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TAXFAMGENSPESUB
Checklist for Subfamily Scarabaeinae Latreille, 1802 (Dung Beetles)
and region Transvaal (Republic South Africa)


Class
 
Insecta
Linnaeus, 1758
Insects, True insects
380
3140
5821
Families
Genera
Species
 
Taxom
 

Subclass
 
Pterygota
Lang, 1888
Winged insects
380
3140
5821
Families
Genera
Species
 
Taxom
 

Infraclass
 
Neoptera
van der Wulp, 1890
Modern Wing-folding insects
357
3054
5696
Families
Genera
Species
 
Taxom
 

Superorder
 
Holometabola
Heider, 1889
Holometabolous Insects
229
2399
4651
Families
Genera
Species
 
Taxom
 

Order
 
Coleoptera
Linnaeus, 1758
Beetles, Coleopterans
63
937
1960
Families
Genera
Species
 
Taxom
 

Suborder
 
Polyphaga
Emery, 1886
Polyphags
56
814
1700
Families
Genera
Species
 
Taxom
 

Infraorder
 
Scarabaeiformia
Crowson, 1960
9
253
765
Families
Genera
Species
 
Taxom
 

Superfamily
 
Scarabaeoidea
Latreille, 1802
Bess Beetles, Chafer, Dung Beetles, D´or Beetles
9
253
765
Families
Genera
Species
 
Taxom
 
FamilyScarabaeidae
Latreille, 1802
Dung beetles, Flower beetles, Fruit chafers, Rain beetles
229
698
Genera
Species
 
Taxom
 
SubfamilyScarabaeinae
Latreille, 1802
Dung Beetles, True dung beetles
48
305
Genera
Species
 
Taxom
 


Genera for Transvaal
48 Genera with 305 Species
#ACDEFGHKLMNOPST
Allogymnopleurus Janssens, 18553SpeciesTaxom
Anachalcos Hope, 18571SpeciesTaxom
Caccobius Thomson, 19045SpeciesTaxom
Canthidium Erichson, 18671SpeciesTaxom
Catharsius Hope, 185713SpeciesTaxom
Cheironitis van Lansberge, 17764SpeciesTaxom
Circellium Latreille, 17811SpeciesTaxom
Cleptocaccobius Cambefort, 18572SpeciesTaxom
Copris Geoffroy, 1962Mexican dung beetle24SpeciesTaxom
Coptodactyla Burmeister, 18771SpeciesTaxom
Coptorhina Hope, 18573SpeciesTaxom
Cyptochirus Lesne, 18512SpeciesTaxom
Delopleurus Erichson, 18571SpeciesTaxom
Digitonthophagus Balthasar, 17871SpeciesTaxom
Drepanocerus Kirby, 18514SpeciesTaxom
Eodrepanus Barbero, Pal..., 19011SpeciesTaxom
Epirinus Reiche, 19016SpeciesTaxom
Euoniticellus Janssens, 19533SpeciesTaxom
Euonthophagus Balthasar, 18551SpeciesTaxom
Frankenbergerius5SpeciesTaxom
Garreta Janssens, 19674SpeciesTaxom
Gymnopleurus Illiger, 18439SpeciesTaxom
Heliocopris Hope, 17989SpeciesTaxom
Heteronitis Gillet, 18621SpeciesTaxom
Kheper Janssens, 19867SpeciesTaxom
Latodrepanus Krikken, 18571SpeciesTaxom
Liatongus Reitter, 18401SpeciesTaxom
Metacatharsius Paulian, 18577SpeciesTaxom
Milichus Péringuey, 19041SpeciesTaxom
Neosisyphus Müller, 18188SpeciesTaxom
Odontoloma Boheman, 19083SpeciesTaxom
Oniticellus Serville, 18404SpeciesTaxom
Onitis Fabricius, 185720SpeciesTaxom
Onthophagus Latreille, 192784SpeciesTaxom
Pachylomera Griffith & P..., 18281SpeciesTaxom
Peckolus Scholtz & Ho..., 20082SpeciesTaxom
Pedaria Laporte, 18576SpeciesTaxom
Phalops Erichson, 18856SpeciesTaxom
Pinacotarsus Harold, 18751SpeciesTaxom
Pleuronitis Lansberge, 18401SpeciesTaxom
Proagoderus Lansberge, 188411SpeciesTaxom
Sarophorus Erichson, 18571SpeciesTaxom
Scarabaeus Linnaeus, 190121SpeciesTaxom
Sceliages Westwood, 18442SpeciesTaxom
Sisyphus Latreille, 181811SpeciesTaxom
Stiptopodius Harold, 19011SpeciesTaxom
Tibiodrepanus Krikken, 18401SpeciesTaxom
Tiniocellus Péringuey, 18512SpeciesTaxom


