|
|
| | | | Checklist for Subfamily Scarabaeinae Latreille, 1802 (Dung Beetles)
and region Southeast Asia (Far East)
Class
| Insecta
Linnaeus, 1758
Insects, True insects | 849
21441
123532 | Families
Genera
Species |
Taxom
|
Subclass
| Pterygota
Lang, 1888
Winged insects | 844
21413
123490 | Families
Genera
Species |
Taxom
|
Infraclass
| Neoptera
van der Wulp, 1890
Modern Wing-folding insects | 796
21025
121869 | Families
Genera
Species |
Taxom
|
Superorder
| Holometabola
Heider, 1889
Holometabolous Insects | 482
15573
99264 | Families
Genera
Species |
Taxom
|
Order
| Coleoptera
Linnaeus, 1758
Beetles, Coleopterans | 140
6932
53341 | Families
Genera
Species |
Taxom
|
Suborder
| Polyphaga
Emery, 1886
Polyphags | 126
6346
47559 | Families
Genera
Species |
Taxom
|
Infraorder
| Scarabaeiformia
Crowson, 1960
| 14
748
8916 | Families
Genera
Species |
Taxom
|
Superfamily
| Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles | 14
748
8916 | Families
Genera
Species |
Taxom
| | Family | Scarabaeidae
Latreille, 1802
Dung beetles, Flower beetles, Fruit chafers, Rain beetles | 564
6821 | Genera
Species |
Taxom
| | Subfamily | Scarabaeinae
Latreille, 1802
Dung Beetles, True dung beetles | 60
969 | Genera
Species |
Taxom
|
Genera for Southeast Asia
60 Genera with 969 Species #ACDEGHIKLMOPSTY
| Amphistomus van Lansberge, 1927 | | 3 | Species | Taxom | | Anoctus Sharp, 1942 | | 4 | Species | Taxom | | Caccobius Thomson, 1990 | | 13 | Species | Taxom | | Canthidium Erichson, 2006 | | 2 | Species | Taxom | Canthon Hoffmannsegg, 1817 | Tumblebug | 0 | Species | Taxom | | Canthonosoma MacLeay, 1871 | | 1 | Species | Taxom | | Cassolus Sharp, 1885 | | 6 | Species | Taxom | | Catharsius Hope, 1996 | | 8 | Species | Taxom | | Cheironitis van Lansberge, 1868 | | 1 | Species | Taxom | | Cleptocaccobius Cambefort, 0 | | 3 | Species | Taxom | | Copris Geoffroy, 1914 | Mexican dung beetle | 111 | Species | Taxom | | Coptodactyla Burmeister, 1976 | | 3 | Species | Taxom | | Cyobius Sharp, 1875 | | 5 | Species | Taxom | | Deltochilum Eschscholtz, 1887 | | 2 | Species | Taxom | Dendropaemon Perty, 1830 | | 0 | Species | Taxom | | Diastellopalpus van Lansberge, 1884 | | 1 | Species | Taxom | | Digitonthophagus Balthasar, 1787 | | 2 | Species | Taxom | | Drepanocerus Kirby, 1868 | | 3 | Species | Taxom | | Eodrepanus Barbero, Pal..., 1945 | | 2 | Species | Taxom | | Euoniticellus Janssens, 1987 | | 1 | Species | Taxom | | Euonthophagus Balthasar, 1790 | | 1 | Species | Taxom | | Garreta Janssens, 1854 | | 3 | Species | Taxom | | Gromphas Brullé, 1869 | | 1 | Species | Taxom | | Gymnopleurus Illiger, 1785 | | 4 | Species | Taxom | | Haroldius Boucomont, 1994 | | 23 | Species | Taxom | | Helictopleurus Orbigny, 1869 | | 1 | Species | Taxom | | Heliocopris Hope, 1859 | | 5 | Species | Taxom | | Hyalonthophagus Palestrini &..., 1913 | | 1 | Species | Taxom | | Ixodina Roth, 1909 | | 1 | Species | Taxom | | Kheper Janssens, 1843 | | 1 | Species | Taxom | | Larhodius Balthasar, 2003 | | 2 | Species | Taxom | | Latodrepanus Krikken, 1871 | | 1 | Species | Taxom | | Lepanus Balthasar, 1888 | | 1 | Species | Taxom | | Liatongus Reitter, 1798 | | 12 | Species | Taxom | | Macropanelus | | 3 | Species | Taxom | | Milichus Péringuey, 1871 | | 1 | Species | Taxom | | Ochicanthon Vaz-de-Mello, 2006 | | 40 | Species | Taxom | | Oniticellus Serville, 1879 | | 5 | Species | Taxom | | Onitis Fabricius, 1857 | | 23 | Species | Taxom | | Ontherus Erichson, 1869 | | 1 | Species | Taxom | | Onthophagus Latreille, 1963 | | 593 | Species | Taxom | | Onychothecus Boucomont, 1992 | | 4 | Species | Taxom | | Panelus Lewis, 1939 | | 18 | Species | Taxom | | Parachorius Harold, 1873 | | 15 | Species | Taxom | | Paragymnopleurus Shipp, 1997 | | 20 | Species | Taxom | | Paraphytus Harold, 1981 | | 7 | Species | Taxom | | Pedaria Laporte, 1922 | | 1 | Species | Taxom | | Phacosoma Boucomont, 1989 | | 2 | Species | Taxom | | Proagoderus Lansberge, 2002 | | 6 | Species | Taxom | | Scarabaeus Linnaeus, 1840 | | 1 | Species | Taxom | | Sinapisoma Boucomont, 1928 | | 1 | Species | Taxom | | Sinodrepanus Simonis, 2004 | | 6 | Species | Taxom | | Sisyphus Latreille, 0 | | 10 | Species | Taxom | | Sulcophanaeus Olsoufieff, 1856 | | 1 | Species | Taxom | | Synapsis Forster, 1933 | | 20 | Species | Taxom | | Temnoplectron Westwood, 2017 | | 2 | Species | Taxom | | Tesserodon Hope, 2017 | | 2 | Species | Taxom | | Tibiodrepanus Krikken, 1868 | | 3 | Species | Taxom | | Tiniocellus Péringuey, 2014 | | 1 | Species | Taxom | | Yvescambefortius Ochi & Kon, 1926 | | 1 | Species | Taxom |
Species list for Southeast Asia
969 Species and 47 Subspecies | Amphistomus alfurorum Huijbregts &..., 2007 | | | Amphistomus fasciculatus Gillet, 1927 | | | Amphistomus speculatus Gillet, 1927 | | | Anoctus laevis Sharp, 1875 | | | Anoctus rasilis Boucomont, 1921 | | | Anoctus scatonomoides Lansberge, 1885 | | | Anoctus siamensis Balthasar, 1942 | | | Caccobius (Caccophilus) tortus Sharp, 1875 | | | Caccobius tortus tortus | | | Caccobius (Caccophilus) unicornis Fabricius, 1798 | | | Caccobius bawangensis Ochi & Kon &..., 1997 | | | Caccobius bidentatus Boucomont, 1923 | | | Caccobius binodulus Harold, 1877 | | | Caccobius demangei Boucomont, 1919 | | | Caccobius dickinsoni Masumoto, Oc..., 2022 | | | Caccobius gilleti Balthasar, 1933 | | | Caccobius gonoderus Fairmaire, 1888 | | | Caccobius jessoensis Harold, 1867 | | | Caccobius maruyamai Masumoto, Oc..., 2012 | | | Caccobius masumotoi Cambefort, 1990 | | | Canthidium (Eucanthidium) macclevei Kohlmann & S..., 2006 | | | Canthidium delgadoi Kohlmann & S..., 2006 | | | Canthonosoma mastersi MacLeay, 1871 | | | Cassolus javanus Boucomont, 1914 | | | Cassolus minutus Boucomont, 1914 | | | Cassolus nudus Sharp, 1875 | | | Cassolus peninsularis Arrow, 1907 | | | Cassolus pongchaii Masumoto, 1989 | | | Cassolus sumatranus Lansberge, 1885 | | | Catharsius aethiops Sharp, 1875 | | | Catharsius birmanensis Lansberge, 1874 | | | Catharsius dayacus Lansberge, 1886 | | | Catharsius dux Harold, 1878 | | | Catharsius javanus Lansberge, 1886 | | | Catharsius molossus Linnaeus, 1758 | | | Catharsius quadridentatus Lansberge, 1885 | | | Catharsius renaudpauliani Ochi & Kon, 1996 | | | Cheironitis hoplosternus Harold, 1868 | | | Cleptocaccobius simplex Boucomont, 1923 | | | Cleptocaccobius simplex meridionalis Cambefort, 1985 | | | Cleptocaccobius simplex simplex | | | Copris (Copris) cambodiensis Ochi, Kon & ..., 2008 | | | Copris cambodiensis cambodiensis | | | Copris cambodiensis masumotoi Ochi, Kon & ..., 2011 | | | Copris (Copris) kachinensis Ochi, Kon & ..., 2008 | | | Copris (Copris) punctatus Gillet, 1910 | | | Copris punctatus punctatus | | | Copris (Microcopris) propinquus Felsche, 1910 | | | Copris (Paracopris) cariniceps Felsche, 1910 | | | Copris (Sinocopris) barclayi Ochi, Kon & Bai, 2009 | | | Copris (Sinocopris) uenoi Hanboonsong ..., 2003 | | | Copris agnus Sharp, 1875 | | | Copris angusticornis Arrow, 1933 | | | Copris anomiopseoides (Boucomont, 1924) | ⇒ Paracopris anomiopseoides Boucomont, 1924 | | Copris arimotoi Ochi & Araya, 1994 | | | Copris arrowi Felsche, 1910 | | | Copris bartolozzi Ochi, Kon & ..., 2019 | | | Copris bellator Chevrolat, 1844 | | | Copris bengalensis Shiino, 1978 | | | Copris calvus Sharp, 1875 | | | Copris caobangensis Bui, Dumack ..., 2018 | | | Copris carinicus Gillet, 1910 | | | Copris celebensis Lansberge, 1886 | | | Copris chinmontanus Ochi, Kon & ..., 2019 | | | Copris confucius Harold, 1877 | | | Copris corpulentus Gillet, 1910 | | | Copris doriae Harold, 1877 | | | Copris elphenor Klug, 1855 | | | Copris erratus Lansberge, 1886 | | | Copris erratus erratus | | | Copris erratus masayukii Ochi & Kon, 2004 | | | Copris fricator Fabricius, 1787 | | | Copris fujiokai Ochi & Kon, 1996 | | | Copris fujiokai poringensis Ochi & Kon, 2005 | | | Copris furciceps Felsche, 1910 | | | Copris gibbulus Lansberge, 1886 | | | Copris gibbulus borneensis Ochi & Kon, 2005 | | | Copris gibbulus gibbulus | | | Copris gladiator Arrow, 1933 | | | Copris hatayamai Ochi & Araya, 1992 | | | Copris hidakai Ochi & Kon, 1996 | | | Copris hosakai Ochi & Kon, 2014 | | | Copris interioris Kolbe, 1897 | | | Copris iris Sharp, 1875 | | | Copris javanensis Bui, Ziegler..., 2019 | | | Copris johkii Ochi & Kon, 2014 | | | Copris jucundus Gillet, 1932 | | | Copris kasagii Ochi & Kon, 1996 | | | Copris kimioi Ochi & Kon, 2021 | | | Copris kiriromensis Ochi & Kon, 2021 | | | Copris kiuchii Masumoto, 1989 | | | Copris kusakabei Ochi, Kon & ..., 2005 | | | Copris laevigatus (Gillet, 1927) | ⇒ Onthophagus laevigatus Fabricius, 1798 | | Copris lannathai Hanboonsong ..., 2003 | | | Copris laosensis Ochi & Kon, 2021 | | | Copris lugubris Boheman, 1858 | | | Copris macacus Lansberge, 1886 | | | Copris magicus Harold, 1881 | | | Copris marcoberrai Ochi, Kon & ..., 2019 | | | Copris marcus Gillet, 1921 | | | Copris micros Ochi & Kon, 2021 | | | Copris mindorensis Ochi, 1992 | | | Copris minutus Drury, 1773 | Small Black Dung Beetle | | Copris miyakei Ochi & Kon, 2004 | | | Copris moffartsi Gillet, 1907 | | | Copris mongkhoni Hanboonsong ..., 2003 | | | Copris montreuili Ochi, Kon & ..., 2019 | | | Copris nevinsoni Waterhouse, 1891 | Warton meshweaver | | Copris nonarmatus Ochi & Kon, 2021 | | | Copris novaki Zelenka, 1992 | | | Copris numa Lansberge, 1886 | Braken Bat Cave meshweaver | | Copris obenbergeri Balthasar, 1933 | Government Canyon Bat Cave meshweaver | | Copris olivieri Illiger, 1800 | | | Copris orphanus (Guérin-Ménev..., 1847) | ⇒ Copris evanidus Klug, 1855 | | Copris parapecuarius Ochi, Kon & ..., 2008 | | | Copris parasonensis Ochi, Kon & ..., 2019 | | | Copris paraszechouanicus Ochi, Kon & ..., 2018 | | | Copris pedarioides Lansberge, 1886 | | | Copris poggii Ochi & Kon, 2005 | | | Copris popaensis Ochi, Kon & ..., 2019 | | | Copris pseudochus basipunctulatus Ochi, Kon & ..., 2020 | | | Copris pseudochus chuduraziensis Ochi, Kon & ..., 2019 | | | Copris pseudochus pseudochus | | | Copris punctipennis Boucomont, 1914 | | | Copris punctulatus Wiedemann, 1823 | | | Copris punctulatus punctulatus | | | Copris punctulatus septentrionalis Ochi, Kon & ..., 2019 | | | Copris ramosiceps Gillet, 1921 | | | Copris reflexus (Fabricius, 1787) | ⇒ Copris minutus Drury, 1773 | | Copris repertus Walker, 1858 | | | Copris ribbei Lansberge, 1886 | | | Copris riekoae Ochi & Kon, 2003 | | | Copris sarpedon Harold, 1868 | | | Copris saundersi Harold, 1869 | | | Copris scorpio Zelenka, 1993 | | | Copris siamensis Gillet, 1910 | | | Copris signatus Walker, 1858 | | | Copris sinicus Hope, 1842 | | | Copris sinon Olivier, 1789 | | | Copris sonensis Bui, Dumack ..., 2018 | | | Copris sphaeropterus Harold, 1877 | | | Copris spinator Harold, 1881 | | | Copris sumatrensis Gillet, 1921 | | | Copris szechouanicus Balthasar, 1958 | | | Copris takanoi Ochi, Kon & Bai, 2020 | | | Copris tetraodon Gillet, 1910 | | | Copris tricorniprothoracicus Ochi, Kon & ..., 2020 | | | Copris tsukamotoi Ochi, Kon & ..., 2008 | | | Copris vietnamicus Kabakov, 1994 | | | Copris vietnamimontanus Ochi, Kon & ..., 2018 | | | Copris vitalisi Gillet, 1921 | | | Copris wiesei Kolbe, 1914 | | | Coptodactyla glabricollis Hope, 1842 | | | Coptodactyla merdeka Reid, 2000 | | | Coptodactyla torresica Mathews, 1976 | | | Cyobius brendelli Ochi, Kon & ..., 2010 | | | Cyobius cheyi Ochi, Kon & ..., 2006 | | | Cyobius mendeli Ochi, Kon & ..., 2010 | | | Cyobius tsukamotoi Ochi, Kon & ..., 2008 | | | Cyobius wallacei Sharp, 1875 | | | Deltochilum mexicanum Burmeister, 1848 | | | Deltochilum scabriusculum Bates, 1887 | Rockpool blenny | | Diastellopalpus lamellicollis Quedenfeldt, 1884 | | | Digitonthophagus bonasus Fabricius, 1775 | | | Digitonthophagus gazella Fabricius, 1787 | Brown dung beetle | | Drepanocerus planus Fabricius, 1801 | | | Drepanocerus runicus (Arrow, 1909) | ⇒ Ixodina runica Arrow, 1909 | | Drepanocerus sinicus (Harold, 1868) | ⇒ Tibiodrepanus sinicus Harold, 1868 | | Eodrepanus integriceps Janssens, 1953 | | | Eodrepanus striatulus Paulian, 1945 | | | Euoniticellus cambeforti Masumoto, 1987 | | | Euonthophagus gibbosus Scriba, 1790 | | | Garreta azareus | | | Garreta gilleti Garreta, 1914 | | | Garreta ruficornis Motschulsky, 1854 | | | Gromphas inermis Harold, 1869 | | | Gymnopleurus aethiops (Sharp, 1875) | ⇒ Paragymnopleurus melanarius Harold, 1867 | | Gymnopleurus geoffroyi Fuessly, 1775 | | | Gymnopleurus parvus MacLeay, 1821 | | | Gymnopleurus sinuatus (Geoffroy, 1785) | ⇒ Gymnopleurus geoffroyi Fuessly, 1775 | | Haroldius borneensis Paulian, 1993 | | | Haroldius brendelli Krikken & Hu..., 2009 | | | Haroldius cambeforti Krikken & Hu..., 2006 | | | Haroldius celebensis Krikken & Hu..., 2006 | | | Haroldius discoidalis Paulian, 1993 | | | Haroldius fairmairei Boucomont, 1914 | | | Haroldius fleutiauxi Paulian, 1945 | | | Haroldius globosus Boucomont, 1925 | | | Haroldius kawadai Masumoto, 1995 | | | Haroldius kolaka Krikken & Hu..., 2006 | | | Haroldius loebli Paulian, 1987 | | | Haroldius maruyamai Utsunomiya &..., 2005 | | | Haroldius oharai Ochi, Kon & ..., 2009 | | | Haroldius pahangensis Kral, 2003 | | | Haroldius pauliani Scheuern, 1995 | | | Haroldius penelopae Krikken & Hu..., 2006 | | | Haroldius perroti Paulian, 1939 | | | Haroldius philippinensis Pereira, 1954 | | | Haroldius rugatulus Boucomont, 1914 | | | Haroldius stevensi Arrow, 1931 | | | Haroldius sumatranus Paulian & Sc..., 1994 | | | Haroldius tangkoko Krikken & Hu..., 2006 | | | Haroldius thailandensis Paulian & Sc..., 1994 | | | Helictopleurus giganteus Harold, 1869 | | | Heliocopris bucephalus Fabricius, 1775 | | | Heliocopris dominus Bates, 1868 | | | Heliocopris hamadryas Fabricius, 1775 | | | Heliocopris staudingeri Kolbe, 1893 | | | Heliocopris tyrannus Thomson, 1859 | | | Hyalonthophagus alcedo Orbigny, 1913 | | | Ixodina runica Arrow, 1909 | | | Kheper prodigiosus Erichson, 1843 | | | Larhodius hashimi Masumoto & U..., 2003 | | | Larhodius maruyamai Masumoto & U..., 2003 | | | Latodrepanus caelatus Gerstaecker, 1871 | | | Lepanus pygmaeus MacLeay, 1888 | | | Liatongus affinis Arrow, 1908 | | | Liatongus capricornis Arrow, 1931 | | | Liatongus femoratus Illiger, 1800 | | | Liatongus gagatinus Hope, 1831 | | | Liatongus martialis Harold, 1879 | | | Liatongus minutus Motschulsky, 1860 | | | Liatongus phanaeoides Westwood, 1839 | | | Liatongus rhadamistus Fabricius, 1775 | | | Liatongus triacanthus Boucomont, 1920 | | | Liatongus tridentatus Boucomont, 1919 | | | Liatongus venator Fabricius, 1801 | | | Liatongus vertagus Fabricius, 1798 | | | Macropanelus fujitai Ochi, Kon & ..., 2008 | | | Macropanelus karasuyamai Ochi, Kon & ..., 2008 | | | Macropanelus sumatrensis Ochi & Kon &..., 1998 | | | Milichus picticollis Gerstaecker, 1871 | | | Ochicanthon cambeforti Ochi & Kon &..., 1997 | | | Ochicanthon crockermontis Krikken & Hu..., 2007 | | | Ochicanthon crypticus Krikken & Hu..., 2007 | | | Ochicanthon danum Krikken & Hu..., 2007 | | | Ochicanthon dulitmontis Krikken & Hu..., 2007 | | | Ochicanthon dytiscoides Boucomont, 1914 | | | Ochicanthon edmondsi Krikken & Hu..., 2007 | | | Ochicanthon fernandoi Sabu & Latha, 2011 | | | Ochicanthon gangkui Ochi & Kon &..., 1997 | | | Ochicanthon hanskii Krikken & Hu..., 2007 | | | Ochicanthon hikidai Ochi & Kon &..., 1997 | | | Ochicanthon javanus Krikken & Hu..., 2007 | | | Ochicanthon karasuyamai Ochi, Kon & ..., 2007 | | | Ochicanthon kikutai Ochi, Ueda &..., 2006 | | | Ochicanthon kimanis Krikken & Hu..., 2007 | | | Ochicanthon laetus Arrow, 1931 | | | Ochicanthon maryatiae Ochi, Ueda &..., 2006 | | | Ochicanthon masumotoi Ochi & Araya, 1996 | | | Ochicanthon mulu Krikken & Hu..., 2007 | | | Ochicanthon neglectus Krikken & Hu..., 2007 | | | Ochicanthon niinoii Ochi & Kon, 2014 | | | Ochicanthon niisatoi Ochi, Kon & ..., 2017 | | | Ochicanthon obscurus Boucomont, 1920 | | | Ochicanthon ochii Hanboonsong ..., 2001 | | | Ochicanthon oharai Ochi, Kon, H..., 2008 | | | Ochicanthon okudai Ochi & Kon, 2014 | | | Ochicanthon parantisae Ochi & Kon &..., 1997 | | | Ochicanthon peninsularis Krikken & Hu..., 2007 | | | Ochicanthon philippinense Ochi, 1990 | | | Ochicanthon punctatus Boucomont, 1914 | | | Ochicanthon ranongensis Ochi, Kon & ..., 2020 | | | Ochicanthon rombauti Krikken & Hu..., 2007 | | | Ochicanthon takakui Ochi, Kon, H..., 2008 | | | Ochicanthon takuapaensis Ochi, Kon & ..., 2020 | | | Ochicanthon tambunan Krikken & Hu..., 2007 | | | Ochicanthon thai Paulian, 1987 | | | Ochicanthon thailandicum Masumoto, 1989 | | | Ochicanthon uedai Ochi, Kon, H..., 2008 | | | Ochicanthon vazdemelloi (Latha & Sabu, 2011) | ⇒ Ochicanthon fernandoi Sabu & Latha, 2011 | | Ochicanthon woroae Ochi, Ueda &..., 2006 | | | Oniticellus caelatus (Gerstaecker, 1871) | ⇒ Latodrepanus caelatus Gerstaecker, 1871 | | Oniticellus cinctus Fabricius, 1775 | Bordered dung beetle | | Oniticellus freyi Balthasar, 1941 | | | Oniticellus rhadamistus Fabricius, 1775 | | | Oniticellus tessellatus Harold, 1879 | | | Onitis bordati Cambefort, 1988 | | | Onitis corydon Boisduval, 1835 | | | Onitis crenatus Reiche, 1847 | | | Onitis cryptodus Gillet, 1918 | | | Onitis excavatus Arrow, 1931 | | | Onitis falcatus Wulfen, 1786 | | | Onitis feae Felsche, 1907 | | | Onitis hageni Lansberge, 1886 | | | Onitis intermedius Frivaldsky, 1892 | | | Onitis lama Lansberge, 1875 | | | Onitis lansbergei Sharp, 1875 | | | Onitis ludekingi Lansberge, 1875 | | | Onitis niger (Lansberge, 1875) | ⇒ Liatongus femoratus Illiger, 1800 | | Onitis phartopus Lansberge, 1875 | | | Onitis philemon (Fabricius, 1801) | ⇒ Onitis subopacus Arrow, 1931 | | Onitis singhalensis Lansberge, 1875 | | | Onitis siva Gillet, 1911 | | | Onitis spinipes Drury, 1770 | | | Onitis subcrenatus Kolbe, 1897 | | | Onitis subopacus Arrow, 1931 | | | Onitis uncinatus Klug, 1855 | | | Onitis virens Lansberge, 1875 | | | Onitis viridulus (Boheman, 1857) | ⇒ Onitis pecuarius Lansberge, 1875 | | Ontherus cephalotes Harold, 1869 | | | Onthophagus (Colobonthophagus) aenescens Wiedemann, 1823 | | | Onthophagus (Colobonthophagus) caprai Frey, 1956 | | | Onthophagus (Colobonthophagus) occipitalis Lansberge, 1885 | | | Onthophagus (Colobonthophagus) piceorufulus Kabakov, 1994 | | | Onthophagus (Colobonthophagus) poggii Scheuern, 1996 | | | Onthophagus (Colobonthophagus) thai Kabakov, 1994 | | | Onthophagus (Colobonthophagus) tragus Fabricius, 1792 | | | Onthophagus (Digitonthophagus) bonasus Fabricius, 1775 | Dung beetle | | Onthophagus (Furconthophagus) boucomonti Paulian, 1931 | | | Onthophagus (Furconthophagus) karenensis Masumoto, Oc..., 2008 | | | Onthophagus (Furconthophagus) khonkaenus Masumoto, Oc..., 2008 | | | Onthophagus (Furconthophagus) lilliputanus Lansberge, 1883 | | | Onthophagus (Furconthophagus) papulatorius Kabakov, 2008 | | | Onthophagus (Furconthophagus) papulatus Boucomont, 1914 | | | Onthophagus (Furconthophagus) promontorii Boucomont, 1924 | | | Onthophagus (Gibbonthophagus) chineicus Kabakov, 1998 | | | Onthophagus (Gibbonthophagus) denticollis Lansberge, 1883 | | | Onthophagus (Gibbonthophagus) duporti Boucomont, 1914 | | | Onthophagus (Gibbonthophagus) euryceros Kabakov, 1998 | | | Onthophagus (Gibbonthophagus) fujiii Ochi & Kon, 1995 | | | Onthophagus (Gibbonthophagus) gracilipes Boucomont, 1914 | | | Onthophagus (Gibbonthophagus) hiabunicus Kabakov, 1998 | | | Onthophagus (Gibbonthophagus) kimioi Ochi, Kon & ..., 2011 | | | Onthophagus (Gibbonthophagus) lenis Kabakov, 1998 | | | Onthophagus (Gibbonthophagus) limbatus Herbst, 1789 | | | Onthophagus (Gibbonthophagus) luridipennis Boheman, 1858 | | | Onthophagus (Gibbonthophagus) nigriobscurior Ochi, Kon & ..., 2009 | | | Onthophagus (Gibbonthophagus) palawanicus Ochi & Kon, 2004 | | | Onthophagus (Gibbonthophagus) parviobscurior Ochi, Kon & ..., 2009 | | | Onthophagus (Gibbonthophagus) penmani Masumoto, Oc..., 2002 | | | Onthophagus (Gibbonthophagus) privus Kabakov, 1998 | | | Onthophagus (Gibbonthophagus) rufiobscurior Ochi, Kon & ..., 2009 | | | Onthophagus (Gibbonthophagus) sasajii Ochi & Kon, 2001 | | | Onthophagus (Gibbonthophagus) semipersonatus Ochi, Kon & ..., 2009 | | | Onthophagus (Gibbonthophagus) sunantaae Masumoto, 1989 | | | Onthophagus (Gibbonthophagus) taurinus White, 1844 | | | Onthophagus (Gibbonthophagus) viridicervicapra Ochi, Kon & ..., 2009 | | | Onthophagus (Indachorius) arai Masumoto, 1989 | | | Onthophagus (Indachorius) baykanus Kabakov, 1994 | | | Onthophagus (Indachorius) bonengus Kabakov, 1994 | | | Onthophagus (Indachorius) danumensis (Ochi, Kon & ..., 2009) | ⇒ Onthophagus danumicus Huijbregts &..., 2012 | | Onthophagus (Indachorius) doisuthepensis Masumoto, 1989 | | | Onthophagus (Indachorius) hsui