & Gill, 1987 | | | Canthon humboldti Solis & Kohl..., 2002 | | | Canthon inusitatus Kohlmann & S..., 2006 | | | Canthon juvencus Harold, 1868 | | | Canthon leechi Martinez, Ha..., 1964 | | | Canthon lituratus Germar, 1813 | | | Canthon meridionalis Martinez, Ha..., 1964 | | | Canthon moniliatus (Bates, 1887) | ⇒ Scybalocanthon moniliatus Bates, 1887 | | Canthon morsei Howden, 1966 | | | Canthon mutabilis Lucas, 1857 | | | Canthon septemmaculatus Latreille, 1812 | | | Canthon septemmaculatus septemmaculatus | | | Canthon silvaticus Solis & Kohl..., 2002 | | | Canthon staigi Pereira, 1953 | | | Canthon subhyalinus Harold, 1867 | | | Canthon subhyalinus subhyalinus | | | Canthon vazquezae Martinez, Ha..., 1964 | | | Copris aspericollis Gillet, 1910 | | | Copris brevicornis Darling & Gé..., 2018 | | | Copris costaricensis Gahan, 1894 | | | Copris costaricensis costaricensis | | | Copris hispanus Linnaeus, 1764 | | | Copris incertus Say, 1835 | Black dung beetle | | Copris laeviceps Harold, 1869 | | | Copris lugubris Boheman, 1858 | | | Copris matthewsi Delgado & Ko..., 2001 | | | Copris megasoma Matthews & H..., 1959 | | | Copris remotus dicyrtus Matthews & H..., 1959 | | | Copris subpunctatus Gillet, 1910 | | | Copris tridentatus Solis & Kohl..., 2003 | | | Coprophanaeus boucardi Nevinson, 1891 | | | Coprophanaeus chiriquensis Olsufieff, 1924 | | | Coprophanaeus corythus Harold, 1863 | | | Coprophanaeus kohlmanni (Arnaud, 2002) | ⇒ Coprophanaeus morenoi Arnaud, 1982 | | Coprophanaeus morenoi Arnaud, 1982 | | | Coprophanaeus pecki Howden & Young, 1981 | | | Coprophanaeus solisi Arnaud, 1997 | | | Coprophanaeus telamon Erichson, 1847 | | | Coprophanaeus uhleri (Maly & Pokorny, 2008) | ⇒ Coprophanaeus chiriquensis Olsufieff, 1924 | | Cryptocanthon denticulum Cook, 2002 | | | Cryptocanthon lindemanae Howden & Gill, 1987 | | | Cryptocanthon osaensis Cook, 2002 | | | Cryptocanthon solisi Cook, 2002 | | | Deltochilum acanthus Kohlmann & S..., 2012 | | | Deltochilum acropyge Bates, 1887 | | | Deltochilum burmeisteri (Harold, 1867) | ⇒ Calhyboma mexicanum Burmeister, 1848 | | Deltochilum carinatum Westwood, 1837 | | | Deltochilum gibbosum Fabricius, 1775 | Humpback Dung Beetle | | Deltochilum lobipes Bates, 1887 | | | Deltochilum mexicanum Burmeister, 1848 | | | Deltochilum panamensis Howden, 1966 | | | Deltochilum parile Bates, 1887 | | | Deltochilum parile parile | | | Deltochilum pseudoparile Paulian, 1938 | | | Deltochilum scabriusculum Bates, 1887 | Rockpool blenny | | Deltochilum valgum Burmeister, 1873 | | | Deltochilum valgum acropyge Bates, 1887 | | | Dichotomius (Luederwaldtinia) agenor Harold, 1869 | | | Dichotomius (Luederwaldtinia) centralis Harold, 1869 | | | Dichotomius (Luederwaldtinia) danieli (Kohlmann & S..., 1997) | ⇒ Luederwaldtinia danieli Kohlmann & S..., 1997 | | Dichotomius (Luederwaldtinia) favi (Kohlmann & S..., 1997) | ⇒ Luederwaldtinia favi Kohlmann & S..., 1997 | | Dichotomius (Luederwaldtinia) femoratus (Howden & Young, 1981) | ⇒ Luederwaldtinia