Species list for Transvaal
305 Species and 7 Subspecies
Allogymnopleurus consocius Péringuey, 1901
Allogymnopleurus splendidus Bertoloni, 1849
Allogymnopleurus thalassinus Klug, 1855
Anachalcos convexus Boheman, 1857
Caccobius cavatus d´Orbigny, 1908
Caccobius ferrugineus Fahraeus, 1857
Caccobius fuliginosus (Roth, 1851)⇒ Caccophilus reticuliger d´Orbigny, 1904
Caccobius obtusus Fahraeus, 1857
Caccobius reticuliger d´Orbigny, 1904
Canthidium smaragdinum Harold, 1867
Catharsius calaharicus Kolbe, 1893
Catharsius dubius (Péringuey, 1901)⇒ Catharsius molossus Linnaeus, 1758
Catharsius longiceps Gillet, 1907
Catharsius marcellus Kolbe, 1893
Catharsius melancholicus (Boheman, 1860)⇒ Catharsius longiceps Gillet, 1907
Catharsius molossus Linnaeus, 1758
Catharsius pandion Harold, 1877
Catharsius philus Kolbe, 1893
Catharsius sesostris Waterhouse, 1888
Catharsius tricornutus DeGeer, 1778
Catharsius ulysses Boheman, 1857
Catharsius vansoni Ferreira, 1960
Catharsius vitulus Boheman, 1857
Cheironitis arrowi Janssens, 1937
Cheironitis hoplosternus Harold, 1868
Cheironitis indicus (Lansberge, 1875)⇒ Cheironitis arrowi Janssens, 1937
Cheironitis scabrosus Fabricius, 1776
Circellium bacchus Fabricius, 1781Flightless dung beetle
Cleptocaccobius seminulum Klug, 1855
Cleptocaccobius viridicollis Fahraeus, 1857
Copris amyntor (Klug, 1855)⇒ Copris harrisi Waterhouse, 1891
Copris armiger Gillet, 1910
Copris caelatus Fabricius, 1794
Copris cambeforti Nguyen-phung, 1988
Copris cornifrons Boheman, 1860
Copris corniger (Sahlberg, 1823)⇒ Onthophagus taurus Schreber, 1759
Copris denticulatus Nguyen-phung, 1988
Copris elphenor Klug, 1855
Copris evanidus Klug, 1855
Copris fallaciosus Gillet, 1907Kenyan dung beetle
Copris fidius Olivier, 1789
Copris harrisi Waterhouse, 1891
Copris jacchoides Nguyen-phung..., 1987
Copris jacchus Lang, 1971
Copris macer Péringuey, 1901
Copris mesacanthus Harold, 1878
Copris mesacanthus transvaalensis Nguyen-phung, 1988
Copris misellus Péringuey, 1901
Copris obesus Boheman, 1857
Copris orion Klug, 1835
Copris orion caffer Gillet, 1908
Copris ritsemae Harold, 1875
Copris urus (Boheman, 1857)⇒ Catharsius molossus Linnaeus, 1758
Copris vilhenai Ferreira, 1962
Coptodactyla subaenea Harold, 1877
Coptorhina auspicata Péringuey, 1901
Coptorhina klugi Hope, 1833
Coptorhina nitidipennis Boheman, 1857
Cyptochirus ambiguus (Kirby, 1828)⇒ Drepanocerus trogiformis Roth, 1851
Cyptochirus trogiformis Roth, 1851
Delopleurus pullus Boheman, 1857
Digitonthophagus gazella Fabricius, 1787Brown dung beetle
Drepanocerus ambiguus Kirby, 1828
Drepanocerus kirbyi Kirby, 1828
Drepanocerus patrizii Boucomont, 1923
Drepanocerus trogiformis Roth, 1851