Masumoto, Ch..., 2004 | | | Onthophagus (Indachorius) koshunensis Balthasar, 1941 | | | Onthophagus (Indachorius) lannamiibun Masumoto, Oc..., 2002 | | | Onthophagus (Indachorius) maesaensis Masumoto, Oc..., 2002 | | | Onthophagus (Indachorius) masaoi Ochi, 1992 | | | Onthophagus (Indachorius) mendeli Ochi, Kon & ..., 2009 | | | Onthophagus (Indachorius) mongkhoni Masumoto, Ha..., 2002 | | | Onthophagus (Indachorius) nongkaiensis Masumoto, Oc..., 2002 | | | Onthophagus (Indachorius) octonaevus Kabakov, 1994 | | | Onthophagus (Indachorius) paramasaoi Ochi, Kon & ..., 2009 | | | Onthophagus (Indachorius) phetchabunensis Masumoto, Oc..., 2002 | | | Onthophagus (Indachorius) phuquoci Paulian, 1945 | | | Onthophagus (Indachorius) pseudoworoae Ochi, Kon & ..., 2009 | | | Onthophagus (Indachorius) scotti Masumoto, Oc..., 2002 | | | Onthophagus (Indachorius) semidanumensis Ochi, Kon & ..., 2009 | | | Onthophagus (Indachorius) subsapaensis Kabakov, 1994 | | | Onthophagus (Indachorius) tiamicus Kabakov, 1994 | | | Onthophagus (Indachorius) tongbantumi Masumoto, Oc..., 2002 | | | Onthophagus (Macronthophagus) curvicarinatus Boucomont, 1914 | | | Onthophagus (Macronthophagus) diabolicus Harold, 1877 | | | Onthophagus (Macronthophagus) manipurensis Arrow, 1907 | | | Onthophagus (Macronthophagus) menieri Ochi, 2003 | | | Onthophagus (Macronthophagus) rubricollis Hope, 1831 | | | Onthophagus (Macronthophagus) uenoi Ochi, 1995 | | | Onthophagus uenoi borneomontanus Ochi, 2003 | | | Onthophagus uenoi uenoi Ochi, 1995 | | | Onthophagus (Matashia) anguliceps (Boucomont, 1914) | ⇒ Onthophagus avocetta Arrow, 1933 | | Onthophagus (Micronthophagus) falsivigilans Masumoto, 1995 | | | Onthophagus (Micronthophagus) gigantivigilans Masumoto, Ha..., 2002 | | | Onthophagus (Micronthophagus) vigilans Boucomont, 1921 | | | Onthophagus (Onthophagiellus) kondaoensis Kabakov, 1994 | | | Onthophagus (Onthophagiellus) opacifalculatus Ochi, Kon & ..., 2009 | | | Onthophagus (Onthophagiellus) parafalculatus Ochi, Kon & ..., 2009 | | | Onthophagus (Onthophagiellus) thanwaakhomus Masumoto, 1992 | | | Onthophagus (Onthophagus) acuticornis Endrödi, 1973 | | | Onthophagus (Onthophagus) angustatus Boucomont, 1914 | | | Onthophagus (Onthophagus) apilularius Masumoto, 1995 | | | Onthophagus (Onthophagus) barbieri Paulian, 1978 | | | Onthophagus (Onthophagus) batesi Howden and C..., 1963 | | | Onthophagus (Onthophagus) bokiaunus Masumoto, 1995 | | | Onthophagus (Onthophagus) chaiyaphumensis Masumoto, Oc..., 2002 | | | Onthophagus (Onthophagus) dapcauensis Boucomont, 1921 | | | Onthophagus (Onthophagus) doiinthanonensis Masumoto, 1989 | | | Onthophagus (Onthophagus) doipuiensis Masumoto, 1989 | | | Onthophagus (Onthophagus) doitungensis Masumoto, Oc..., 2002 | | | Onthophagus (Onthophagus) dubernardi Boucomont, 1914 | | | Onthophagus (Onthophagus) embersoni Masumoto, Ha..., 2002 | | | Onthophagus (Onthophagus) gorochovi Kabakov, 1998 | | | Onthophagus (Onthophagus) gosoli Masumoto, 1989 | | | Onthophagus (Onthophagus) grandivigilans Masumoto, 1995 | | | Onthophagus (Onthophagus) hastifer Lansberge, 1885 | | | Onthophagus (Onthophagus) horii Ochi, Kon & ..., 2009 | | | Onthophagus (Onthophagus) ishiii Ochi & Kon, 1995 | | | Onthophagus (Onthophagus) javaecola Balthasar, 1959 | | | Onthophagus (Onthophagus) kanyaayonus Masumoto, 1992 | | | Onthophagus (Onthophagus) khonmiinitnoi Masumoto, 1990 | | | Onthophagus (Onthophagus) kinhthaicus Kabakov, 1998 | | | Onthophagus (Onthophagus) kiuchii Masumoto, 1995 | | | Onthophagus (Onthophagus) krakadaakhomus Masumoto, 1992 | | | Onthophagus (Onthophagus) lakyim Masumoto, 1990 | | | Onthophagus (Onthophagus) lindaae Masumoto, 1989 | | | Onthophagus (Onthophagus) maleengnaafon Masumoto, 1990 | | | Onthophagus (Onthophagus) maleengnoi Masumoto, 1990 | | | Onthophagus (Onthophagus) maniti Masumoto, 1989 | | | Onthophagus (Onthophagus) matsudai Ochi & Kon, 2004 | | | Onthophagus (Onthophagus) mulleri van Lansberge, 1883 | | | Onthophagus (Onthophagus) naaroon Masumoto, 1990 | | | Onthophagus (Onthophagus) namnaoensis Masumoto, Oc..., 2002 | | | Onthophagus (Onthophagus) nasonis Kabakov, 1998 | | | Onthophagus (Onthophagus) orientalis (Harold, 1868) | ⇒ Onthophagus fissinasus Fairmaire, 1895 | | Onthophagus (Onthophagus) phraoensis Masumoto, 1990 | | | Onthophagus (Onthophagus) phrutsaphaakhomus Masumoto, 1992 | | | Onthophagus (Onthophagus) proletarius Harold, 1875 | | | Onthophagus (Onthophagus) pseudoarmatus Balthasar, 1944 | | | Onthophagus (Onthophagus) pseudohystrix Masumoto, 1995 | | | Onthophagus (Onthophagus) punneeae Masumoto, 1989 | | | Onthophagus (Onthophagus) ratchasimaensis Masumoto, Ha..., 2002 | | | Onthophagus (Onthophagus) rugulosus Harold, 1886 | | | Onthophagus (Onthophagus) sakaeratensis Masumoto, Ha..., 2002 | | | Onthophagus (Onthophagus) salvazai Paulian, 1978 | | | Onthophagus (Onthophagus) singhaakhomus Masumoto, 1992 | | | Onthophagus (Onthophagus) taayai Masumoto, 1995 | | | Onthophagus (Onthophagus) taurus Schreber, 1759 | Bull Headed Dung Beetle | | Onthophagus (Onthophagus) terminatus Eschscholtz, 1822 | | | Onthophagus (Onthophagus) tholaayi Masumoto, 1990 | | | Onthophagus (Onthophagus) trituber Wiedemann, 1823 | | | Onthophagus (Onthophagus) tumami Masumoto, Oc..., 2002 | | | Onthophagus (Onthophagus) wangnamkhieoensis Masumoto, Ha..., 2002 | | | Onthophagus (Onthophagus) yamaokai Masumoto, Oc..., 2002 | | | Onthophagus (Palaeonthophagus) lemur Fabricius, 1781 | | | Onthophagus (Palaeonthophagus) muelleri (Lansberge, 1883) | ⇒ Onthophagus statzi Krell, 1990 | | Onthophagus (Palaeonthophagus) novaki Boucomont, 1927 | | | Onthophagus (Palaeonthophagus) olsoufieffi Boucomont, 1924 | | | Onthophagus (Palaeonthophagus) vacca Linnaeus, 1767 | | | Onthophagus (Paraphanaeomorphus) carinensis Boucomont, 1914 | | | Onthophagus (Paraphanaeomorphus) comottoides Kabakov, 1998 | | | Onthophagus (Paraphanaeomorphus) janushevi Kabakov, 1983 | | | Onthophagus (Paraphanaeomorphus) napolovi Kabakov, 1998 | | | Onthophagus (Paraphanaeomorphus) pseudojavanus Paulian, 1931 | | | Onthophagus (Paraphanaeomorphus) quadricolor Kabakov, 1983 | | | Onthophagus (Paraphanaeomorphus) semipiceus Kabakov, 1998 | | | Onthophagus (Paraphanaeomorphus) strnadi Kabakov, 1998 | | | Onthophagus (Paraphanaeomorphus) vaulogeri Boucomont, 1923 | | | Onthophagus (Parascatonomus) anceyi Boucomont, 1921 | | | Onthophagus (Parascatonomus) damaki Masumoto, Ha..., 2002 | | | Onthophagus (Parascatonomus) koichii (Ochi & Araya, 1994) | ⇒ Caccobius jessoensis Harold, 1867 | | Onthophagus (Parascatonomus) leytensis Ochi & Araya, 1994 | | | Onthophagus (Parascatonomus) negrosensis Ochi & Araya, 1994 | | | Onthophagus (Parascatonomus) pseudorudis Ochi & Araya, 1994 | | | Onthophagus (Parascatonomus) quezonensis Ochi & Araya, 1994 | | | Onthophagus (Parascatonomus) satoi Ochi, Shimad..., 2003 | | | Onthophagus (Parascatonomus) shizumui Ochi & Araya, 1994 | | | Onthophagus (Parascatonomus) takedai Ochi & Araya, 1994 | | | Onthophagus (Parascatonomus) takeshii Ochi & Araya, 1994 | | | Onthophagus (Parascatonomus) yukae Masumoto, Oc..., 2002 | | | Onthophagus (Parentius) kabakovi Martin-Piera, 1985 | | | Onthophagus (Paronthophagus) hidakai Ochi & Kon, 1995 | | | Onthophagus (Phanaeomorphus) johkii Ochi & Kon, 1994 | | | Onthophagus (Phanaeomorphus) zetteli Masumoto, Oc..., 2002 | | | Onthophagus (Pseudophanaeomorphus) parachandrai Ochi, Kon & ..., 2009 | | | Onthophagus (Serrophorus) avocettoides Kabakov, 1983 | | | Onthophagus (Serrophorus) laevis (Harold, 1880) | ⇒ Onthophagus laevissimus d´Orbigny, 1905 | | Onthophagus laevis asiaticus Boucomont, 1919 | | | Onthophagus laevis laevis | | | Onthophagus laevis lampromelas Fairmaire, 1891 | | | Onthophagus (Serrophorus) rectecornutus Lansberge, 1883 | | | Onthophagus (Serrophorus) sagittarius Fabricius, 1775 | Sagittarius dung beetle | | Onthophagus (Sinonthophagus) nampatensis Tarasov & Ka..., 2010 | | | Onthophagus (Sunenaga) streltsovi Tarasov & Ka..., 2010 | | | Onthophagus abei Ochi & Kon, 2015 | | | Onthophagus academus Balthasar, 1946 | | | Onthophagus aceroides Krikken & Hu..., 2013 | | | Onthophagus aemulus Gillet, 1930 | | | Onthophagus aereomaculatus Boucomont, 1914 | | | Onthophagus aereopictus Boucomont, 1914 | | | Onthophagus aereopictus aereopictus | | | Onthophagus aereopictus rubropictus Boucomont, 1924 | | | Onthophagus akhaus Masumoto, Oc..., 2007 | | | Onthophagus alfuricus Huijbregts &..., 2012 | | | Onthophagus allojavanus Huijbregts &..., 2011 | | | Onthophagus aloysiellus Zunino, 1977 | | | Onthophagus ambang Krikken & Hu..., 2011 | | | Onthophagus ambang ambang | | | Onthophagus ambang morowali Krikken & Hu..., 2011 | | | Onthophagus amphicoma Boucomont, 1914 | | | Onthophagus andonarensis Boucomont, 1914 | | | Onthophagus angkhanensis Masumoto, Oc..., 2013 | | | Onthophagus annulopunctatus Krikken & Hu..., 2017 | | | Onthophagus aphodioides Lansberge, 1883 | | | Onthophagus apomontanus Ochi, Kon & ..., 2023 | | | Onthophagus apunneea Masumoto, Oc..., 2007 | | | Onthophagus arayai Ochi & Kon, 2007 | | | Onthophagus ardoini Frey, 1971 | | | Onthophagus armatus Blanchard, 1853 | | | Onthophagus asiaticus Endrödi, 1973 | | | Onthophagus atropolitus d´Orbigny, 1902 | | | Onthophagus aureopilosus Boucomont, 1914 | | | Onthophagus australis Guérin-Ménev..., 1830 | Southern dung beetle | | Onthophagus avocetta Arrow, 1933 | | | Onthophagus axanicus Ochi, Kon & ..., 2021 | | | Onthophagus azusae Ochi & Kon, 2006 | | | Onthophagus babirussa Eschscholtz, 1822 | | | Onthophagus baenzigeri Masumoto, Oc..., 2007 | | | Onthophagus baeri Boucomont, 1914 | | | Onthophagus bakeri (Boucomont, 1919) | ⇒ Panelus bakeri Boucomont, 1921 | | Onthophagus balthasari Vsetecka, 1939 | | | Onthophagus bangueyensis Boucomont, 1914 | | | Onthophagus baolocensis Ochi & Kon, 2015 | | | Onthophagus baramtagal Ochi, Kon & ..., 2016 | | | Onthophagus batillifer Harold, 1875 | | | Onthophagus batui Huijbregts &..., 2009 | | | Onthophagus begoniophilus Krikken & Hu..., 2017 | | | Onthophagus bellus d´Orbigny, 1905 | | | Onthophagus bicolensis Ochi & Kon, 2006 | | | Onthophagus bicolensis asin Ochi & Kon, 2006 | | | Onthophagus bicolensis bicolensis | | | Onthophagus bicuneus Kabakov, 2008 | | | Onthophagus bifasciatus Fabricius, 1781 | | | Onthophagus biscarinulatus Huijbregts &..., 2012 | | | Onthophagus bison Boucomont, 1919 | | | Onthophagus bisscrutator Krikken & Hu..., 2017 | | | Onthophagus bistiniocelloides Krikken, 1986 | | | Onthophagus bituberoculus Krikken & Hu..., 2013 | | | Onthophagus blanchardi Harold, 1869 | | | Onthophagus blumei (Lansberge, 1883) | ⇒ Onthophagus avocetta Arrow, 1933 | | Onthophagus bongkudai Krikken & Hu..., 2017 | | | Onthophagus bonorae Zunino, 1976 | | | Onthophagus borneensis Harold, 1877 | | | Onthophagus borneotagal Ochi, Kon & ..., 2016 | | | Onthophagus bourgognei Paulian, 1945 | | | Onthophagus brutus Arrow, 1931 | | | Onthophagus bunamin Matthews, 1972 | | | Onthophagus calcaratus Boucomont, 1920 | | | Onthophagus cameloides d´Orbigny, 1900 | | | Onthophagus carinulatus Harold, 1877 | | | Onthophagus carpophilus Pereira & Ha..., 1961 | | | Onthophagus caviceps Frey, 1962 | | | Onthophagus cervicapra Boucomont, 1914 | | | Onthophagus cervus (Fabricius, 1798) | ⇒ Onthophagus falsus Gillet, 1925 | | Onthophagus chandrai Ochi, 2007 | | | Onthophagus chebaicus Boucomont, 1919 | | | Onthophagus chetroiensis Masumoto, Oc..., 2012 | | | Onthophagus cheyi Ochi & Kon, 2006 | | | Onthophagus chiangdaolekus Masumoto, Oc..., 2020 | | | Onthophagus chinbucephalus Ochi & Kon, 2018 | | | Onthophagus chinvigilans Ochi & Kon, 2018 | | | Onthophagus chumphonensis Masumoto, Oc..., 2008 | | | Onthophagus circulifer Arrow, 1931 | | | Onthophagus citreum Boucomont, 1919 | | | Onthophagus clermonti Paulian, 1931 | | | Onthophagus clivimerus Huijbregts &..., 2011 | | | Onthophagus cognatus Boucomont, 1921 | | | Onthophagus colffsi Lansberge, 1883 | | | Onthophagus collinsi Krikken & Hu..., 1987 | | | Onthophagus commottoides Kabakov, 1998 | | | Onthophagus comottoi Lansberge, 1885 | | | Onthophagus consentaneus Harold, 1867 | Common northern dung beetle | | Onthophagus coomani Paulian, 1931 | | | Onthophagus coprimorphoides Huijbregts &..., 2012 | | | Onthophagus coracinoides Kabakov, 1983 | | | Onthophagus coracinus Boucomont, 1914 | | | Onthophagus crassicollis Boucomont, 1913 | | | Onthophagus cribratus Lansberge, 1883 | | | Onthophagus cuneus Boucomont, 1924 | | | Onthophagus cupreopastillatus Ochi & Kon, 2006 | | | Onthophagus cupreotagal Ochi, Kon & ..., 2016 | | | Onthophagus dandalu Matthews, 1972 | | | Onthophagus danumicus Huijbregts &..., 2012 | | | Onthophagus dapitanensis Boucomont, 1919 | | | Onthophagus dayacus Boucomont, 1914 | | | Onthophagus deflexicollis Lansberge, 1883 | | | Onthophagus deliensis Lansberge, 1885 | | | Onthophagus deliensis bawangicus Ochi, Kon & ..., 2009 | | | Onthophagus deliensis deliensis | | | Onthophagus denticornis Boucomont, 1914 | | | Onthophagus depressicollis Boucomont, 1914 | | | Onthophagus difficilis (Le Guillou, 1844) | ⇒ Onthophagus vacca Linnaeus, 1767 | | Onthophagus dinoderus d´Orbigny, 1913 | | | Onthophagus doriae Harold, 1877 | | | Onthophagus dorsofasciatus Fairmaire, 1893 | | | Onthophagus drescheri Paulian, 1939 | | | Onthophagus echinus Boucomont, 1914 | | | Onthophagus egenus (Harold, 1877) | ⇒ Onthophagus lemur Fabricius, 1781 | | Onthophagus eschscholtzi Boucomont, 1924 | | | Onthophagus falculatus Boucomont, 1914 | | | Onthophagus falsus Gillet, 1925 | | | Onthophagus fissinasus Fairmaire, 1895 | | | Onthophagus foedus Boucomont, 1914 | | | Onthophagus fossor Arrow, 1931 | | | Onthophagus frankrozendaali Huijbregts &..., 2012 | | | Onthophagus fukuyamai Ochi, Kon & ..., 