femoratus Howden & Young, 1981 | | Dichotomius (Luederwaldtinia) gamboaensis (Howden & Young, 1981) | ⇒ Luederwaldtinia gamboaensis Howden & Young, 1981 | | Dichotomius (Luederwaldtinia) inachus Erichson, 1847 | | | Dichotomius (Luederwaldtinia) mysticus Luederwaldt, 1935 | | | Dichotomius (Luederwaldtinia) rodrigoi (Kohlmann & S..., 1997) | ⇒ Luederwaldtinia rodrigoi Kohlmann & S..., 1997 | | Dichotomius (Luederwaldtinia) rugatus (Luederwaldt, 1935) | ⇒ Luederwaldtinia rugatus Luederwaldt, 1935 | | Dichotomius (Luederwaldtinia) simplicicornis Luederwaldt, 1935 | | | Dichotomius (Luederwaldtinia) yucatanus Bates, 1887 | | | Dichotomius (Selenocopris) ingens Luederwaldt, 1935 | | | Dichotomius amicitiae Kohlmann & S..., 1997 | | | Dichotomius annae Linnaeus, 1767 | | | Dichotomius bitiensis Gillet, 1911 | | | Dichotomius costaricensis Luederwaldt, 1935 | | | Dichotomius enioi Montoya-Moli..., 2021 | | | Dichotomius latistriatus (Luederwaldt, 1935) | ⇒ Luederwaldtinia latistriatus Luederwaldt, 1935 | | Dichotomius nevermanni Luederwaldt, 1935 | | | Dichotomius puncticollis Luederwaldt, 1935 | | | Dichotomius robustus Luederwaldt, 1935 | Apex mesh weaver | | Dichotomius rugosicollis Luederwaldt, 1935 | | | Dichotomius satanas Harold, 1867 | | | Dichotomius satanas satanas | | | Dichotomius virescens Luederwaldt, 1935 | | | Dichotomius woodruffi Solis & Kohl..., 2022 | | | Digitonthophagus gazella Fabricius, 1787 | Brown dung beetle | | Disphysema candezei Harold, 1873 | | | Euoniticellus fulvus Goeze, 1777 | | | Euoniticellus intermedius Reiche, 1848 | Intermediate sandy dung beetle | | Eurysternus angustulus Harold, 1869 | | | Eurysternus caribaeus Herbst, 1789 | | | Eurysternus foedus Guérin-Ménev..., 1844 | | | Eurysternus hamaticollis Balthasar, 1939 | | | Eurysternus magnus Castelnau, 1840 | | | Eurysternus mexicanus Harold, 1869 | | | Eurysternus olivaceus Génier, 2009 | | | Eurysternus plebejus Harold, 1880 | | | Eurysternus streblus Génier, 2009 | | | Eurysternus velutinus Bates, 1887 | | | Eutrichillum arcus Solis & Kohl..., 2003 | | | Helictopleurus neoamplicollis Krell, 2000 | | | Heliocopris hamadryas Fabricius, 1775 | | | Homocopris achamas Harold, 1867 | | | Isocopris foveolatus Luederwaldt, 1931 | | | Malagoniella astyanax Olivier, 1789 | | | Malagoniella astyanax yucateca Harold, 1863 | | | Megathoposoma candezei Harold, 1873 | | | Onitis belial Fabricius, 1798 | | | Onoreidium bottimeri Howden & Young, 1981 | | | Ontherus azteca Harold, 1869 | | | Ontherus brevicollis Kirsch, 1870 | | | Ontherus brevipennis Harold, 1867 | | | Ontherus pseudodidymus Génier, 1996 | | | Ontherus sextuberculatus Génier, 1996 | | | Onthophagus (Furconthophagus) furcatus (Fabricius, 1781) | ⇒ Onthophagus furcaticeps Masters, 1886 | | Onthophagus (Onthophagus) alluvius Howden and C..., 1963 | | | Onthophagus (Onthophagus) batesi Howden and C..., 1963 | | | Onthophagus (Onthophagus) cyanellus Bates, 1887 | | | Onthophagus (Onthophagus) hoepfneri Harold, 1869 | | | Onthophagus (Onthophagus) incensus Say, 1835 | Dung beetle | | Onthophagus (Onthophagus) landolti (Harold, 1880) | ⇒ Onthophagus schaefferi Howden and C..., 1963 | | Onthophagus landolti landolti Harold, 1880 | | | Onthophagus (Onthophagus) schaefferi Howden and C..., 1963 | | | Onthophagus (Palaeonthophagus) fracticornis (Preyssler, 1790) | ⇒ Onthophagus similis Scriba, 1790 | | Onthophagus (Palaeonthophagus) similis (Scriba, 1790) | ⇒ Onthophagus fracticornis Preyssler, 1790 | | Onthophagus (Palaeonthophagus) vacca Linnaeus, 1767 | | | Onthophagus (Paleonthophagus) nuchicornis Linnaeus, 1758 | | | Onthophagus (Paleonthophagus) similis Scriba, 1790 | | | Onthophagus brucei cyanopterus (d´Orbigny, 1905) | ⇒ Proagoderus cyanopterus d´Orbigny, 1905 | | Onthophagus acuminatus (Harold, 1880) | ⇒ Onthophagus embrikianus Paulian, 1936 | | Onthophagus andersoni Howden & Gill, 1987 | | | Onthophagus anthracinus (Harold, 1873) | ⇒ Caccobius histeroides Ménétriés, 1832 | | Onthophagus atriglabrus Howden & Gill, 1987 | | | Onthophagus atrosericeus Boucomont, 1932 | | | Onthophagus belorhinus Bates, 1887 | | | Onthophagus bidentatus (Drapiez, 1819) | ⇒ Caccobius schreberi Linnaeus, 1767 | | Onthophagus carpophilus Pereira & Ha..., 1961 | | | Onthophagus championi Bates, 1887 | | | Onthophagus chryses Bates, 1887 | | | Onthophagus coriaceoumbrosus Kohlmann & S..., 2001 | | | Onthophagus coscineus Bates, 1887 | | | Onthophagus criniger d´Orbigny, 1904 | | | Onthophagus crinitus (Harold, 1869) | ⇒ Onthophagus criniger d´Orbigny, 1904 | | Onthophagus crinitus crinitus | | | Onthophagus crinitus panamensis Bates, 1887 | | | Onthophagus cryptodicranius Kohlmann & S..., 2001 | | | Onthophagus curvicornis Latreille, 1811 | | | Onthophagus dicranius Bates, 1887 | | | Onthophagus dorsipilulus Howden & Gill, 1987 | | | Onthophagus embrikianus Paulian, 1936 | | | Onthophagus gazellinus Bates, 1887 | | | Onthophagus genuinus Kohlmann & S..., 2001 | | | Onthophagus grataehelenae Kohlmann & S..., 2001 | | | Onthophagus hopfneri | | | Onthophagus humboldti Kohlmann, So..., 2019 | | | Onthophagus inediapterus Kohlmann & S..., 2001 | | | Onthophagus kohlmanni Génier, 2017 | | | Onthophagus limonensis Kohlmann & S..., 2001 | | | Onthophagus marginicollis Harold, 1880 | | | Onthophagus maya Zunino, 1981 | | | Onthophagus micropterus Zunino & Hal..., 1981 | | | Onthophagus mirabilis Bates, 1887 | | | Onthophagus nasicornis Harold, 1869 | | | Onthophagus nasutus Guérin-Ménev..., 1855 | | | Onthophagus nemorivagus Kohlmann & S..., 2001 | | | Onthophagus notiodes Solis & Kohl..., 2003 | | | Onthophagus nubilus Kohlmann & S..., 2001 | | | Onthophagus nyctopus Bates, 1887 | | | Onthophagus orphnoides (Bates, 1887) | ⇒ Onthophagus mirabilis Bates, 1887 | | Onthophagus praecellens Bates, 1887 | | | Onthophagus propraecellens Howden & Gill, 1987 | | | Onthophagus quetzalis Howden & Gill, 1993 | | | Onthophagus rhinolophus Harold, 1869 | | | Onthophagus sharpi Harold, 1875 | | | Onthophagus singulariformis Kohlmann & S..., 2001 | | | Onthophagus solisi Howden & Gill, 1993 | | | Onthophagus stockwelli Howden & Young, 1956 | | | Onthophagus tapirus Sharp, 1877 | | | Onthophagus viridivinosus Kohlmann & S..., 2001 | | | Onthophagus xiphias (Solis & Kohl..., 2003) | ⇒ Onthophagus nuchicornis Linnaeus, 1758 | | Oxysternon festivum Linnaeus, 1758 | | | Oxysternon silenum Castelnau, 1840 | | | Oxysternon silenus Castelnau, 1840 | | | Pedaridium pilosum (Robinson, 1948) | ⇒ Trichillidium pilosum Robinson, 1948 | | Phanaeus (Notiophanaeus) endymion Harold, 1863 | | | Phanaeus (Notiophanaeus) pyrois Bates, 1887 | | | Phanaeus beltianus Bates, 1887 | | | Phanaeus changdiazi Kohlmann & S..., 2001 | | | Phanaeus demon Castelnau, 1840 | | | Phanaeus demon excelsus Bates, 1887 | | | Phanaeus excelsus (Bates, 1887) | ⇒ Phanaeus demon excelsus Bates, 1887 | | Phanaeus eximius Bates, 1887 | | | Phanaeus hermes Harold, 1868 | | | Phanaeus malyi Arnaud, 2002 | | | Phanaeus olsoufieffi (Balthasar, 1939) | ⇒ Phanaeus pyrois olsoufieffi Balthasar, 1939 | | Phanaeus wagneri Harold, 1863 | | | Pinotus foveolatus (Luederwaldt, 1931) | ⇒ Isocopris foveolatus Luederwaldt, 1931 | | Pseudocanthon perplexus LeConte, 1847 | | | Scatimus erinnyos Kohlmann & S..., 1996 | | | Scatimus ovatus Harold, 1862 | | | Scatimus quadridentatus (Balthasar, 1939) | ⇒ Scatimus ovatus Harold, 1862 | | Scybalocanthon moniliatus Bates, 1887 | | | Sisyphus crispatus Gory, 1833 | | | Sisyphus crispatus mexicanus Harold, 1863 | | | Sisyphus mexicanus (Harold, 1863) | ⇒ Sisyphus crispatus mexicanus Harold, 1863 | | Sulcophanaeus auricollis Harold, 1880 | | | Sulcophanaeus noctis Bates, 1887 | | | Sulcophanaeus noctis cupricollis Nevinson, 1891 | | | Sulcophanaeus noctis noctis | | | Sulcophanaeus velutinus Murray, 1856 | | | Sylvicanthon aequinoctialis Harold, 1868 | | | Synapsis tmolus Fischer von ..., 1821 | | | Tetraechma liturata Germar, 1813 | | | Trichillidium pilosum Robinson, 1948 | | | Trichillum bradyporum (Boucomont, 1928) | ⇒ Bradypodidium bradyporum Boucomont, 1928 | | Uroxys boneti Pereira & Ha..., 1961 | | | Uroxys bonplandi Kohlmann, So..., 2019 | | | Uroxys coarctatus Harold, 1867 | | | Uroxys deavilai Delgado & Ko..., 2007 | | | Uroxys depressifrons (Howden & Young, 1981) | ⇒ Uroxys pauliani Balthasar, 1940 | | Uroxys dybasi Howden & Young, 1981 | | | Uroxys gatunensis Howden & Young, 1981 | | | Uroxys gorgon Arrow, 1933 | | | Uroxys macrocularis (Howden & Young, 1981) | ⇒ Uroxys boneti Pereira & Ha..., 1961 | | Uroxys metagorgon Howden & Young, 1981 | | | Uroxys microcularis Howden & Young, 1981 | | | Uroxys micros Bates, 1887 | | | Uroxys nebulinus Howden & Gill, 1987 | | | Uroxys pauliani Balthasar, 1940 | | | Uroxys platypyga Howden & Young, 1981 | | | Uroxys transversifrons Howden & Gill, 1987 | |
Subnodes of Scarabaeinae with occurrences in Costa Rica
Links, sources and further information Scarabaeinae at artfakta.se
Scarabaeinae at dyntaxa.se
Scarabaeinae at itis.gov
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