Eodrepanus fastiditus Péringuey, 1901
Epirinus asper Péringuey, 1901
Epirinus gratus Péringuey, 1901
Epirinus mucrodentatus Scholtz & Ho..., 1987
Epirinus punctatus Scholtz & Ho..., 1987
Epirinus pygidialis Scholtz & Ho..., 1987
Epirinus validus Péringuey, 1901
Euoniticellus africanus Harold, 1873Greater sandy dung beetle
Euoniticellus triangulatus Harold, 1873
Euoniticellus zumpti Janssens, 1953
Euonthophagus carbonarius Klug, 1855
Frankenbergerius armatus Boheman, 1857
Frankenbergerius barratti Waterhouse, 1876
Frankenbergerius forcipatus Harold, 1880
Frankenbergerius gomesi Ferreira, 1954
Frankenbergerius nanus Frolov & Sch..., 2005
Garreta caffer Fahraeus, 1857
Garreta nitens Olivier, 1789
Garreta unicolor Fahraeus, 1857
Garreta zumpti Frey, 1967
Gymnopleurus aenescens Wiedemann, 1821
Gymnopleurus humanus MacLeay, 1821
Gymnopleurus humeralis Klug, 1855
Gymnopleurus ignitus Klug, 1855
Gymnopleurus leei Fabricius, 1792
Gymnopleurus reichei Waterhouse, 1890
Gymnopleurus sericeifrons Fairmaire, 1887
Gymnopleurus thelwalli Waterhouse, 1890
Gymnopleurus virens Erichson, 1843
Heliocopris andersoni Bates, 1868
Heliocopris colossus Bates, 1868
Heliocopris dilloni Guérin-Ménev..., 1847
Heliocopris erycoides Felsche, 1907
Heliocopris hamadryas Fabricius, 1775
Heliocopris hamifer Harold, 1878
Heliocopris japetus Klug, 1855
Heliocopris neptunus Boheman, 1857
Heliocopris pirmal Fabricius, 1798
Heteronitis castelnaui Harold, 1862
Kheper clericus Boheman, 1857
Kheper cupreus Castelnau, 1840
Kheper lamarcki MacLeay, 1821
Kheper nigroaeneus Boheman, 1857
Kheper platynotus Bates, 1888
Kheper subaeneus Harold, 1869
Kheper zurstrasseni Davis, 1986
Latodrepanus laticollis Fahraeus, 1857
Liatongus militaris Castelnau, 1840Yellow Shouldered Dung Beetle
Metacatharsius anderseni Waterhouse, 1891
Metacatharsius exiguus Boheman, 1860
Metacatharsius latifrons Harold, 1868
Metacatharsius opacus Waterhouse, 1891
Metacatharsius pumilionis Wallengren, 1881
Metacatharsius transvaalensis Balthasar, 1968
Metacatharsius troglodytes (Boheman, 1857)⇒ Metacatharsius anderseni Waterhouse, 1891
Milichus rhodesianus Péringuey, 1904
Neosisyphus calcaratus Klug, 1855
Neosisyphus confrater Kolbe, 1914
Neosisyphus fortuitus Péringuey, 1901
Neosisyphus kuehni Haaf, 1955
Neosisyphus rubripes Péringuey, 1901
Neosisyphus rubrus Paschalidis, 1974
Neosisyphus rugosus Gory, 1833
Neosisyphus spinipes Thunberg, 1818Grey dungball roller
Odontoloma louwi Howden & Sch..., 1987
Odontoloma pauxillum Boheman, 1857
Odontoloma quadridens d´Orbigny, 1908
Oniticellus egregius Klug, 1855
Oniticellus formosus (Chevrolat, 1830)⇒ Oniticellus pictus Hausmann, 1807