2018 | | | Onthophagus fulvus Sharp, 1875 | | | Onthophagus fuscostriatus Boucomont, 1914 | | | Onthophagus gaesatus Boucomont, 1925 | | | Onthophagus gagates Hope, 1831 | | | Onthophagus gajo Krikken, 1977 | | | Onthophagus galeatus Boucomont, 1919 | | | Onthophagus gestroi (Harold, 1877) | ⇒ Onthophagus bellus d´Orbigny, 1905 | | Onthophagus gibbicollis Lansberge, 1885 | | | Onthophagus gonipa Krikken & Hu..., 2011 | | | Onthophagus griseoaeneus Lansberge, 1885 | | | Onthophagus hageni Lansberge, 1883 | | | Onthophagus hanboonsongae Masumoto, Oc..., 2021 | | | Onthophagus hartiniae Ochi & Kon, 2009 | | | Onthophagus heterorrhinus Lansberge, 1885 | | | Onthophagus hielkemai Krikken & Hu..., 2011 | | | Onthophagus hikidai Ochi & Kon, 2006 | | | Onthophagus hiranoyasutoshii Ochi & Kon, 2018 | | | Onthophagus hiroyukii Ochi, 2007 | | | Onthophagus hirsutulus Lansberge, 1883 | | | Onthophagus histeriformis Boucomont, 1914 | | | Onthophagus histrio d´Orbigny, 1902 | | | Onthophagus hoberlandti Balthasar, 1966 | | | Onthophagus hollowayi Krikken & Hu..., 2017 | | | Onthophagus holosericeus Harold, 1877 | | | Onthophagus holosericus Harold, 1877 | | | Onthophagus holzi Boucomont, 1914 | | | Onthophagus hosomai Ochi & Kon, 2014 | | | Onthophagus ifugaoensis Ochi & Kon, 2006 | | | Onthophagus imbutus Sharp, 1875 | | | Onthophagus imperator Castelnau, 1840 | | | Onthophagus incisus Harold, 1877 | | | Onthophagus indosinicus Boucomont, 1921 | | | Onthophagus inermivertex Boucomont, 1921 | | | Onthophagus infucatus Harold, 1877 | | | Onthophagus insulindicus Boucomont, 1914 | | | Onthophagus isanus Masumoto, Oc..., 2008 | | | Onthophagus itoianus Masumoto, Oc..., 2022 | | | Onthophagus itonoborui Masumoto, Oc..., 2012 | | | Onthophagus iumienus Masumoto, Oc..., 2007 | | | Onthophagus jacobeus Boucomont, 1924 | | | Onthophagus javanensis Balthasar, 1963 | | | Onthophagus javanoides Huijbregts &..., 2011 | | | Onthophagus jeannelianus Paulian, 1945 | | | Onthophagus jeannelianus jeannelianus | | | Onthophagus jingping Masumoto, Oc..., 2007 | | | Onthophagus kachinicus Kabakov, 2006 | | | Onthophagus kaengkrachangus Masumoto, Oc..., 2008 | | | Onthophagus kangeanus Paulian, 1936 | | | Onthophagus kaosoidowensis Masumoto, Oc..., 2008 | | | Onthophagus kapitensis Frey, 1971 | | | Onthophagus kashizakii Kon & Ochi, 2005 | | | Onthophagus katayamai Masumoto, Oc..., 2012 | | | Onthophagus kawaharai Ochi & Kon, 2007 | | | Onthophagus keikoae Ochi & Kon, 2014 | | | Onthophagus kiyoshii Ochi & Kon, 2007 | | | Onthophagus knapperti Krikken, 1981 | | | Onthophagus koechlei Masumoto, Oc..., 2011 | | | Onthophagus kolaka Huijbregts &..., 2009 | | | Onthophagus kongkaewensis Masumoto, Oc..., 2012 | | | Onthophagus koni Ochi, 2007 | | | Onthophagus konsarnensis Masumoto, Oc..., 2008 | | | Onthophagus kontumicus Kabakov, 1983 | | | Onthophagus kuluensis Bates, 1891 | | | Onthophagus kyokoae Masumoto & Ochi, 2015 | | | Onthophagus laevigatus Fabricius, 1798 | | | Onthophagus laevissimus d´Orbigny, 1905 | | | Onthophagus laichauensis Ochi, Kon & ..., 2021 | | | Onthophagus laminatus (MacLeay, 1864) | ⇒ Onthophagus lunatus Harold, 1868 | | Onthophagus lamyi Paulian, 1945 | | | Onthophagus langkawiensis Ochi & Kon, 2015 | | | Onthophagus laratinus Arrow, 1916 | | | Onthophagus lekmaengtibanphrao Masumoto & Ochi, 2015 | | | Onthophagus leknamnaous Masumoto & Ochi, 2015 | | | Onthophagus leomontanus Boucomont, 1914 | | | Onthophagus leusermontis Huijbregts &..., 2011 | | | Onthophagus lindu Huijbregts &..., 2009 | | | Onthophagus liwagensis Ochi & Kon, 2006 | | | Onthophagus lore Huijbregts &..., 2009 | | | Onthophagus luctuosus Boucomont, 1914 | | | Onthophagus lunatus Harold, 1868 | | | Onthophagus lunulifer Boucomont, 1914 | | | Onthophagus luteosignatus Lansberge, 1883 | | | Onthophagus luzonwallacei Ochi, Kawaha..., 2021 | | | Onthophagus maephaluangus Masumoto, Oc..., 2007 | | | Onthophagus maesalongensis Masumoto, Oc..., 2013 | | | Onthophagus magnini Paulian, 1933 | | | Onthophagus magnioculus Ochi & Kon, 2006 | | | Onthophagus magnipygus Boucomont, 1914 | | | Onthophagus malangensis Boucomont, 1914 | | | Onthophagus malasiacus Gillet, 1927 | | | Onthophagus manguliensis Boucomont, 1914 | | | Onthophagus maruchanus Masumoto, Oc..., 2012 | | | Onthophagus maruyamai Ochi, Kon & ..., 2014 | | | Onthophagus maryatiae Ochi & Kon, 2005 | | | Onthophagus matanyo Huijbregts &..., 2009 | | | Onthophagus matsuii Ochi & Kon, 2006 | | | Onthophagus megapacificus Ochi & Kon, 2006 | | | Onthophagus megathorax Gillet, 1921 | | | Onthophagus mekara Huijbregts &..., 2009 | | | Onthophagus mentaveiensis Boucomont, 1914 | | | Onthophagus miichunus Masumoto, Oc..., 2020 | | | Onthophagus mimikanus Balthasar, 1969 | | | Onthophagus mindanaensis Boucomont, 1914 | | | Onthophagus mindanaoensis Frey, 1971 | | | Onthophagus mniszechi Harold, 1869 | Kershaw´s burying beetle | | Onthophagus moajat Huijbregts &..., 2009 | | | Onthophagus mogo Krikken & Hu..., 2008 | | | Onthophagus montishannoniae Krikken & Hu..., 2008 | | | Onthophagus mordindangus Masumoto, Oc..., 2008 | | | Onthophagus morimotoi Masumoto, Oc..., 2021 | | | Onthophagus mouhoti Harold, 1875 | | | Onthophagus multipunctatus Boucomont, 1914 | | | Onthophagus mulutagal Ochi, Kon & ..., 2016 | | | Onthophagus namnaolekus Masumoto, Oc..., 2020 | | | Onthophagus namnaonus Masumoto, Oc..., 2020 | | | Onthophagus namnaous Masumoto, Oc..., 2013 | | | Onthophagus neervoorti Boucomont, 1914 | | | Onthophagus neofurcatus Goidanich, 1926 | | | Onthophagus nitefactus Harold, 1877 | | | Onthophagus obscurior Boucomont, 1914 | | | Onthophagus ochii Masumoto, 1988 | | | Onthophagus ochromerus Harold, 1877 | | | Onthophagus ohkuboi Ochi & Kon, 2006 | | | Onthophagus ontosatu Huijbregts &..., 2012 | | | Onthophagus opacihartiniae Ochi & Kon, 2015 | | | Onthophagus otai Ochi & Kon, 2006 | | | Onthophagus pacificus Lansberge, 1885 | | | Onthophagus palatus Boucomont, 1914 | | | Onthophagus palavanus (Huijbregts &..., 2011) | ⇒ Onthophagus javanensis Balthasar, 1963 | | Onthophagus palawantagal Ochi, Kon & ..., 2016 | | | Onthophagus pangmaphanus Masumoto, Oc..., 2022 | | | Onthophagus paracenricornis Ochi & Kon, 2014 | | | Onthophagus paracentricornis Ochi & Kon, 2014 | | | Onthophagus parapalatus Krikken & Hu..., 1988 | | | Onthophagus parenthesis Boucomont, 1913 | | | Onthophagus paroculus Krikken & Hu..., 1987 | | | Onthophagus parryi Harold, 1869 | | | Onthophagus parvidens (Frey, 1971) | ⇒ Onthophagus clermonti Paulian, 1931 | | Onthophagus pasotagal Ochi, Kon & ..., 2016 | | | Onthophagus pastillatus Boucomont, 1919 | | | Onthophagus pauper Boucomont, 1914 | | | Onthophagus pavidus Harold, 1877 | | | Onthophagus pedator Sharp, 1875 | | | Onthophagus peninsularis Boucomont, 1914 | | | Onthophagus peninsulomerus Huijbregts &..., 2011 | | | Onthophagus perakensis Ochi, Kon & ..., 2014 | | | Onthophagus personatus Boucomont, 1914 | | | Onthophagus phahompokus Masumoto & Ochi, 2015 | | | Onthophagus phakuaiensis Masumoto, Oc..., 2013 | | | Onthophagus phanaeicollis Lansberge, 1883 | | | Onthophagus phanaeides Frey, 1956 | | | Onthophagus phanaeiformis Boucomont, 1914 | | | Onthophagus phatoensis Masumoto, Oc..., 2008 | | | Onthophagus philippinensis Boucomont, 1919 | | | Onthophagus phillippsorum Krikken & Hu..., 1987 | | | Onthophagus phukhieoensis Masumoto, Oc..., 2007 | | | Onthophagus pimpasaleei Masumoto, Oc..., 2008 | | | Onthophagus planicollis Harold, 1880 | | | Onthophagus pollicatus Harold, 1879 | | | Onthophagus poringensis Ochi & Kon, 2005 | | | Onthophagus praedatus Harold, 1862 | | | Onthophagus pseudoarai Ochi & Kon, 2018 | | | Onthophagus pseudobrutus Kabakov, 1983 | | | Onthophagus pseudocoracinus Kabakov, 1983 | | | Onthophagus ptox Erichson, 1842 | | | Onthophagus purpurascens Boucomont, 1914 | | | Onthophagus pygidialis Lansberge, 1883 | | | Onthophagus quadripustulatus Fabricius, 1775 | | | Onthophagus quangnamensis Ochi, Kon & ..., 2021 | | | Onthophagus quasijohkii Ochi & Kon, 2005 | | | Onthophagus quasitagal Ochi & Kon, 2005 | | | Onthophagus ramosellus Bates, 1891 | | | Onthophagus ranongensis Masumoto, Oc..., 2008 | | | Onthophagus ranongjohkii Ochi, Kon & ..., 2016 | | | Onthophagus ratchataniensis Masumoto, Oc..., 2013 | | | Onthophagus remotus Harold, 1879 | | | Onthophagus rhyncophorus | | | Onthophagus ribbei Boucomont, 1914 | | | Onthophagus rizali Ochi, Kawaha..., 2021 | | | Onthophagus robertopoggii Ochi & Kon, 2006 | | | Onthophagus rorarius Harold, 1877 | | | Onthophagus rosenbergi Krikken & Hu..., 2011 | | | Onthophagus rosenbergi divergens Krikken & Hu..., 2011 | | | Onthophagus rosenbergi rosenbergi | | | Onthophagus rotundicollis Lansberge, 1883 | | | Onthophagus rouyeri Boucomont, 1914 | | | Onthophagus rubripennis (Arrow, 1907) | ⇒ Onthophagus taurus Schreber, 1759 | | Onthophagus ruficauda Arrow, 1931 | | | Onthophagus rugicollis Harold, 1880 | | | Onthophagus rutilans (Sharp, 1875) | ⇒ Onthophagus borneensis Harold, 1877 | | Onthophagus rutilans aborneensis Ochi, Kon & ..., 2009 | | | Onthophagus rutilans rutilans | | | Onthophagus sabahensis Ochi & Kon, 2006 | | | Onthophagus sabai Masumoto, Oc..., 2012 | | | Onthophagus saigonensis Boucomont, 1924 | | | Onthophagus saksphus Masumoto, Oc..., 2021 | | | Onthophagus samarensis Ochi, Kon & ..., 2023 | | | Onthophagus samoengus Masumoto, Oc..., 2008 | | | Onthophagus sanggona Huijbregts &..., 2009 | | | Onthophagus sangirensis Boucomont, 1914 | | | Onthophagus sangwalus Masumoto, Oc..., 2008 | | | Onthophagus sarasinorum Krikken & Hu..., 2011 | | | Onthophagus sceptrifer Boucomont, 1924 | | | Onthophagus schillhammeri Kabakov, 2006 | | | Onthophagus sciron Balthasar, 1969 | | | Onthophagus scrutator Harold, 1877 | | | Onthophagus sembeli Krikken & Hu..., 2008 | | | Onthophagus semiaereopictus Ochi, Kawaha..., 2021 | | | Onthophagus semigranosus Lansberge, 1883 | | | Onthophagus semipacificus Ochi & Kon, 2006 | | | Onthophagus semiperakensis Ochi, Kon & ..., 2014 | | | Onthophagus semiworoae Ochi, Kon & ..., 2014 | | | Onthophagus senescens Péringuey, 1908 | | | Onthophagus senex Boucomont, 1914 | | | Onthophagus seniculus (Fabricius, 1781) | ⇒ Onthophagus senescens Péringuey, 1908 | | Onthophagus sepilokensis Ochi & Kon, 2006 | | | Onthophagus seramicus Huijbregts &..., 2012 | | | Onthophagus serdangensis Lansberge, 1886 | | | Onthophagus seseba Krikken & Hu..., 2017 | | | Onthophagus setoculus Krikken & Hu..., 1987 | | | Onthophagus sexdentatus Boucomont, 1919 | | | Onthophagus shinichii Ochi & Kon, 2018 | | | Onthophagus shunichii Ochi, Kon & ..., 2021 | | | Onthophagus sibela Huijbregts &..., 2012 | | | Onthophagus sibuyanus Boucomont, 1924 | | | Onthophagus sidangeaus Masumoto, Oc..., 2021 | | | Onthophagus sideki Krikken & Hu..., 1987 | | | Onthophagus sihkahonoi Huijbregts &..., 2009 | | | Onthophagus simboroni Ochi & Kon, 2006 | | | Onthophagus sinagai Krikken & Hu..., 2008 | | | Onthophagus sopu Krikken & Hu..., 2017 | | | Onthophagus spathatus Boucomont, 1914 | | | Onthophagus spiculatus Boucomont, 1914 | | | Onthophagus statzi ♰ Krell, 1990 | | | Onthophagus subcornutus Boucomont, 1914 | | | Onthophagus sugihartoi Ochi, 2007 | | | Onthophagus suginokoichii Ochi & Kon, 2008 | | | Onthophagus sulawesiensis Krikken & Hu..., 2011 | | | Onthophagus sulawesijohkii Ochi, Kon & ..., 2016 | | | Onthophagus sumatramontanus Ochi & Kon, 2008 | | | Onthophagus sumatranus Lansberge, 1883 | | | Onthophagus sundanensis Lansberge, 1883 | | | Onthophagus surdus Boucomont, 1925 | | | Onthophagus sycophanta (Fairmaire, 1887) | ⇒ Euonthophagus gibbosus Scriba, 1790 | | Onthophagus sydneyensis Blackburn, 1903 | | | Onthophagus taeniatus Boucomont, 1914 | | | Onthophagus tagal Boucomont, 1924 | | | Onthophagus takaoi Ochi & Kon, 2014 | | | Onthophagus tambing Krikken & Hu..., 2011 | | | Onthophagus taoi Ochi & Kon, 2006 | | | Onthophagus thailaevis Masumoto, Oc..., 2008 | | | Onthophagus thainuaensis Masumoto, Oc..., 2008 | | | Onthophagus thungphraensis Masumoto, Oc..., 2020 | | | Onthophagus timorensis Boucomont, 1914 | | | Onthophagus tiniocelloides Boucomont, 1925 | | | Onthophagus tonkineus Gillet, 1921 | | | Onthophagus tonywhitteni Krikken & Hu..., 2011 | | | Onthophagus toraut Krikken & Hu..., 2011 | | | Onthophagus traginus Frey, 1972 | | | Onthophagus tragoides Boucomont, 1914 | | | Onthophagus transvestitus Huijbregts &..., 2009 | | | Onthophagus tricolor Boucomont, 1914 | | | Onthophagus tridentitibialis Ochi & Kon, 2008 | | | Onthophagus trochilus Arrow, 1931 | | | Onthophagus tsubakii Ochi & Kon, 2009 | | | Onthophagus tsutomui Ochi & Kon, 2016 | | | Onthophagus tungkamangensis Masumoto, Oc..., 2007 | | | Onthophagus tungkamungensis Masumoto, Oc..., 2008 | | | Onthophagus ubaidillahi Krikken & Hu..., 2012 | | | Onthophagus uedai Ochi & Kon, 2006 | | | Onthophagus ulugombakensis Ochi, Kon & ..., 2014 | | | Onthophagus unidentatus Boucomont, 1927 | | | Onthophagus urellus Boucomont, 1919 | | | Onthophagus vanasseni Krikken & Hu..., 2008 | | | Onthophagus vanderblomi Krikken & Hu..., 2008 | | | Onthophagus vanofwegeni Krikken & Hu..., 2008 | | | Onthophagus variolaris Lansberge, 1883 | | | Onthophagus ventralis Lansberge, 1883 | | | Onthophagus vethi Krikken, 1977 | | | Onthophagus viridiperakensis Ochi, Kon & ..., 2014 | | | Onthophagus volucer Balthasar, 1959 | | | Onthophagus vulpes Harold, 1877 | | | Onthophagus vulpinus Arrow, 1931 | | | Onthophagus wallacei Harold, 1871 | | | Onthophagus wangnamkieous Masumoto, Oc..., 2008 | | | Onthophagus wangtaichieni Masumoto, Oc..., 2020 | | | Onthophagus waterstradti Boucomont, 1914 | | | Onthophagus watuwila Krikken & Hu..., 2008 | | | Onthophagus wayaua Huijbregts &..., 2012 | | | Onthophagus wiebesi Krikken, 1977 | | | Onthophagus