Oniticellus pictus Hausmann, 1807
Oniticellus planatus Castelnau, 1840
Onitis aeruginosus Klug, 1855
Onitis alexis (Klug, 1835)⇒ Cheironitis ungaricus ungaricus Herbst, 1789
Onitis caffer Boheman, 1857
Onitis confusus Boheman, 1860
Onitis crenatus Reiche, 1847
Onitis cryptodus Gillet, 1918
Onitis curvipes Lansberge, 1875
Onitis deceptor Péringuey, 1901
Onitis fulgidus (Klug, 1855)⇒ Pleuronitis fulgidus Castelnau, 1840
Onitis mendax Gillet, 1918
Onitis minutus Lansberge, 1875
Onitis obenbergeri Balthasar, 1942
Onitis paraconfusus Ferreira, 1976
Onitis parainflaticollis Ferreira, 1977
Onitis perbrincki Ferreira, 1958
Onitis picticollis Boheman, 1857
Onitis pseudosetosus Ferreira, 1976
Onitis tortuosus Houston, 1983
Onitis uncinatus Klug, 1855
Onitis viridulus (Boheman, 1857)⇒ Onitis pecuarius Lansberge, 1875
Onthophagus (Euonthophagus) carbonarius Klug, 1855
Onthophagus (Onthophagus) taurus Schreber, 1759Bull Headed Dung Beetle
Onthophagus (Onthophagus) trituber Wiedemann, 1823
Onthophagus marginalis przewalski Kabakov, 2006
Onthophagus (Proagoderus) alcedo d´Orbigny, 1913
Onthophagus (Proagoderus) alcyon (Klug, 1855)⇒ Onthophagus alcedo d´Orbigny, 1913
Onthophagus (Proagoderus) alcyonides d´Orbigny, 1913
Onthophagus (Proagoderus) aureiceps d´Orbigny, 1902
Onthophagus (Proagoderus) pegesimallus Marcus, 1917
Onthophagus absyrtus Balthasar, 1946
Onthophagus acutus d´Orbigny, 1908
Onthophagus aequepubens d´Orbigny, 1905
Onthophagus aeruginosus Roth, 1851
Onthophagus albipodex d´Orbigny, 1902
Onthophagus apiciosus d´Orbigny, 1902
Onthophagus aschenborni Frey, 1975
Onthophagus asperulus d´Orbigny, 1905
Onthophagus beiranus Péringuey, 1908
Onthophagus bicavifrons d´Orbigny, 1902
Onthophagus biconifer d´Orbigny, 1905
Onthophagus binodis Thunberg, 1818Humpbacked Dung Beetle
Onthophagus bovinus Péringuey, 1892
Onthophagus carpophilus Pereira & Ha..., 1961
Onthophagus chalcostolus d´Orbigny, 1902
Onthophagus crassicollis Boucomont, 1913
Onthophagus cribripennis d´Orbigny, 1902
Onthophagus croesulus Bates, 1888
Onthophagus cupricollis Péringuey, 1888
Onthophagus cyaneoniger d´Orbigny, 1902
Onthophagus dedecor Wallengren, 1881
Onthophagus deterrens Péringuey, 1901
Onthophagus discretus Péringuey, 1901
Onthophagus ebenus Péringuey, 1888
Onthophagus echinus Boucomont, 1914
Onthophagus filicornis Harold, 1873
Onthophagus fimetarius Roth, 1851
Onthophagus flavimargo d´Orbigny, 1902
Onthophagus flavolimbatus Klug, 1855
Onthophagus fritschi d´Orbigny, 1902
Onthophagus fugitivus Péringuey, 1901
Onthophagus giraffa Hausmann, 1807
Onthophagus granulum d´Orbigny, 1904
Onthophagus graphicus Wallengren, 1881
Onthophagus hyaena Fabricius, 1801