woroae Ochi & Kon, 2006 | | | Onthophagus woroae bruneiensis Ochi, Kon & ..., 2009 | | | Onthophagus woroae woroae | | | Onthophagus yangmunensis Masumoto & Ochi, 2015 | | | Onthophagus yasuhikoi Masumoto, Oc..., 2012 | | | Onthophagus yifer Krikken & Hu..., 2012 | | | Onthophagus yujii Masumoto, Oc..., 2012 | | | Onthophagus yumotoi Ochi & Kon, 2006 | | | Onthophagus zebu Boucomont, 1921 | | | Onthophagus zymoticus Moxey, 1963 | | | Onychothecus ateuchoides Boucomont, 1912 | | | Onychothecus corniclypeus Zhang, 1980 | | | Onychothecus riekoae Ochi & Kon, 1998 | | | Onychothecus tridentigeris Zelenka, 1992 | | | Panelus bakeri Boucomont, 1921 | | | Panelus borneensis Paulian, 1992 | | | Panelus brendelli Ochi, Kon & ..., 2009 | | | Panelus danumensis Ochi, Kon & ..., 2009 | | | Panelus fujitai Ochi, Kon & ..., 2008 | | | Panelus jaccoudi Paulian, 1980 | | | Panelus kalimantanicus Ochi, Kon & ..., 2009 | | | Panelus karasuyamai Ochi, Kon & ..., 2008 | | | Panelus manasi Paulian, 1983 | | | Panelus mendeli Ochi, Kon & ..., 2009 | | | Panelus minutus Boucomont, 1914 | | | Panelus orousseti Paulian, 1981 | | | Panelus quadridentatus Balthasar, 1952 | | | Panelus sabahi Paulian, 1992 | | | Panelus saigusai Ochi & Kon, 2016 | | | Panelus sumatranus Lansberge, 1885 | | | Panelus sumatrensis Ochi, Kon & ..., 1998 | | | Panelus tonkinensis Paulian, 1939 | | | Parachorius bolavensis Tarasov, 2017 | | | Parachorius fukiensis Balthasar, 1960 | | | Parachorius fungorum Kryzhanovsky..., 1966 | | | Parachorius javanus Boucomont, 1914 | | | Parachorius lannathai Hanboonsong ..., 2001 | | | Parachorius longipenis Tarasov, 2017 | | | Parachorius maruyamai Masumoto, Oc..., 2012 | | | Parachorius matsudai Ochi, Pham &..., 2016 | | | Parachorius newthayerae Tarasov, 2017 | | | Parachorius nudus Sharp, 1875 | | | Parachorius peninsularis Arrow, 1907 | | | Parachorius pseudojavanus Tarasov, 2017 | | | Parachorius semsanganus Tarasov & Keith, 2011 | | | Parachorius solodovnikovi Tarasov, 2017 | | | Parachorius thomsoni Harold, 1873 | | | Paragymnopleurus brahminus Waterhouse, 1890 | | | Paragymnopleurus maurus Sharp, 1875 | | | Paragymnopleurus melanarius Harold, 1867 | | | Paragymnopleurus planus Sharp, 1875 | | | Paragymnopleurus rudis Sharp, 1875 | | | Paragymnopleurus sinuatus Olivier, 1789 | | | Paragymnopleurus sinuatus abax Sharp, 1875 | | | Paragymnopleurus sinuatus assamensis Waterhouse, 1890 | | | Paragymnopleurus sinuatus productus Sharp, 1875 | | | Paragymnopleurus sinuatus sinuatus | | | Paragymnopleurus sinuatus szechouanicus Balthasar, 1934 | | | Paragymnopleurus sparsus Sharp, 1875 | | | Paragymnopleurus sparsus arnoldi Krikken & Hu..., 1987 | | | Paragymnopleurus sparsus javanus Krikken & Hu..., 1987 | | | Paragymnopleurus sparsus palawanicus Ochi & Kon, 1997 | | | Paragymnopleurus sparsus sparsus | | | Paragymnopleurus spinotus Boucomont, 1914 | | | Paragymnopleurus stipes Sharp, 1875 | | | Paragymnopleurus striatus Sharp, 1875 | | | Paragymnopleurus sumatrensis Ochi & Kon, 1997 | | | Paraphytus andoi Ochi & Kon, 2015 | | | Paraphytus dentifrons Lewis, 1895 | | | Paraphytus doriae Harold, 1877 | | | Paraphytus foveatus Boucomont, 1923 | | | Paraphytus harmandi Boucomont, 1923 | | | Paraphytus ritsemae Harold, 1880 | | | Paraphytus sumatranus Paulian & Ca..., 1981 | | | Pedaria decorsei (Boucomont, 1922) | ⇒ Pedaria taylori Waterhouse, 1890 | | Phacosoma fallacilaetum (Masumoto, 1989) | ⇒ Ochicanthon thai Paulian, 1987 | | Phacosoma thailandicum (Masumoto, 1989) | ⇒ Ochicanthon thailandicum Masumoto, 1989 | | Proagoderus amplexus Sharp, 1875 | | | Proagoderus gladiator Arrow, 1907 | | | Proagoderus mouhoti Harold, 1875 | | | Proagoderus schwaneri Snellen van ..., 1864 | | | Proagoderus victor Sharp, 1875 | | | Proagoderus watanabei Ochi & Kon, 2002 | | | Scarabaeus gangeticus Castelnau, 1840 | | | Sinapisoma minuta Boucomont, 1928 | | | Sinodrepanus besucheti Simonis, 1985 | | | Sinodrepanus exsul Sharp, 1875 | | | Sinodrepanus falsus Sharp, 1875 | | | Sinodrepanus rex Boucomont, 1912 | | | Sinodrepanus similis Simonis, 1985 | | | Sinodrepanus thailandicus Ochi, Kon & ..., 2004 | | | Sisyphus bowringi White, 1844 | | | Sisyphus denticrus (Fairmaire, 1886) | ⇒ Sisyphus neglectus Gory, 1833 | | Sisyphus indicus Hope, 1831 | | | Sisyphus laoticus Arrow, 1927 | | | Sisyphus longipes Olivier, 1789 | | | Sisyphus maniti Masumoto, 1988 | | | Sisyphus neglectus Gory, 1833 | | | Sisyphus thoracicus Sharp, 1875 | | | Sisyphus thoracicus chaiyaphumensis Hanboonsong ..., 2000 | | | Sisyphus thoracicus thoracicus | | | Sulcophanaeus velutinus Murray, 1856 | | | Synapsis birmanicus Gillet, 1907 | | | Synapsis boonlongi Hanboonsong ..., 1999 | | | Synapsis cambeforti (Ochi, Kon & ..., 2008) | ⇒ Synapsis ritsemae Lansberge, 1874 | | Synapsis dickinsoni Hanboonsong ..., 1999 | | | Synapsis horaki Zidek & Pokorny, 2010 | | | Synapsis kiuchii Hanboonsong ..., 1999 | | | Synapsis ochii Masumoto, 1995 | | | Synapsis ovalis Boucomont, 1920 | | | Synapsis puluongensis Bui & Bonkowski, 2018 | | | Synapsis punctatus Ochi, Kon & ..., 2008 | | | Synapsis ritsemae Lansberge, 1874 | | | Synapsis roslihashimi Ochi, Kon & ..., 2008 | | | Synapsis satoi Ochi & Kon, 2007 | | | Synapsis simplex Sharp, 1875 | | | Synapsis strnadi Kral, 2002 | | | Synapsis thoas (Sharp, 1875) | ⇒ Synapsis ritsemae Lansberge, 1874 | | Synapsis tridens Sharp, 1881 | | | Synapsis vietnamicus Ochi, Kon & ..., 2020 | | | Synapsis yama Gillet, 1911 | | | Synapsis yunnanus Arrow, 1933 | | | Temnoplectron arfakensis Ochi, Kon & ..., 2017 | | | Temnoplectron waigeoensis Ochi, Kon & ..., 2017 | | | Tesserodon seramicum Krikken & Hu..., 2009 | | | Tesserodon waigeoensis Ochi, Kon & ..., 2017 | | | Tibiodrepanus hircus Wiedemann, 1823 | | | Tibiodrepanus setosus Wiedemann, 1823 | | | Tibiodrepanus sinicus Harold, 1868 | | | Tiniocellus averyanovi Kabakov, 2014 | | | Yvescambefortius sarawacus Gillet, 1926 | |
Subnodes of Scarabaeinae with occurrences in Southeast Asia | Tribe | Ateuchini
Montreuil, 1998
| 3
12 | Genera
Species |
Taxom
| | Tribe | Canthonini
Péringuey, 1900
| 10
44 | Genera
Species |
Taxom
| | Tribe | Coprini
Kolbe, 1805
| 7
138 | Genera
Species |
Taxom
| | Tribe | Deltochilini
Lacordaire, 1856
| 1
3 | Genus
Species |
Taxom
| | Tribe | Gymnopleurini
Lacordaire, 1856
| 3
18 | Genera
Species |
Taxom
| | Tribe | Oniticellini
d´Orbigny, 1916
| 12
37 | Genera
Species |
Taxom
| | Tribe | Onitini
Hope, 1837
| 2
24 | Genera
Species |
Taxom
| | Tribe | Onthophagini
Lacordaire, 1856
| 13
636 | Genera
Species |
Taxom
| | Tribe | Phanaeini
MacLeay, 1819
| 3
2 | Genera
Species |
Taxom
| | Tribe | Scarabaeini
Latreille, 1802
| 2
2 | Genera
Species |
Taxom
| | Tribe | Sisyphini
Mulsant, 1842
| 1
8 | Genus
Species |
Taxom
|
Links, sources and further information Scarabaeinae at artfakta.se
Scarabaeinae at dyntaxa.se
Scarabaeinae at itis.gov
|
|
|
|