Onthophagus interstitialis (Fahraeus, 1857)⇒ Euonthophagus carbonarius Klug, 1855
Onthophagus jeanneli diversiceps Frey, 1975
Onthophagus lamnifer d´Orbigny, 1902
Onthophagus leucopygus (Harold, 1867)⇒ Onthophagus albipodex d´Orbigny, 1902
Onthophagus lituratus Roth, 1851
Onthophagus lugens (Fahraeus, 1857)⇒ Onthophagus fimetarius Roth, 1851
Onthophagus lugubris (Fahraeus, 1857)⇒ Onthophagus lituratus Roth, 1851
Onthophagus marshalli d´Orbigny, 1908
Onthophagus minutus (Hausmann, 1807)⇒ Onthophagus centricornis Fabricius, 1798
Onthophagus modestus Harold, 1862
Onthophagus monodon Fahraeus, 1857
Onthophagus nanus (Harold, 1878)⇒ Onthophagus waterhousei Gillet, 1927
Onthophagus obtusicornis (Fahraeus, 1857)⇒ Onthophagus marginalis przewalski Kabakov, 2006
Onthophagus obtutus Péringuey, 1901
Onthophagus pallidipennis (Fahraeus, 1857)⇒ Onthophagus rasipennis d´Orbigny, 1908
Onthophagus parenthesis Boucomont, 1913
Onthophagus pauxillus d´Orbigny, 1902
Onthophagus pilosus (Fahraeus, 1857)⇒ Onthophagus echinus Boucomont, 1914
Onthophagus pinguis Gerstaecker, 1871
Onthophagus praedatus Harold, 1862
Onthophagus pugionatus Fahraeus, 1857
Onthophagus pullus (Roth, 1851)⇒ Onthophagus parenthesis Boucomont, 1913
Onthophagus quadrinodosus Fahraeus, 1857
Onthophagus rasipennis d´Orbigny, 1908
Onthophagus setosus Fahraeus, 1857
Onthophagus signatus Fahraeus, 1857
Onthophagus simoni d´Orbigny, 1902
Onthophagus subplanus d´Orbigny, 1902
Onthophagus sugillatus Klug, 1855
Onthophagus suturalis (Péringuey, 1888)⇒ Onthophagus trituber Wiedemann, 1823
Onthophagus talpa Fahraeus, 1857
Onthophagus temporalis d´Orbigny, 1902
Onthophagus tonsus d´Orbigny, 1902
Onthophagus trinodosus Fahraeus, 1857
Onthophagus variegatus (Fabricius, 1798)⇒ Onthophagus filicornis Harold, 1873
Onthophagus ventrosus d´Orbigny, 1905
Onthophagus verticalis (Fahraeus, 1857)⇒ Onthophagus praedatus Harold, 1862
Onthophagus vigens Péringuey, 1901
Onthophagus vinctus Erichson, 1843
Onthophagus waterhousei Gillet, 1927
Pachylomera femoralis Kirby, 1828
Peckolus alpinus Howden & Sch..., 1988
Peckolus poenskopius Deschodt & C..., 2008
Pedaria barrei Josso & Prév..., 2006
Pedaria conformis Péringuey, 1901
Pedaria cylindrica Fahraeus, 1857
Pedaria dellacasai Josso & Prév..., 2009
Pedaria murphyi murphyi
Pedaria picea Fahraeus, 1857
Phalops ardea Klug, 1855
Phalops boschas Klug, 1855
Phalops bubalus Harold, 1867
Phalops dregei Harold, 1867
Phalops flavocinctus Klug, 1855
Phalops rufosignatus (Lansberge, 1885)⇒ Phalops dregei Harold, 1867
Pinacotarsus dohrni Harold, 1875
Pleuronitis fulgidus Castelnau, 1840
Proagoderus aureiceps d´Orbigny, 1902
Proagoderus bicallosus Klug, 1855
Proagoderus brucei (d´Orbigny, 1902)⇒ Onthophagus chalcostolus d´Orbigny, 1902
Proagoderus chalcostolus d´Orbigny, 1902
Proagoderus dives Harold, 1877
Proagoderus lanista Castelnau, 1840
Proagoderus loricatus Klug, 1855
Proagoderus rangifer Klug, 1855
Proagoderus sapphirinus (Péringuey, 1901)⇒ Onthophagus aureiceps d´Orbigny, 1902
Proagoderus tersidorsis d´Orbigny, 1902
Proagoderus unidens Quedenfeldt, 1884
Sarophorus costatus Fahraeus, 1857
Scarabaeus ambiguus (Boheman, 1857)⇒ Gnorimus variabilis Linnaeus, 1758
Scarabaeus anderseni Waterhouse, 1890
Scarabaeus bohemani Harold, 1868
Scarabaeus caffer Boheman, 1857
Scarabaeus clanceyi Ferreira, 1954
Scarabaeus convexus Hausmann, 1807
Scarabaeus deludens zur Strassen, 1961
Scarabaeus ebenus Klug, 1855
Scarabaeus funebris Boheman, 1857
Scarabaeus galenus Westwood, 1847
Scarabaeus goryi Castelnau, 1840
Scarabaeus heqvisti zur Strassen, 1962
Scarabaeus inguisitus
Scarabaeus inquisitus Péringuey, 1908
Scarabaeus interstitialis Boheman, 1857
Scarabaeus natalensis zur Strassen, 1962
Scarabaeus rusticus Boheman, 1857
Scarabaeus schulzeae Ferreira, 1969
Scarabaeus vicinus (Janssens, 1940)⇒ Lucanus cervus Linnaeus, 1758
Scarabaeus westwoodi (Harold, 1869)⇒ Scarabaeus galenus Westwood, 1847
Scarabaeus zambesianus Péringuey, 1901
Sceliages difficilis zur Strassen, 1965
Sceliages hippias Westwood, 1844
Sisyphus alveatus Boucomont, 1935
Sisyphus caffer Boheman, 1857
Sisyphus costatus Thunberg, 1818
Sisyphus crispatus Gory, 1833
Sisyphus crispatus crispatus
Sisyphus fasciculatus Boheman, 1857
Sisyphus goryi (Harold, 1859)⇒ Sisyphus caffer goryi Harold, 1859
Sisyphus impressipennis Lansberge, 1886
Sisyphus muricatus Olivier, 1789
Sisyphus ocellatus nanniscus Péringuey, 1901
Sisyphus spinipes (Thunberg, 1818)⇒ Neosisyphus spinipes Thunberg, 1818
Stiptopodius singularis Péringuey, 1901
Tibiodrepanus sulcicollis Castelnau, 1840
Tiniocellus eurypygus Branco, 2010
Tiniocellus spinipes Roth, 1851

Subnodes of Scarabaeinae with occurrences in Transvaal
TribeAteuchini
Montreuil, 1998
4
11
Genera
Species

Taxom

TribeCanthonini
Péringuey, 1900
5
13
Genera
Species

Taxom

TribeCoprini
Kolbe, 1805
6
53
Genera
Species

Taxom

TribeGymnopleurini
Lacordaire, 1856
3
16
Genera
Species

Taxom

TribeOniticellini
d´Orbigny, 1916
9
19
Genera
Species

Taxom

TribeOnitini
Hope, 1837
4
26
Genera
Species

Taxom

TribeOnthophagini
Lacordaire, 1856
10
113
Genera
Species

Taxom

TribeScarabaeini
Latreille, 1802
4
31
Genera
Species

Taxom

TribeSisyphini
Mulsant, 1842
2
18
Genera
Species

Taxom



Links, sources and further information
Scarabaeinae at artfakta.se
Scarabaeinae at dyntaxa.se
Scarabaeinae at itis.gov